447 lines
55 KiB
XML
447 lines
55 KiB
XML
<document id="91B5BFC411143A278A5660BCFA19F7F9" ID-DOI="10.5281/zenodo.175850" ID-GBIF-Dataset="df6a72a8-bc5e-43e5-9471-3700a7533e76" ID-ISSN="1175-5326" ID-Zenodo-Dep="175850" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459821720877" checkinUser="plazi" docAuthor="Arango, Claudia P. & Krapp, Franz" docDate="2007" docId="437AD90D2B20D518FF12FA4B006C7DEA" docLanguage="en" docName="zt01435p024.pdf" docOrigin="Zootaxa 1435" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Anoplodactylus perissoporus Arango & Krapp, 2007, new species" docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="BF43A1752B21D51CFF85FFF1033B7F61" masterDocTitle="A new species of Anoplodactylus (Arthropoda: Pycnogonida) from the Great Barrier Reef and discussion on the A. tenuicorpus - complex *" masterLastPageNumber="24" masterPageNumber="19" pageNumber="20" updateTime="1698221899491" updateUser="plazi">
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<mods:titleInfo id="901388FBFD39526EF0D5700B81270669">
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<mods:title id="A0C10B81F30A71C4EB9F297435586A7D">A new species of Anoplodactylus (Arthropoda: Pycnogonida) from the Great Barrier Reef and discussion on the A. tenuicorpus - complex *</mods:title>
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<mods:roleTerm id="7B18ECD5BF4CBC09258907624630CC91">Author</mods:roleTerm>
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<mods:namePart id="17B022AB64315D1F8DA5519DA97B5E8D">Arango, Claudia P.</mods:namePart>
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<mods:roleTerm id="6990F7284676A783D9E785314EEE5E78">Author</mods:roleTerm>
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<mods:namePart id="F8695BAACF964794E088060D421D1DA5">Krapp, Franz</mods:namePart>
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<mods:typeOfResource id="8DD35D93C5A1078413655D5AA0DCC552">text</mods:typeOfResource>
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<mods:title id="DD82E56299D50ABEA22CD75360F50429">Zootaxa</mods:title>
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<mods:part id="0FA1F36AA2354F314C1A39A6615D5E9A">
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<mods:date id="136DB577F05D0CDC8808EABC6492832E">2007</mods:date>
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<mods:detail id="32DE3084EC987C81A9B909E792C04CA7" type="volume">
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<mods:identifier id="97DDBC656B53380B5FE9C6EACEB1D88D" type="DOI">10.5281/zenodo.175850</mods:identifier>
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<mods:identifier id="D5C758047A2D18023D77063CF6129F16" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DF11D6C8989507699FBAC740D21481E2" type="Zenodo-Dep">175850</mods:identifier>
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<treatment id="437AD90D2B20D518FF12FA4B006C7DEA" ID-DOI="http://doi.org/10.5281/zenodo.5671733" ID-GBIF-Taxon="119355937" ID-Zenodo-Dep="5671733" LSID="urn:lsid:plazi:treatment:437AD90D2B20D518FF12FA4B006C7DEA" httpUri="http://treatment.plazi.org/id/437AD90D2B20D518FF12FA4B006C7DEA" lastPageId="4" lastPageNumber="23" pageId="1" pageNumber="20">
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<subSubSection id="83C93B902B20D51DFF12FA4B00367AB5" box="[151,781,1466,1492]" pageId="1" pageNumber="20" type="nomenclature">
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<paragraph id="CB6C681B2B20D51DFF12FA4B00367AB5" blockId="1.[151,781,1466,1492]" box="[151,781,1466,1492]" pageId="1" pageNumber="20">
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<heading id="9024DF772B20D51DFF12FA4B00367AB5" bold="true" box="[151,781,1466,1492]" fontSize="11" level="1" pageId="1" pageNumber="20" reason="1">
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<emphasis id="F9A7B4092B20D51DFF12FA4B00367AB5" bold="true" box="[151,781,1466,1492]" pageId="1" pageNumber="20">
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<taxonomicName id="0CD313982B20D51DFF12FA4B02C97AB5" ID-CoL="5V93C" box="[151,498,1466,1492]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="perissoporus" status="sