738 lines
94 KiB
XML
738 lines
94 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.855.33013" ID-GBIF-Dataset="74c6bbe7-c9cc-468d-904e-61408ceff72e" ID-PMC="PMC6586674" ID-Pensoft-Pub="1313-2970-855-109" ID-Pensoft-UUID="F460A02DFFD1FB56FFF2FF94FFB2FFAC" ID-PubMed="31244545" ID-ZBK="F5A7AE8B38834CFD859FB2F3F9A079C0" ID-Zenodo-Dep="3251190" ID-ZooBank="F5A7AE8B38834CFD859FB2F3F9A079C0" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-855-109" ModsDocOrigin="ZooKeys 855" ModsDocTitle="Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera, Hydrophilidae, Acidocerinae)" checkinTime="1560464664927" checkinUser="pensoft" docAuthor="Giron, Jennifer C. & Short, Andrew Edward Z." docDate="2019" docId="43F651D6F22DCEE79CC1C8646EB153FF" docLanguage="en" docName="ZooKeys 855: 109-154" docOrigin="ZooKeys 855" docSource="http://dx.doi.org/10.3897/zookeys.855.33013" docTitle="Primocerus Girón & Short 2019, gen. nov." docType="treatment" docUuid="0EA5176F-B2BB-4E50-8799-C93F09B412B6" docUuidSource="ZooBank" docVersion="6" id="F460A02DFFD1FB56FFF2FF94FFB2FFAC" lastPageNumber="137" masterDocId="F460A02DFFD1FB56FFF2FF94FFB2FFAC" masterDocTitle="Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="154" masterPageNumber="109" pageNumber="129" updateTime="1732488762355" updateUser="ExternalLinkService">
|
||
<mods:mods id="DC3167085A58D4A6987512824CC27D05" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5C32071B5F5B4635AD25A8FBB8917A69">
|
||
<mods:title id="20DDC1712EA727DC8E11B7DE23AF25B9">Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FC8461E9284A159926F24BD049B1DD06" type="personal">
|
||
<mods:role id="8C1F7356EB60FCEA38B9CEA2D3011C32">
|
||
<mods:roleTerm id="8B12003B5FDD43D46082A0E36CFAF687">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="128D1AEEDFCF1AE6DBD6300893ACCCCD">Giron, Jennifer C.</mods:namePart>
|
||
<mods:affiliation id="23009D5BFE039F6B7FED92004744BEA2">Department of Ecology & Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="5686ABCB822620559DE73E35655D9740" type="orcid">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="E351C95354C2ED8A1A20B34964BE1EF5" type="email">entiminae@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="256CB7E13FCC96A0D120B81CC041CC05" type="personal">
|
||
<mods:role id="BF65F1089A89632B80D56DD973EBC6B3">
|
||
<mods:roleTerm id="01AC3FF932051C2CD10198208D751599">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B0C2E84F0170F34EC2185798F7C390AE">Short, Andrew Edward Z.</mods:namePart>
|
||
<mods:affiliation id="7280D09F980471F4BDF912605717EA86">Department of Ecology & Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="23155AEF5C5FDF47C23E6D4801F4B8B1">text</mods:typeOfResource>
|
||
<mods:relatedItem id="127CF082376379CAA00507BCE0FEE54F" type="host">
|
||
<mods:titleInfo id="B389C99E93133623197423C346505D30">
|
||
<mods:title id="95DD254303F8721DB95D828F9B131F25">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9C4DF7EA660830179DC1856D18510A49">
|
||
<mods:date id="419913C3B016FC5B121A121A9E4CEA75">2019</mods:date>
|
||
<mods:detail id="6571F01D43499313C29CD0D623D14B1C" type="volume">
|
||
<mods:number id="8C3880D70628BAE485EC19A525558A48">855</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B3F702122D248D5736F44D9EF1D87E75" unit="page">
|
||
<mods:start id="E16D377F2CBF9394DCC76850B1CEC37A">109</mods:start>
|
||
<mods:end id="50386200F3BEB32A6ACD4C798A976BCB">154</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="C178198135F3E3DBB79FAEA44F3C64BE">
|
||
<mods:url id="BC7C4C3242930CA0E5558778C5855595">http://dx.doi.org/10.3897/zookeys.855.33013</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="B541DF889B34336F5A22A6C3A4932282">journal article</mods:classification>
|
||
<mods:identifier id="F4E5D2BE42D0606F9485577E189B85EC" type="DOI">http://dx.doi.org/10.3897/zookeys.855.33013</mods:identifier>
|
||
<mods:identifier id="D0494DAA867A32929A770464C5696398" type="Pensoft-Pub">1313-2970-855-109</mods:identifier>
|
||
<mods:identifier id="3194BA27D39A28E27F4173E6F7BE928A" type="ZooBank">F5A7AE8B38834CFD859FB2F3F9A079C0</mods:identifier>
|
||
<mods:identifier id="B94AE16A09DC10E580028170F3CCDBC1" type="Pensoft-UUID">F460A02DFFD1FB56FFF2FF94FFB2FFAC</mods:identifier>
|
||
<mods:identifier id="C41909F99931DFDB82FC31681542D519" type="Zenodo-Dep">3251190</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="43F651D6F22DCEE79CC1C8646EB153FF" ID-GBIF-Taxon="158217533" LSID="urn:lsid:zoobank.org:act:0EA5176F-B2BB-4E50-8799-C93F09B412B6" httpUri="http://treatment.plazi.org/id/43F651D6F22DCEE79CC1C8646EB153FF" lastPageId="28" lastPageNumber="137" pageId="20" pageNumber="129" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_subFamily="Acidocerinae">
|
||
<subSubSection id="0D5FC87C3F08D7E87820E8EDC07508EE" pageId="20" pageNumber="129" type="nomenclature">
|
||
<paragraph id="11D446BBF747A0007A5A79A295051FA5" pageId="20" pageNumber="129">
|
||
<taxonomicName id="7FA33BD4746DFE16622EB59697D669AC" ID-CoL="8XP36" LSID="43F651D6-F22D-CEE7-9CC1-C8646EB153FF" authority="Girón & Short, 2019" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="20" pageNumber="129" phylum="Arthropoda" rank="genus" status="gen. nov.">Primocerus</taxonomicName>
|
||
<taxonomicNameLabel id="12AFFEC80B6EEEB1D8608B167DF5A564" pageId="20" pageNumber="129">gen. nov.</taxonomicNameLabel>
|
||
<figureCitation id="82E2D22153834313C09B2CC303F6CBFB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="20" pageNumber="129">Figs 10</figureCitation>
|
||
<figureCitation id="42B0D46402B131A3F0426055A54D2F24" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="20" pageNumber="129">, 11</figureCitation>
|
||
<figureCitation id="40A8BBBCF91763933C24D7BB4E8BA367" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="20" pageNumber="129">, 12</figureCitation>
|
||
<figureCitation id="5ABC39DE53F33A28CECFA8C0E5B785BC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="20" pageNumber="129">, 13</figureCitation>
|
||
<figureCitation id="2A4374F14E9010E65F353F777A3936C0" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="20" pageNumber="129">, 14</figureCitation>
|
||
<figureCitation id="D4CE95B3B1C8CCCEAFB0B4792444DD8F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of Primocerus spp." figureDoi="10.3897/zookeys.855.33013.figure15" httpUri="https://binary.pensoft.net/fig/310072" pageId="20" pageNumber="129">, 15</figureCitation>
|
||
<figureCitation id="E0AC75E1804611B0FD2D0F3EAF119372" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Habitat of Primocerus spp. A habitat and type locality for P. cuspidis, Venezuela, Tobogan de la Selva, collecting event AS- 08 - 080 b B habitat and type locality for P. pijiguaense, Venezuela, Los Pijiguaos, collecting event AS- 07 - 015 C habitat and type locality for P. neutrum, Venezuela, along La Escalera, collecting event AS- 08 - 058 D habitat and type locality for P. petilus, Brazil, Vale do Paraiso, collecting event BR 18 - 0203 - 01 G." figureDoi="10.3897/zookeys.855.33013.figure16" httpUri="https://binary.pensoft.net/fig/310073" pageId="20" pageNumber="129">, 16</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="541CC444ED5CB9442F30C83A8DD5E1EB" pageId="21" pageNumber="130" type="type species">
|
||
<paragraph id="0612DF406C82C344B99FEFA46CD3BC77" pageId="21" pageNumber="130">
|
||
<pageBreakToken id="1CE2D05409EDE3D092CCBB70ED47376A" pageId="21" pageNumber="130" start="start">Type</pageBreakToken>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="64A57918B4C595713724DA167402C230" pageId="21" pageNumber="130">
|
||
<taxonomicName id="096EC10C95869D5896794851C21CAB1A" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus neutrum" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="neutrum">
|
||
<emphasis id="765C3A3907C147DA45E0FB316F6E3825" italics="true" pageId="21" pageNumber="130">Primocerus neutrum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9816D080B915837537D75CF76F97F2B8" lastPageId="24" lastPageNumber="133" pageId="21" pageNumber="130" type="differential diagnosis">
|
||
<paragraph id="FFADC1BBE85E00AB830BFB713B605F43" pageId="21" pageNumber="130">Differential diagnosis.</paragraph>
|
||
<paragraph id="65E8FC82C3F7F76C7319C861FFA81C55" pageId="21" pageNumber="130">
|
||
Small to medium sized beetles (2.4-4.9 mm), elongate oval in dorsal view, moderate to strongly convex in lateral view (e.g., Figs
|
||
<figureCitation id="E09E3805697C4B15173089AA21D5FC68" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="21" pageNumber="130">11F</figureCitation>
|
||
,
|
||
<figureCitation id="0CFFC06D840D426F59C4FBEB207FA6FE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="21" pageNumber="130">12B</figureCitation>
|
||
), brown, dark brown, reddish brown, or rather orange. Antennae with eight antennomeres (e.g., Fig.
|
||
<figureCitation id="349827A5DBB9C17B5B4B3E72AE06D975" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="21" pageNumber="130">10G</figureCitation>
|
||
). Maxillary palps short to moderately long (e.g., shorter to nearly as long as the width of the head; e.g., Figs
|
||
<figureCitation id="2EA314B12528451CEF377334D194B32F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="21" pageNumber="130">10H</figureCitation>
|
||
,
|
||
<figureCitation id="D8A03B4E8D0EF4720FBD72A2BED95EDD" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="21" pageNumber="130">12H</figureCitation>
|
||
). Elytra with sutural striae; elytral punctures from shallow to sharply marked (e.g., Figs
|
||
<figureCitation id="3AF26FD850C682BA818070820C01BA1F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="21" pageNumber="130">11E</figureCitation>
|
||
,
|
||
<figureCitation id="467AE8361B309BA77DF121D4BD2CAC9A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="21" pageNumber="130">12E</figureCitation>
|
||
); serial punctures, ground punctures and systematic punctures similar in size and degree of impression throughout elytra; all punctures seemingly arranged in rows, sometimes evidently so. Prosternum flat to mesally only slightly produced. Posterior elevation of mesoventrite with a curved transverse ridge, rather sharp and low (Fig.
|
||
<figureCitation id="0CF2E1F4FF17DAB767BAAD240946751A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="21" pageNumber="130">14A</figureCitation>
|
||
), except in
|
||
<taxonomicName id="BCCBE460188B2C726E90932180921BCA" lsidName="P. cuspidis" pageId="21" pageNumber="130" rank="species" species="cuspidis">
|
||
<emphasis id="12FAFA0A07B11C46DDD9C6CC8F7CA5B4" italics="true" pageId="21" pageNumber="130">P. cuspidis</emphasis>
|
||
</taxonomicName>
|
||
which bears a sharp, pyramidal (triangular) projection. Posteromesal glabrous patch on metaventrite nearly as wide as long. Pubescence on anterior surface of metafemora ranging from sparse to densely covering basal three fourths of the femur (e.g., Figs
|
||
<figureCitation id="F985187B4EBF38898048110FA1650121" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="21" pageNumber="130">12C</figureCitation>
|
||
,
|
||
<figureCitation id="43787D690F741536CF2A49CCCB47A06B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="21" pageNumber="130">10G</figureCitation>
|
||
,
|
||
<figureCitation id="B667B05DDFA4D5A7C4E24EEEDA481A05" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="21" pageNumber="130">11G</figureCitation>
|
||
). Fifth abdominal ventrite apically rounded, truncate or slightly emarginate, usually with stout setae (e.g., Fig.
|
||
<figureCitation id="B641EF1548951EA534BCE761ED36FB7B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="21" pageNumber="130">14C</figureCitation>
|
||
). Basal piece of aedeagus as long as or longer than parameres; median lobe triangular, nearly as wide at base as basal width of one paramere, with apical projection (Fig.
|
||
<figureCitation id="8143401DF3F986523B9537B4F746517A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="21" pageNumber="130">
|
||
14
|
||
<normalizedToken id="2AB57C0D1DFD25989C4C3232E27AA9BC" originalValue="D–L">D-L</normalizedToken>
|
||
</figureCitation>
|
||
); gonopore absent (Fig.
|
||
<figureCitation id="6D92BEB62FA94C3A7C3BE9FF9D446585" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="21" pageNumber="130">
|
||
14
|
||
<normalizedToken id="D9354E888657F37112E72C57AFA98811" originalValue="D–L">D-L</normalizedToken>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B4A45A6D6B392754427706655B1626D5" ID-Zenodo-Dep="3251381" doi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="21" pageNumber="130" start="Figure 10" startId="F10">
|
||
<paragraph id="FE106074E3BF6122A92A15A1BC50E1CE" pageId="21" pageNumber="130">
|
||
<emphasis id="949776517B578CD92DEC4C31B1499854" bold="true" pageId="21" pageNumber="130">Figure 10.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="E1A1A82D5984604BB9AD8D1E748C7C1D" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1F76D5107EBA849BED48FF85EE0882B5" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="227EB413DDCB4977D7F8AC00B279A2AD" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="991CBB63F3A4164EAF17272E2689250D" originalValue="A–D">A-D</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="1DFA48E59138C839BBA81BCD4CE68511" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus ocellatus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="ocellatus">
|
||
<emphasis id="AF70CD5A3F7F5B1CC321AE2738C39743" italics="true" pageId="21" pageNumber="130">Primocerus ocellatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="6A788C1E1793AA4833ADF3A3711495AF" bold="true" pageId="21" pageNumber="130">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="0598F0BF3897FB6A11356B936B3B30D7" bold="true" pageId="21" pageNumber="130">B</emphasis>
|
||
lateral view
|
||
<emphasis id="447F929EC9CE80B656C7A12D4B44E818" bold="true" pageId="21" pageNumber="130">C</emphasis>
|
||
ventral view
|
||
<emphasis id="5F62416C2C78D0172A643B7F3067BA8C" bold="true" pageId="21" pageNumber="130">D</emphasis>
|
||
head, dorsal view.
|
||
<emphasis id="9AAA2B747B5D86BDBE02B2B7E5B00211" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="AC1E2361995DE879208FB459DC7E3AFF" originalValue="E–H">E-H</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="486A8826B74309588E6E02AD023CF075" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus gigas" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="D50E2B8D9D99465915F775B117D4F52F" italics="true" pageId="21" pageNumber="130">Primocerus gigas</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="333C55532BD6032A5E90E7948C2FCC41" bold="true" pageId="21" pageNumber="130">E</emphasis>
|
||
dorsal view
|
||
<emphasis id="4395C4B0B3C0C34ED3A4C775D4938FA6" bold="true" pageId="21" pageNumber="130">F</emphasis>
|
||
lateral view
|
||
<emphasis id="E9A03B92C8675EFB8542906D13AD2CDE" bold="true" pageId="21" pageNumber="130">G</emphasis>
|
||
ventral view
|
||
<emphasis id="7EA05ACC64E2C4C8D5E4FB22A112F264" bold="true" pageId="21" pageNumber="130">H</emphasis>
|
||
head, dorsal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5704D5180592E059F2789FB63763748F" ID-Zenodo-Dep="3251383" doi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="21" pageNumber="130" start="Figure 11" startId="F11">
|
||
<paragraph id="C4C0283D31F219424C1987B71FF4A389" pageId="21" pageNumber="130">
|
||
<emphasis id="DE62FCC8F07BDF8DC8761EE12C0C6D7E" bold="true" pageId="21" pageNumber="130">Figure 11.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4EF3822E20604A202A774F2F75FFDF96" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B8A5C665DFC04E324A7254ABDAB065F6" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="D319C41E24568593614471B88F753435" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="AE646228A7690316A10E83937757C207" originalValue="A–D">A-D</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="018C1625B59DA3DF55ADF4B5CD6BFAB9" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus cuspidis" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="cuspidis">
|
||
<emphasis id="644E2CA2C3AE2C1421FDF554C6A01284" italics="true" pageId="21" pageNumber="130">Primocerus cuspidis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="8D9B77AB6AA4D831360D51FDBA0026E2" bold="true" pageId="21" pageNumber="130">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="18A33252CA11FA6783C3D9B1D034C5B6" bold="true" pageId="21" pageNumber="130">B</emphasis>
|
||
lateral view
|
||
<emphasis id="D4B449658597A4BFA5187384AD4633E7" bold="true" pageId="21" pageNumber="130">C</emphasis>
|
||
ventral view
|
||
<emphasis id="E5287115869BCE34E8A78D3F04B2262C" bold="true" pageId="21" pageNumber="130">D</emphasis>
|
||
head, dorsal view.
|
||
<emphasis id="0D8FEFE8D6A26CEA1AF44BF3160526CB" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="15E4B5D94F84A312B3A8A545C73C25A5" originalValue="E–H">E-H</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="73F6B086E36A58157B922AA59680F0B1" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus neutrum" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="neutrum">
|
||
<emphasis id="910C91B28027834642932DA9D5F5503E" italics="true" pageId="21" pageNumber="130">Primocerus neutrum</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="9EE771819BBABA27D62C83C44B85D8E2" bold="true" pageId="21" pageNumber="130">E</emphasis>
|
||
dorsal view
|
||
<emphasis id="157465CBA98F4F823F8B2634A7EF6B3A" bold="true" pageId="21" pageNumber="130">F</emphasis>
|
||
lateral view
|
||
<emphasis id="37DA33269263AE5BC26E820FDF6327AC" bold="true" pageId="21" pageNumber="130">G</emphasis>
|
||
ventral view
|
||
<emphasis id="3459DF83C51A893E8E22F89BF82209E5" bold="true" pageId="21" pageNumber="130">H</emphasis>
|
||
head, dorsal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AE4FB9174BA19FC53203054658F93581" ID-Zenodo-Dep="3251385" doi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="21" pageNumber="130" start="Figure 12" startId="F12">
|
||
<paragraph id="6B144D9BA0053EF5057F019C941B1C41" pageId="21" pageNumber="130">
|
||
<emphasis id="DDE3BE3507B0105397BDA39049AE8DC0" bold="true" pageId="21" pageNumber="130">Figure 12.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="8B44921ABC0928C703D790482EEFB8F2" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9C88507054182AF153EE73B9A8FAF275" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="09337BEF10F3B6E91DAA2AA8B3D12123" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="84D301E33B03688985A3DACF877D2F32" originalValue="A–D">A-D</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="FAE3A167526C740EB7E871D5F7B5A907" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus maipure" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="maipure">
|
||
<emphasis id="0A01D0905036CCF3D0D196210448E911" italics="true" pageId="21" pageNumber="130">Primocerus maipure</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="E8F8DDB5713319890D99B25D9897B6A8" bold="true" pageId="21" pageNumber="130">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="340972F60A41D8346A515A147BE74629" bold="true" pageId="21" pageNumber="130">B</emphasis>
|
||
lateral view
|
||
<emphasis id="84E2CE84F7A47E835C20F3B9FFB469C8" bold="true" pageId="21" pageNumber="130">C</emphasis>
|
||
ventral view
|
||
<emphasis id="D46BA438FAE40468FCB027389BD4F2CD" bold="true" pageId="21" pageNumber="130">D</emphasis>
|
||
head, dorsal view.
|
||
<emphasis id="F67A209CC411061C337CF2D3C57DC05F" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="DC2FA2012DE3F59A7DE7C983232ACA53" originalValue="E–H">E-H</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="F8AF1AB71E1810226701137821A5E8AB" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus pijiguaense" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="pijiguaense">
|
||
<emphasis id="6CF4B1704727C5A9B23281709A255F67" italics="true" pageId="21" pageNumber="130">Primocerus pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="D40726E8DC37BF8A43B63A4B62DBE501" bold="true" pageId="21" pageNumber="130">E</emphasis>
|
||
dorsal view
|
||
<emphasis id="0956F3347E300AA2FCA72A2A22A94172" bold="true" pageId="21" pageNumber="130">F</emphasis>
|
||
lateral view
|
||
<emphasis id="A9309237C16BEF45C063A7DFD40BE8F0" bold="true" pageId="21" pageNumber="130">G</emphasis>
|
||
ventral view
|
||
<emphasis id="9792E3EC2907FD39B510E2B7BFB916F2" bold="true" pageId="21" pageNumber="130">H</emphasis>
|
||
head, dorsal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D4F5D02D9B876AA7D9BA5957CB7B2482" ID-Zenodo-Dep="3251387" doi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="21" pageNumber="130" start="Figure 13" startId="F13">
|
||
<paragraph id="8F9687DB210609E75706075696EAD0E7" pageId="21" pageNumber="130">
|
||
<emphasis id="DE4F65518F773D0A7391D15605138B9A" bold="true" pageId="21" pageNumber="130">Figure 13.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="DA9D1C0037A9F0CBA9F553F99C3FB90B" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="55E613E97DD212AC5C1B5BDB2B85D67B" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="C2E69FCED792FC15F36B76EC9923701A" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="3B564275D43031B5C7FF25561A87085E" originalValue="A–C">A-C</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="4DAF2A6A050C124E985603AE53589A9C" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus petilus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="petilus">
|
||
<emphasis id="003553B6B4F52D77F62B51CAB529FB40" italics="true" pageId="21" pageNumber="130">Primocerus petilus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="3C0C1AD7855D51A4FEFFB340747E5E16" bold="true" pageId="21" pageNumber="130">A</emphasis>
|
||
dorsal view
|
||
<emphasis id="0A0256E027D5D8AFE2B483985FA363CF" bold="true" pageId="21" pageNumber="130">B</emphasis>
|
||
lateral view
|
||
<emphasis id="120AC62F975C31079CDAAC9AED1EC130" bold="true" pageId="21" pageNumber="130">C</emphasis>
|
||
ventral view.
|
||
<emphasis id="5FE5C0C97B790BD4557E431F45120966" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="59B2726CF6C1365AE7B22844E64E07B4" originalValue="D–F">D-F</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="4D57D8476302DF866D83828C46319B6A" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus striatolatus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="striatolatus">
|
||
<emphasis id="37AD7C677C75BCF26C5B96432B6B3F50" italics="true" pageId="21" pageNumber="130">Primocerus striatolatus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="CF93B80A2132CE15711EA62F4F892021" bold="true" pageId="21" pageNumber="130">D</emphasis>
|
||
dorsal view
|
||
<emphasis id="B640BFFBAB7FD0CCC78D792A2F7F64A2" bold="true" pageId="21" pageNumber="130">E</emphasis>
|
||
lateral view
|
||
<emphasis id="25A0BBF0BD4663A3CDF3EB66738AAC01" bold="true" pageId="21" pageNumber="130">F</emphasis>
|
||
ventral view.
|
||
<emphasis id="D8B9E923BF99CC24269629E7AC5C7EB6" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="E780E5A394482A4AA232EB6975CBA40E" originalValue="G–I">G-I</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="FC114F73328E24896509D30319408A11" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus semipubescens" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="semipubescens">
|
||
<emphasis id="F8E12BDF6420B4F7D56002D030D2F4FB" italics="true" pageId="21" pageNumber="130">Primocerus semipubescens</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="F0E82773DC646BFA638A106BBD4930F7" bold="true" pageId="21" pageNumber="130">G</emphasis>
|
||
dorsal view
|
||
<emphasis id="09659F28345F015DD9D626CB0253DF70" bold="true" pageId="21" pageNumber="130">H</emphasis>
|
||
lateral view
|
||
<emphasis id="6683B23F6BA7178464DE0A89DEEEC8C9" bold="true" pageId="21" pageNumber="130">I</emphasis>
|
||
ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6AFBD1EEBFAF6138E2CD4B15C17C7DD1" ID-Zenodo-Dep="3251389" doi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="21" pageNumber="130" start="Figure 14" startId="F14">
|
||
<paragraph id="BA001B15D7098EB737F174EA9E132E54" pageId="21" pageNumber="130">
|
||
<emphasis id="F1A01BB79062758F2F9EB648BDE4B611" bold="true" pageId="21" pageNumber="130">Figure 14.</emphasis>
|
||
Thorax, abdomen and aedeagus of
|
||
<taxonomicName id="18DE2E0065C13501DBF8C0CF47BD8F6A" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0965224DF7DAFC813DAD3046B3896B15" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.:
|
||
<emphasis id="2C4FFF385D47E0C2D19E60CBA1F84FB9" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="E2A09A349BB1902FC9D676CBA4C814BB" originalValue="A–D">A-D</normalizedToken>
|
||
</emphasis>
|
||
<taxonomicName id="14731C2BF8D27708CC2497931F8BA33D" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus neutrum" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="neutrum">
|
||
<emphasis id="889E6751DE193F60D58446F085FA988D" italics="true" pageId="21" pageNumber="130">Primocerus neutrum</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="8F18CE7904156302B842E38493F01116" bold="true" pageId="21" pageNumber="130">A</emphasis>
|
||
ventral view of mesoventrite (white arrow pointing transverse ridge)
|
||
<emphasis id="A52199CE3C558D271274874FB62C962C" bold="true" pageId="21" pageNumber="130">B</emphasis>
|
||
posterior view of metafurca
|
||
<emphasis id="9264951507BF220D2220FB4A239927AD" bold="true" pageId="21" pageNumber="130">C</emphasis>
|
||
fifth abdominal ventrite
|
||
<emphasis id="D4000E59B0D100AF882C9056B5400377" bold="true" pageId="21" pageNumber="130">D</emphasis>
|
||
aedeagus.
|
||
<emphasis id="B36BB3FF39A725CCAAEF962D887EEBD2" bold="true" pageId="21" pageNumber="130">
|
||
<normalizedToken id="0C84579E44DECDBAB9739CDE464ADDDC" originalValue="E–L">E-L</normalizedToken>
|
||
</emphasis>
|
||
aedeagus:
|
||
<emphasis id="51257C09395F2BBF646C77B367DB76CA" bold="true" pageId="21" pageNumber="130">E, F</emphasis>
|
||
<taxonomicName id="77293DE8F3178D27F425A5112B787E46" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus maipure" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="maipure">
|
||
<emphasis id="FA5F4EBDC45E5A1ADA193E0B5BDD60B8" italics="true" pageId="21" pageNumber="130">Primocerus maipure</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="E4D91E572C6750F6DEC24BADDD52623D" bold="true" pageId="21" pageNumber="130">E</emphasis>
|
||
dorsal view
|
||
<emphasis id="08E07456E29FEE6EF3432DBE4D3BBF60" bold="true" pageId="21" pageNumber="130">F</emphasis>
|
||
lateral view.
|
||
<emphasis id="F1306A08AE091578AA8A28BCC4AFCEDF" bold="true" pageId="21" pageNumber="130">G, H</emphasis>
|
||
<taxonomicName id="AD033E4BEFBEFE477DFF83830D65ACE4" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus pijiguaense" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="pijiguaense">
|
||
<emphasis id="E8C547D991A8CC5AC0F47BB55C4913A5" italics="true" pageId="21" pageNumber="130">Primocerus pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="641A527AA77D52C56EE14B52B791BA76" bold="true" pageId="21" pageNumber="130">G</emphasis>
|
||
dorsal view
|
||
<emphasis id="E8366C64BBE93F48E321D75836918D7A" bold="true" pageId="21" pageNumber="130">H</emphasis>
|
||
lateral view
|
||
<emphasis id="2DFAE8D4489B5321605D7AB0B273A321" bold="true" pageId="21" pageNumber="130">I</emphasis>
|
||
<taxonomicName id="3289A019C544B0AE7A306C8BC16A2026" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus gigas" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="DE41724ED6E8417737612C400C65B601" italics="true" pageId="21" pageNumber="130">Primocerus gigas</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1BA12D4481CCF3ABF76EC518928E51B4" bold="true" pageId="21" pageNumber="130">J</emphasis>
|
||
<taxonomicName id="E868E5ECBAD89BAF33CCE230F9C4ED5F" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus petilus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="petilus">
|
||
<emphasis id="80308330149EBDB62136427EE374BA72" italics="true" pageId="21" pageNumber="130">Primocerus petilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5E7E1186A54E7DF7DEF643158E9A3774" bold="true" pageId="21" pageNumber="130">K</emphasis>
|
||
<taxonomicName id="4DE51852DFA387FA3B0EBD476A210DA7" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus striatolatus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="striatolatus">
|
||
<emphasis id="D9639E30D1FFD0653457A81B2485A8E3" italics="true" pageId="21" pageNumber="130">Primocerus striatolatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3DE44F4A075513FABC0171920A861CC8" bold="true" pageId="21" pageNumber="130">L</emphasis>
|
||
<taxonomicName id="B2491E09544A2F43D15BBC7C9799932C" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus cuspidis." order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="species" species="cuspidis.">
|
||
<emphasis id="85CC70BBCBA2329650A13650ABA8414B" italics="true" pageId="21" pageNumber="130">Primocerus cuspidis.</emphasis>
|
||
</taxonomicName>
|
||
Scale bars: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0C3AAD40259FF23945C43AE8AC656B59" lastPageId="22" lastPageNumber="131" pageId="21" pageNumber="130">
|
||
At first sight, the dorsally smoother members of
|
||
<taxonomicName id="AF241D387F21006BC487E6AB49CB6A1F" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9E95242DBFA1B88DB82EA72B72F3F559" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="05160AFD83232D0ABA9E237E9D618833" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="21" pageNumber="130">10</figureCitation>
|
||
,
|
||
<figureCitation id="CB8967C2EE47D8C66C7A698CFFE4C825" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="21" pageNumber="130">11</figureCitation>
|
||
) can be mistaken for
|
||
<taxonomicName id="660FDD00978C9BD410EDF53D0E3E04D4" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C28F219067881ADE77074ED45213378D" italics="true" pageId="21" pageNumber="130">Chasmogenus</emphasis>
|
||
</taxonomicName>
|
||
, given that both genera exhibit sutural striae. The presence of a transverse curved ridge (sometimes very low) on the posterior elevation of the mesoventrite distinguishes
|
||
<taxonomicName id="3867A16577288811F85B1C62E1260FE7" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBABFC4ABB2365A8D6DEF9F1A98E0090" italics="true" pageId="21" pageNumber="130">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="3AF84DF644F261F4338A93AA3E06FCFE" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DC43ADF3563D9E542AC5F1B572509876" italics="true" pageId="21" pageNumber="130">Chasmogenus</emphasis>
|
||
</taxonomicName>
|
||
, in which the mesoventrite is either flat, broadly elevated or with a longitudinal elevation (e.g., figs 2, 4 in Clarkson and Ferreira 2014). In addition, the maxillary palps of most
|
||
<taxonomicName id="F9B49F0F8B12CB67F9BF757A8ADED30A" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="21" pageNumber="130" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F8474CEA4F459A7D410A55FC7D6A9F01" italics="true" pageId="21" pageNumber="130">Chasmogenus</emphasis>
|
||
</taxonomicName>
|
||
species are
|
||
<pageBreakToken id="432B62DFFFFCEB1401824F572A84392E" pageId="22" pageNumber="131" start="start">nearly</pageBreakToken>
|
||
1.5
|
||
<normalizedToken id="EE293C0244ADF0308C42F6B6A12BB987" originalValue="×">x</normalizedToken>
|
||
longer than the maximum width of the head, whereas in
|
||
<taxonomicName id="0AC66CF498E69F083859236836134295" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="22" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="06D65B26C34D0464A8F8C279DF61B5F9" italics="true" pageId="22" pageNumber="131">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
the maxillary palps are nearly as long as the width of the head.
|
||
</paragraph>
|
||
<paragraph id="234CCEA6D293D100903342913A5B988D" lastPageId="23" lastPageNumber="132" pageId="22" pageNumber="131">
|
||
Punctate members of
|
||
<taxonomicName id="258EC5D276FFC889F39CBDAD3EE85FCB" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="22" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4549A7EDA2CD7D6458CA06E311C4F9F8" italics="true" pageId="22" pageNumber="131">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
(in particular
|
||
<taxonomicName id="960591DEFB557E6853F2B56EF4B447C8" lsidName="P. maipure" pageId="22" pageNumber="131" rank="species" species="maipure">
|
||
<emphasis id="79A0321FCBE69DC6A90E6DA9B9653001" italics="true" pageId="22" pageNumber="131">P. maipure</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60B2682416379478161100002C0F3ADB" lsidName="P. pijiguaense" pageId="22" pageNumber="131" rank="species" species="pijiguaense">
|
||
<emphasis id="DB573A9D692183DF94905DF9DBEA58CB" italics="true" pageId="22" pageNumber="131">P. pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="B0F961594DA6EC8B4F7381BA21EF40CB" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="22" pageNumber="131">12</figureCitation>
|
||
) may resemble some species of
|
||
<taxonomicName id="29EE6095249D69B420D421A57B0DAD93" authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="22" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EF32D7E6A34A1D65D5A45318D4DA6B19" italics="true" pageId="22" pageNumber="131">Tobochares</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="22F46F223AE05E7F8C4D17D6D86BE72F" lsidName="T. canthus" pageId="22" pageNumber="131" rank="species" species="canthus">
|
||
<emphasis id="D95783C30EEFF3C3BD4E05B63F76BAA2" italics="true" pageId="22" pageNumber="131">T. canthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D370289F621422B16AB3D1C531F9906E" lsidName="T. pallidus" pageId="22" pageNumber="131" rank="species" species="pallidus">
|
||
<emphasis id="0B84B4920287E801CC3B5143D1DCC898" italics="true" pageId="22" pageNumber="131">T. pallidus</emphasis>
|
||
</taxonomicName>
|
||
; Kohenberg
|
||
<pageBreakToken id="4A94A9F25752292B0CDDB2F9850031B5" pageId="23" pageNumber="132" start="start">and</pageBreakToken>
|
||
Short 2017); striate
|
||
<taxonomicName id="528C3A3528FAB82A962C83E2F460FD6E" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BF6B30331670C162CA7AC3088F0F60C3" italics="true" pageId="23" pageNumber="132">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
may resemble a very small
|
||
<taxonomicName id="576987C709C24B3D5EA8B241D8376FA5" class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C33505CF74F21F060459B737D6DF6DEF" italics="true" pageId="23" pageNumber="132">Radicitus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="CA2F951F317D607D1EAD0FB6C8E7E34E" DOI="https://doi.org/10.11646/zootaxa.3835.2.5" author="Short, AEZ" journalOrPublisher="Zootaxa" pageId="41" pageNumber="150" pagination="251 - 262" refId="B13" refString="Short, AEZ, Garcia, M, 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 3835 (2): 251 - 262, DOI: https://doi.org/10.11646/zootaxa.3835.2.5" title="A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae)." url="https://doi.org/10.11646/zootaxa.3835.2.5" volume="3835" year="2014">
|
||
Short and
|
||
<normalizedToken id="172D4C8D5958F36E5F5995F07B805DE9" originalValue="García">Garcia</normalizedToken>
|
||
2014
|
||
</bibRefCitation>
|
||
). In those cases,
|
||
<taxonomicName id="189AD2CC8331A9DAA04B8029D979E5A3" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="222FA05A4959045546F43185449E403C" italics="true" pageId="23" pageNumber="132">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
can be easily recognized by the presence of sutural striae. Some
|
||
<taxonomicName id="79CDD38FB1B87BDA9FB7558216E13B44" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B257D77694A660861ACFA4A3B078EB68" italics="true" pageId="23" pageNumber="132">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
may also superficially resemble certain New World cylomine genera, such as
|
||
<taxonomicName id="FB3C9EBCB31142366DD58D8239CD25E0" authorityName="Spangler" authorityYear="1979" class="Insecta" family="Hydrophilidae" genus="Andotypus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andotypus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7A7EF30701C75CD84424DDA968F0A7B2" italics="true" pageId="23" pageNumber="132">Andotypus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<normalizedToken id="84F23E1EEC2A105E29C1414C11D84553" originalValue="Fikáček">Fikacek</normalizedToken>
|
||
et al. 2014), from which it may be distinguished by the fully exposed labrum of
|
||
<taxonomicName id="9E46DD33C811B1FC0865FF06225F8C26" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="329FA0A0824F6B96A341882D1BC0CD7F" italics="true" pageId="23" pageNumber="132">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="67021854C58FD446D5C72DF79287A381" lastPageId="24" lastPageNumber="133" pageId="23" pageNumber="132">
|
||
In addition, the presence of sutural striae and the relative size of the basal piece of the aedeagus resemble some species of
|
||
<taxonomicName id="6EA87A50C8FE46476412464C0F929226" authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus" order="Coleoptera" pageId="23" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="474AE24CE488F86F88A3255CDA72ACD5" italics="true" pageId="23" pageNumber="132">Enochrus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="64A929292FBEA708D882F0DC508C60E8" authorityName="Short & Fikacek" authorityYear="2013" lsidName="" pageId="23" pageNumber="132" rank="subFamily" subFamily="Enochrinae">Enochrinae</taxonomicName>
|
||
) in that the basal piece is as long as or longer than the median lobe + parameres (e.g. see figs 11 and 14 in
|
||
<bibRefCitation id="1BC6BB3AD12BF5075BE4A2EB9CEB56AB" DOI="https://doi.org/10.3157/0002-8320(2006)132[279:TNSOET]2.0.CO;2" author="Fernandez, LA" journalOrPublisher="Insect Systematics & Evolution" pageId="40" pageNumber="149" refId="B7" refString="Fernandez, LA, 2006. Two new species of Enochrus Thomson (Coleoptera, Hydrophilidae) from the Neotropical Region. Transactions of the American Entomological Society 132(3): 279-284. https://doi.org/10.3157/0002-8320(2006)132[279:TNSOET]2.0.CO;2" title="Two new species of Enochrus Thomson (Coleoptera, Hydrophilidae) from the Neotropical Region. Transactions of the American Entomological Society 132 (3): 279 - 284." url="https://doi.org/10.3157/0002-8320(2006)132[279:TNSOET]2.0.CO;2" year="2006">
|
||
<pageBreakToken id="6B3FB5C1C1AEC7CB043EB90615A3E5A5" pageId="24" pageNumber="133" start="start">
|
||
<normalizedToken id="5ED9EB1AC584D7E8F8135102974ADD44" originalValue="Fernández">Fernandez</normalizedToken>
|
||
</pageBreakToken>
|
||
2006
|
||
</bibRefCitation>
|
||
). The maxillary palps curved inwards in
|
||
<taxonomicName id="4C92CFE3B965F0ED9312835DA1138876" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="24" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BA3B2168F21863B740CD1446CCEB1A28" italics="true" pageId="24" pageNumber="133">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
(as opposed to zig-zag-like as in
|
||
<taxonomicName id="E25FCB0D93E0A74ED72BDECA2C2F908C" authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Hydrophilidae" genus="Enochrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enochrus" order="Coleoptera" pageId="24" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9FEA8E10E953101F9FD9E170F3D5BB58" italics="true" pageId="24" pageNumber="133">Enochrus</emphasis>
|
||
</taxonomicName>
|
||
) allows for its recognition.
|
||
</paragraph>
|
||
<paragraph id="873902BDBD06ACD186CDBC4F1B0965D2" pageId="24" pageNumber="133">
|
||
The aedeagus of
|
||
<taxonomicName id="EDF768CD78EEEF20DE4AA2078778C7EB" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="24" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A4EF45AFF37E7E98BEEF471CB3D73E93" italics="true" pageId="24" pageNumber="133">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
is so far unique among the
|
||
<taxonomicName id="C04FC73C61B4FEE2EF185059C9CA18F4" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" lsidName="" pageId="24" pageNumber="133" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
|
||
in the lack of a well-developed gonopore, and the presence of a lightly sclerotized projection beyond the apex of the median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="666F9601584E2D357D73ED7AA8EAB0D4" lastPageId="28" lastPageNumber="137" pageId="24" pageNumber="133" type="description">
|
||
<paragraph id="A8D8C8A6C8617B18D555B6A9BA4C1F27" pageId="24" pageNumber="133">Description.</paragraph>
|
||
<paragraph id="EB6A83BE8942C2782810E9B01AFA8EC6" lastPageId="28" lastPageNumber="137" pageId="24" pageNumber="133">
|
||
Small to medium sized beetles, total body length 2.4-4.9 mm; body elongate oval, moderate to strongly convex in lateral view (e.g., Figs
|
||
<figureCitation id="8879DDA545068C24160E1619CDAE5FA8" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="24" pageNumber="133">11F</figureCitation>
|
||
,
|
||
<figureCitation id="FAA7AF2BCB50D90918BBAA6CBC6CB71F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="24" pageNumber="133">12B</figureCitation>
|
||
); orange brown (Fig.
|
||
<figureCitation id="054B4ED602F9F69687C44EF139553A81" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="24" pageNumber="133">
|
||
11
|
||
<normalizedToken id="2B7203F9293B02F405A44570F2B3E755" originalValue="A–D">A-D</normalizedToken>
|
||
</figureCitation>
|
||
), reddish brown (Fig.
|
||
<figureCitation id="C75F17FCF4BDC1303B2817855AFECA07" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">
|
||
10
|
||
<normalizedToken id="C80061A7F89E84C318D2908BFFB7E0D2" originalValue="A–D">A-D</normalizedToken>
|
||
</figureCitation>
|
||
), to dark brown in coloration (e.g., Fig.
|
||
<figureCitation id="EBB4A2F4E4A984F5C084683101D15B90" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="24" pageNumber="133">13</figureCitation>
|
||
), usually uniform along body regions, sometimes slightly paler along margins, pronotum, ventral surfaces, and appendages, particularly maxillary palps and tarsi.
|
||
<emphasis id="EC50BBEEFC632C6324829186F4531D45" bold="true" italics="true" pageId="24" pageNumber="133">Head.</emphasis>
|
||
Frons and clypeus with either shallow (e.g., Fig.
|
||
<figureCitation id="A38B582271D9A83BAB60C1804EFDC57F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">10D</figureCitation>
|
||
) or sharply marked (e.g., Fig.
|
||
<figureCitation id="2592CB15CB4DD152B676307F03E19DC7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="24" pageNumber="133">12D</figureCitation>
|
||
) ground punctures, irregularly and rather densely distributed over the surface, accompanied by scattered seta-bearing systematic punctures, particularly noticeable along anterior and inner margins of eyes, and lateral areas of clypeus; surface between punctures smooth and shiny. Clypeus roughly trapezoid, with posterior margin wider than anterior margin; anterior corners roundly angulated, anterior margin widely roundly emarginate; membranous preclypeal area not visible (visible in
|
||
<taxonomicName id="4FF34A279B41BEC0862FA55C9C6BD690" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="24" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F873A950DB4A2E784F2E93E2748A2773" italics="true" pageId="24" pageNumber="133">Chasmogenus</emphasis>
|
||
</taxonomicName>
|
||
; e.g., fig. 28 in Clarkson and Ferreira 2014); surface mesally moderately convex, laterally flattened to slightly concave (Figs
|
||
<figureCitation id="21ABB41FBC3C80C60EE7C6B1FE2435B2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="24" pageNumber="133">12D, H</figureCitation>
|
||
). Eyes subquadrate in dorsal view, usually protruding from outer outline of head. Labrum wide, fully exposed, collinear to perpendicular to clypeus, and usually around 0.3 times as long as clypeus (e.g., Fig.
|
||
<figureCitation id="152974BC3D63F439BAAE99C8002F9265" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">10D</figureCitation>
|
||
); dorsal surface flat to convex, with scattered fine punctures; anterior margin markedly roundly bent inwards, mesally emarginate, with tiny denticles along emargination, and setae on lateral areas of anterior margin. Temporae densely covered by very short and fine setae (hydrofuge pubescence). Mentum parallel sided, often with lateral margins densely fringed by short setae; surface rather flat, smooth, and glabrous, sometimes with lateral oblique longitudinal ridges, and few crenulations; anterior margin with wide, deep, concave median impression, sometimes marked by a transverse carina. Submentum sunken, concave, and pubescent at base, glabrous, shiny, flat and ascending at apex; ocular ridge of variable development. Maxilla (e.g., Fig.
|
||
<figureCitation id="AF9865EFAEE121A6EF34B5084B55AD5E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">10G</figureCitation>
|
||
) with ventral surface of cardo and stipes smooth, shiny, and glabrous; outer dorsal margin of palpifer with a row of stiff, decumbent, spiniform setae; limit between cardo and stipes oblique; maxillary palps curved inward, brown to orange or yellow, longer than antennae, short to moderately long (e.g., shorter to nearly as long as the width of the head; e.g., Figs
|
||
<figureCitation id="7577CB50AEC58E315F000C076EB76D9B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">10H</figureCitation>
|
||
,
|
||
<figureCitation id="0673D5CDAACE11E16E0FDB998E03AFD7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="24" pageNumber="133">12H</figureCitation>
|
||
); maxillary palpomere 1 gradually broadening towards apex, with inner margin straight and outer margin apically convex; apex of palpomere 3 bearing sensilla; palpomeres 1 and 3 similar in length, palpomere 2 only slightly shorter. Mandibles with apex bifid (observed in
|
||
<taxonomicName id="0655AD6F8F15D994070ECCE418783CD1" lsidName="P. gigas" pageId="24" pageNumber="133" rank="species" species="gigas">
|
||
<emphasis id="5C684405FCC2BD36F914CB303259A49C" italics="true" pageId="24" pageNumber="133">P. gigas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8174BAA62767EFE1CF458456A2408100" lsidName="P. pijiguaense" pageId="24" pageNumber="133" rank="species" species="pijiguaense">
|
||
<emphasis id="5B443D19BAF85FA1D570F0976380187C" italics="true" pageId="24" pageNumber="133">P. pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="ECC0AC6FB7CC9489E311A88A7F64674B" lsidName="P. striatolatus" pageId="24" pageNumber="133" rank="species" species="striatolatus">
|
||
<emphasis id="86241EB4D0DDDD5291A8A4CD8BE7BF72" italics="true" pageId="24" pageNumber="133">P. striatolatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="795AB4E3B791C6A5A0B25D2B4EEB20D6" lsidName="P. petilus" pageId="24" pageNumber="133" rank="species" species="petilus">
|
||
<emphasis id="52011A0DEB5BA1C61952D7C9A3035E11" italics="true" pageId="24" pageNumber="133">P. petilus</emphasis>
|
||
</taxonomicName>
|
||
; e.g., Fig.
|
||
<figureCitation id="729C6D91D536E923A98F227DFBEA4138" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="24" pageNumber="133">12H</figureCitation>
|
||
). Labial palps yellowish to brown, usually nearly as long as mentum, dorsoventrally flattened; palpomere 2 with outer margin convex apicad of midpoint, sometimes with setae near apex; palpomere 3 digitiform to somewhat kidney-shaped, with one or two long subapical setae on outer margin. Antennae (e.g., Fig.
|
||
<figureCitation id="FDA43F46214839934885DC84CAE6D23A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="24" pageNumber="133">10G</figureCitation>
|
||
) with eight antennomeres, slightly paler than general coloration of head; antennomere 1 an
|
||
<pageBreakToken id="9CB28D824AB458BE3B8705ACDCCB9CA3" pageId="25" pageNumber="134" start="start">teriorly</pageBreakToken>
|
||
projected near base, at most reaching midpoint of ventral surface of eye, reaching to surpassing cardo-stipes joint, nearly 2.0
|
||
<normalizedToken id="4069D672D0F18ACA581CF35377DA9847" originalValue="×">x</normalizedToken>
|
||
longer than antennomere 2; antennomere 2 nearly as long as antennomeres 3-4 combined; antennomere 5 forming a well differentiated, symmetric cupule; antennomeres 6-8 slightly flattened, forming a loosely articulated, pubescent club (antennomere 7 shortest, 8 longest); apex of antennomere 8 with longer setae than general pubescence of club.
|
||
<emphasis id="875B873F693933DB102C597FA76CC6E8" bold="true" italics="true" pageId="25" pageNumber="134">Thorax.</emphasis>
|
||
Pronotum widest
|
||
<pageBreakToken id="2540A1F90F9E19FC471D1ADF5BF0BF3A" pageId="26" pageNumber="135" start="start">at</pageBreakToken>
|
||
base, narrowed anteriorly, surface evenly convex; anterior and posterior corners widely rounded, sometimes posterior corners almost forming a sharp straight angle (e.g., Fig.
|
||
<figureCitation id="FE45BD165280B33A20D011A7D628536E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="26" pageNumber="135">10G</figureCitation>
|
||
); anterior and posterior margins nearly straight; ground punctation either shallow or sharp, uniformly dense, with surface between punctures smooth and shiny; seta-bearing systematic punctures forming paired anterolateral semicircles. Scutellar shield of moderate size, triangular, nearly as long as wide, with punctation as in pronotum. Prosternum nearly as long as 0.7
|
||
<normalizedToken id="AF446BB4DD9177ABCF3A87B383D8F2CB" originalValue="×">x</normalizedToken>
|
||
the length of a procoxa; anterior margin of prosternum mesally projected as a wide triangle, apically either acute or rounded (except in
|
||
<taxonomicName id="A05EA98C0E18741205337FD6AB396CE1" lsidName="P. ocellatus" pageId="26" pageNumber="135" rank="species" species="ocellatus">
|
||
<emphasis id="C6AA2B14B57D9E16375C8DD199279423" italics="true" pageId="26" pageNumber="135">P. ocellatus</emphasis>
|
||
</taxonomicName>
|
||
); surface of prosternum flat to only weakly broadly convex, covered by scattered, fine, rather long setae; intercoxal process projected from posterior margin of procoxal cavities, rectangular in outline, mesally longitudinally carinate. Mesoventrite (Fig.
|
||
<figureCitation id="27CB5B0456FC967CC901A483221B4F28" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="26" pageNumber="135">14A</figureCitation>
|
||
) not fused to mesepisterna, with anterior margin nearly 0.3
|
||
<normalizedToken id="9FB8447B53F8EFECB6A329570B3694BF" originalValue="×">x</normalizedToken>
|
||
as wide as anterior margin of mesepisternum; anterior rib of mesoventrite bearing paired medial teardrop-shaped, pearlescent maculae; posterior elevation of mesoventrite with a transverse curved ridge, rather sharp and low, reduced in
|
||
<taxonomicName id="4A3AA9E47764C72F80BA78E629EBD298" lsidName="P. maipure" pageId="26" pageNumber="135" rank="species" species="maipure">
|
||
<emphasis id="AAF37BB3941C533149022D1B6F39FABB" italics="true" pageId="26" pageNumber="135">P. maipure</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="AC7794B190F1F3FFC334760428849419" lsidName="P. pijiguaense" pageId="26" pageNumber="135" rank="species" species="pijiguaense">
|
||
<emphasis id="A5B6062B6AE45A1A4876313D31087231" italics="true" pageId="26" pageNumber="135">P. pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E2B0B44EF907071E86101A05C7A96188" lsidName="P. ocellatus" pageId="26" pageNumber="135" rank="species" species="ocellatus">
|
||
<emphasis id="C5B4D312AB5E95CD3A0FCC1802D9A48E" italics="true" pageId="26" pageNumber="135">P. ocellatus</emphasis>
|
||
</taxonomicName>
|
||
, with a sharp, pyramidal (triangular) spine-like projection in
|
||
<taxonomicName id="E2DAD6E7AA049D499FF5C2DE4B1C6741" lsidName="P. cuspidis" pageId="26" pageNumber="135" rank="species" species="cuspidis">
|
||
<emphasis id="6A2A58913905918BA10E24AB4A2BC43C" italics="true" pageId="26" pageNumber="135">P. cuspidis</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="F0B170E085B6F64FB53831AFFB03DBDB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="26" pageNumber="135">11C</figureCitation>
|
||
); surface of mesoventrite reticulated for the most part, covered by scattered, fine and rather long setae, with anteromedial depression, and posterolateral smooth and glabrous areas; mesepisternum obliquely widely concave, with reticulated surface; mesepimeron trapezoid, with reticulate and pubescent surface. Mesofurca (examined in
|
||
<taxonomicName id="4A211AE21ED382BA0346DBDC2BAF41E8" lsidName="P. neutrum" pageId="26" pageNumber="135" rank="species" species="neutrum">
|
||
<emphasis id="94CAF6314724F35144FAE3CC32C388FF" italics="true" pageId="26" pageNumber="135">P. neutrum</emphasis>
|
||
</taxonomicName>
|
||
) with short arms, 0.75
|
||
<normalizedToken id="8E71EC93F70F72D63505FAB3CB616285" originalValue="×">x</normalizedToken>
|
||
length of mesocoxae; apical half of arms free, explanate at apex, somewhat square. Metaventrite mesally widely elevated, rather wide throughout and flat posteromesally; surface densely pubescent, except for posteromesal nearly as wide as long glabrous patch, and soemtimes postero-lateral areas (Fig.
|
||
<figureCitation id="F982F01CEE008AD5BC3540BF1A38CAFE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="26" pageNumber="135">10G</figureCitation>
|
||
; except in
|
||
<taxonomicName id="B9C14E66929682DCD70F9176F6F15944" lsidName="P. ocellatus" pageId="26" pageNumber="135" rank="species" species="ocellatus">
|
||
<emphasis id="7680A9F8BBEA7D2B0EF1B6AB6E3AB233" italics="true" pageId="26" pageNumber="135">P. ocellatus</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="CC7F118AC6B3994055A7BDFC11CFBDB1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus of Primocerus spp.: A-D Primocerus ocellatus: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus gigas: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure10" httpUri="https://binary.pensoft.net/fig/310067" pageId="26" pageNumber="135">10D</figureCitation>
|
||
). Metepisterna 3-4
|
||
<normalizedToken id="3C168767ADF8F456E96D1ED70F077AEB" originalValue="×">x</normalizedToken>
|
||
longer than wide, narrowing only at posterior end. Metepimeron clearly visible, triangular. Metafurca (examined in
|
||
<taxonomicName id="32CC68EDBA875F4BF96E4DB33B8978F2" lsidName="P. neutrum" pageId="26" pageNumber="135" rank="species" species="neutrum">
|
||
<emphasis id="BC5CC683034884EF5B12E014B5315EEF" italics="true" pageId="26" pageNumber="135">P. neutrum</emphasis>
|
||
</taxonomicName>
|
||
, Fig.
|
||
<figureCitation id="3746FC1CBBC05B2F0017569CC0A3ED8F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="26" pageNumber="135">14B</figureCitation>
|
||
) 1.3
|
||
<normalizedToken id="9AF7B29CB0AAB51CF0CF75CE58C489F4" originalValue="×">x</normalizedToken>
|
||
wider than long, with furcal arms slightly shorter than stalk; stalk triangular (wider near the crux, gradually narrowing ventrally), with paired longitudinal keels extending along basal third of posterior face, fusing together towards crux, with a well-developed median keel on anterior face extending to anterior margin of dorsal sheets; outer margins of stalk gradually diverging from base towards basal third of furcal arms; furcal arms somewhat parallelogram-shaped, with apex (hemiductus) only slightly explanate, with apex pointing obliquely; anterior tendons inserted basad of mid length of dorsal edge of furcal arms; well-developed dorsal sheaths, narrower than widest point of lateral sheaths.
|
||
<emphasis id="5F40FB885BB0AB56160110770D45E98E" bold="true" italics="true" pageId="26" pageNumber="135">Elytra.</emphasis>
|
||
Surface even (without elevations or depressions), with sutural striae; ground punctures and systematic punctures either shallow or sharply marked (e.g., Figs
|
||
<figureCitation id="05601B57CD43345DED18DB0EBEDA28C0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="26" pageNumber="135">11E</figureCitation>
|
||
,
|
||
<figureCitation id="D73D74A257D4E6C8B1D639CD6D5F1551" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="26" pageNumber="135">12E</figureCitation>
|
||
), similar in size and degree of impression throughout elytra, seemingly arranged in rows; serial punctures, when present (e.g., Fig.
|
||
<figureCitation id="9F0C1DDAFC1BE63EEB9951E6A31BF762" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="26" pageNumber="135">13A, D, G</figureCitation>
|
||
), larger and deeper than ground punctures, and clearly arranged in longitudinal rows (striae); serial punctures only very slightly impressed into grooves along posterior half of elytra in striate species (e.g.,
|
||
<taxonomicName id="1FE9E5F1990B25317A6DA41C014BBDF0" lsidName="P. petilus" pageId="26" pageNumber="135" rank="species" species="petilus">
|
||
<emphasis id="35B152E558EAB4E9A7E949AB2F3223C2" italics="true" pageId="26" pageNumber="135">P. petilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FBFF07A848F957F0DE5194508BF62FF3" lsidName="P. striatolatus" pageId="26" pageNumber="135" rank="species" species="striatolatus">
|
||
<emphasis id="7EF64833AAD5632E72E9BF04DBD84E71" italics="true" pageId="26" pageNumber="135">P. striatolatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="C4E148D5E016A82636594A1C67B69D98" lsidName="P. semipubescens" pageId="26" pageNumber="135" rank="species" species="semipubescens">
|
||
<emphasis id="BFC477FFF27BD35AF9C915710B088E93" italics="true" pageId="26" pageNumber="135">P. semipubescens</emphasis>
|
||
</taxonomicName>
|
||
; see Fig.
|
||
<figureCitation id="061AA93BD28F5201B366CC9A6A4AE06C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="26" pageNumber="135">13</figureCitation>
|
||
); seta-bearing systematic punctures rather scarce; elytral outer margins flared, usually along entire length. Epipleura usually well developed, surface either flat or oblique,
|
||
<pageBreakToken id="A8DE84B1C7C42E015EFCC234FE4C9268" pageId="27" pageNumber="136" start="start">with</pageBreakToken>
|
||
sparse setae and irregular sculpture, anteriorly wide, gradually narrowing posteriorly, extending up to midlength of first abdominal ventrite; inner margin of epipleura only slightly indented at anterior outer corner of metepisternum; pseudepipleura usually well developed and perpendicularly positioned, ranging in width from nearly as wide as anterior portion of epipleura, to half as wide, extending up to basal half of abdomen along outer margin of elytra. Hind wings well developed.
|
||
<emphasis id="1DF626F1FDA5D211DE7BC5C9EBDDADA0" bold="true" italics="true" pageId="27" pageNumber="136">Legs.</emphasis>
|
||
Pubescence on anterior surface of metafemora ranging from scarce and limited to anterior margin (e.g., Fig.
|
||
<figureCitation id="06B96AF950B837A8C90432DC6F059FDB" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="27" pageNumber="136">12C</figureCitation>
|
||
), to densely covering most surface up to apical fifth (e.g., Fig.
|
||
<figureCitation id="C23945881343464C0F2C95F29E164F4A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Habitus of Primocerus spp.: A-D Primocerus cuspidis: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus neutrum: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure11" httpUri="https://binary.pensoft.net/fig/310068" pageId="27" pageNumber="136">11G</figureCitation>
|
||
); glabrous area of metafemur with shiny and sometimes slightly reticulated surface; all femora antero-posteriorly flattened; metafemora usually with sharply marked tibial grooves. Tibiae slender, rather cylindrical; longitudinal rows of well-developed spines along pro-, meso- and metatibiae, composed of rather sharp and stout spines, slightly sparser along metatibiae; protibiae with a median longitudinal row of rather long and thick setae along anterior surface; protibial apical spurs large, extending beyond apex of protarsomere 2, sometimes reaching apex of protarsomere 3. All tarsi with five tar
|
||
<pageBreakToken id="099C43706D66A7A708F8388B42686516" pageId="28" pageNumber="137" start="start">someres</pageBreakToken>
|
||
, bearing long apical hair-like setae on dorsal face, and spine-like or hair-like setae on ventral face of tarsomeres 2-4, sometimes also tarsomere 5; pro- and mesotarsomeres 1-4 similar in size and shape; pro- and mesotarsomere 5 approximately as long as 3-4 combined; metatarsomere 2 similar in length to metatarsomere 5; claws rather large, curved; well-developed empodium, bearing a pair of long, curved apical setae.
|
||
<emphasis id="A116BABAD0ECB89F2F5DDD73A3840CA0" bold="true" italics="true" pageId="28" pageNumber="137">Abdomen.</emphasis>
|
||
Abdomen with five ventrites, rather flat to medially convex; all ventrites with uniform, dense, fine pubescence; posterior margin of fifth ventrite either rounded, truncate, or slightly emarginate, usually fringed with spine-like setae (Fig.
|
||
<figureCitation id="543D386AB68BAAF5796661A690501E50" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="28" pageNumber="137">14C</figureCitation>
|
||
). Aedeagus (Fig.
|
||
<figureCitation id="1A789B51E8A3FE56F1844B91D5D7AB28" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="28" pageNumber="137">
|
||
14
|
||
<normalizedToken id="0F423209019C41AF1B1D61EBB753DDD7" originalValue="D–L">D-L</normalizedToken>
|
||
</figureCitation>
|
||
) with basal piece as long or longer than parameres; median lobe triangular, with base nearly as wide as base of a paramere, with well-developed lateral basal apodemes; apex of median lobe variable, with a membranous to lightly sclerotized apical projection; gonopore not differentiated; parameres nearly as long as median lobe, with outer margins usually straight along basal 3/4, with setae at apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3CAAEACB8C29F0EC4A10E8994C6BC43" pageId="28" pageNumber="137" type="larvae">
|
||
<paragraph id="9E559BE52AE65DE4F28CFA7A33F27DAE" pageId="28" pageNumber="137">Larvae.</paragraph>
|
||
<paragraph id="7A93042F684300577FF92B8A7CDC5E07" pageId="28" pageNumber="137">The immature stages are unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67B13CBCA6E4CDB34FAD8EC83B1653A2" pageId="28" pageNumber="137" type="etymology">
|
||
<paragraph id="D4319D9157F62E6ABF5694D9CAEE6121" pageId="28" pageNumber="137">Etymology.</paragraph>
|
||
<paragraph id="7A788B7889AB84A096860D5F2A529D10" pageId="28" pageNumber="137">
|
||
Named from the Latin
|
||
<emphasis id="5B6E1337B34292D3AB53C532AD61F87B" italics="true" pageId="28" pageNumber="137">primus</emphasis>
|
||
, meaning first, with the ending -
|
||
<emphasis id="B9B8C6D563DB00D131F9A71B2C97754F" italics="true" pageId="28" pageNumber="137">cerus</emphasis>
|
||
, in reference to the belonging of the genus to the
|
||
<taxonomicName id="A5751A21BE6A6C1BEC46ED1A558F9B79" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" lsidName="" pageId="28" pageNumber="137" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
|
||
. To be treated as masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="847FB134E8243A31C44E66465FD43259" pageId="28" pageNumber="137" type="distribution">
|
||
<paragraph id="C00F358CA65B849D5419D3929B891BC3" pageId="28" pageNumber="137">Distribution.</paragraph>
|
||
<paragraph id="213890322DC04C0D56A416ED78B6BA5A" pageId="28" pageNumber="137">
|
||
Broadly distributed across the Guiana Shield region of South America, including Brazil (
|
||
<normalizedToken id="6FCF62A31CA85AC0560C6569CE8EF058" originalValue="Pará">Para</normalizedToken>
|
||
), Guyana, Suriname and southern Venezuela (Amazonas,
|
||
<normalizedToken id="0E8BFBC76B2C6CAB03A3E5552F667C6A" originalValue="Bolívar">Bolivar</normalizedToken>
|
||
) (Fig.
|
||
<figureCitation id="E2BEE27FFD1973581B28D4096D0C60F5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of Primocerus spp." figureDoi="10.3897/zookeys.855.33013.figure15" httpUri="https://binary.pensoft.net/fig/310072" pageId="28" pageNumber="137">15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B4138C731A3D0EDAA7EC4EE3A68C4774" ID-Zenodo-Dep="3251391" doi="10.3897/zookeys.855.33013.figure15" httpUri="https://binary.pensoft.net/fig/310072" pageId="28" pageNumber="137" start="Figure 15" startId="F15">
|
||
<paragraph id="32CD0598FF04195566E6BC860F6B944E" pageId="28" pageNumber="137">
|
||
<emphasis id="B66D3C76B8DC9F39EC261C3C0EF1991C" bold="true" pageId="28" pageNumber="137">Figure 15.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="EED77E6E1F1225E4051B407C746C2D4B" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="28" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="80C763430B5BF5084940A72FEEFBA35F" italics="true" pageId="28" pageNumber="137">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="8DA483C76926C75802F231E315CF1B12" pageId="28" pageNumber="137" type="remarks">
|
||
<paragraph id="A972305F4D204F1BA91DA797416396DD" pageId="28" pageNumber="137">Remarks.</paragraph>
|
||
<paragraph id="AC966A5B03921821E5D83E1DAE5A85F5" pageId="28" pageNumber="137">
|
||
The habitats occupied by members of
|
||
<taxonomicName id="C079E19BA7513139E6EF750A3964186B" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="28" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A279C1DA98CF82C5D78C60D7E8E3C168" italics="true" pageId="28" pageNumber="137">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
range from forested pools to seepages (Fig.
|
||
<figureCitation id="5FCAA641DB495CEE1557C6C313F9384D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Habitat of Primocerus spp. A habitat and type locality for P. cuspidis, Venezuela, Tobogan de la Selva, collecting event AS- 08 - 080 b B habitat and type locality for P. pijiguaense, Venezuela, Los Pijiguaos, collecting event AS- 07 - 015 C habitat and type locality for P. neutrum, Venezuela, along La Escalera, collecting event AS- 08 - 058 D habitat and type locality for P. petilus, Brazil, Vale do Paraiso, collecting event BR 18 - 0203 - 01 G." figureDoi="10.3897/zookeys.855.33013.figure16" httpUri="https://binary.pensoft.net/fig/310073" pageId="28" pageNumber="137">16</figureCitation>
|
||
), in an elevational range from 80 to 1950 m. Only one specimen has been collected with a flight intercept trap. Specimens of
|
||
<taxonomicName id="EE41120BF113305B42DAC6FB846402A1" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="28" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2783D076AAF22C78016C3B15795278D9" italics="true" pageId="28" pageNumber="137">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
are relatively rare, given that so far have only been found in low numbers of specimens per collecting event.
|
||
</paragraph>
|
||
<caption id="E76F4247939299759E77E6AC63097C1C" ID-Zenodo-Dep="3251393" doi="10.3897/zookeys.855.33013.figure16" httpUri="https://binary.pensoft.net/fig/310073" pageId="28" pageNumber="137" start="Figure 16" startId="F16">
|
||
<paragraph id="2F65A1FD6D9D66B249467EDD09F01555" pageId="28" pageNumber="137">
|
||
<emphasis id="4B8B75DCEA12D149DEC066E1FCD2B32E" bold="true" pageId="28" pageNumber="137">Figure 16.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="21CBE73D0AC9C8C029274300494FA62D" authorityName="Girón & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="28" pageNumber="137" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="01ECCBE5905C2FD986941F974A1EA0C0" italics="true" pageId="28" pageNumber="137">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="CEA3BCEB1C7F5AB3C0374651898CB2BD" bold="true" pageId="28" pageNumber="137">A</emphasis>
|
||
habitat and type locality for
|
||
<taxonomicName id="24B430398390065F85C82DC265E2A4C2" lsidName="P. cuspidis" pageId="28" pageNumber="137" rank="species" species="cuspidis">
|
||
<emphasis id="7E638B7A34E11DD34317AC616BE4D5A4" italics="true" pageId="28" pageNumber="137">P. cuspidis</emphasis>
|
||
</taxonomicName>
|
||
, Venezuela,
|
||
<normalizedToken id="FC915D1120A81C3A5BE78D60AA07005E" originalValue="Tobogán">Tobogan</normalizedToken>
|
||
de la Selva, collecting event AS-08-080b
|
||
<emphasis id="770880FCCD8FB34050D2BE93014DF42E" bold="true" pageId="28" pageNumber="137">B</emphasis>
|
||
habitat and type locality for
|
||
<taxonomicName id="547203FCA1FAA220EE8934768863B803" lsidName="P. pijiguaense" pageId="28" pageNumber="137" rank="species" species="pijiguaense">
|
||
<emphasis id="EF6308498DED530F7AAE993D37F143A4" italics="true" pageId="28" pageNumber="137">P. pijiguaense</emphasis>
|
||
</taxonomicName>
|
||
, Venezuela, Los Pijiguaos, collecting event AS-07-015
|
||
<emphasis id="4A49B1715DF46AD9A78974E258E737B6" bold="true" pageId="28" pageNumber="137">C</emphasis>
|
||
habitat and type locality for
|
||
<taxonomicName id="588752D25947FF4BCCDA3579AB037C59" lsidName="P. neutrum" pageId="28" pageNumber="137" rank="species" species="neutrum">
|
||
<emphasis id="3E1758898603507662B6AC423B3DD5D3" italics="true" pageId="28" pageNumber="137">P. neutrum</emphasis>
|
||
</taxonomicName>
|
||
, Venezuela, along La Escalera, collecting event AS-08-058
|
||
<emphasis id="DBBD2AB6BFD6973EE792009C2B1A7A9A" bold="true" pageId="28" pageNumber="137">D</emphasis>
|
||
habitat and type locality for
|
||
<taxonomicName id="B48534CB7718152C455C4381629315B9" lsidName="P. petilus" pageId="28" pageNumber="137" rank="species" species="petilus">
|
||
<emphasis id="AA22D9240BF2604998EBB6007C355B67" italics="true" pageId="28" pageNumber="137">P. petilus</emphasis>
|
||
</taxonomicName>
|
||
, Brazil, Vale do Paraiso, collecting event BR18-0203-01G.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |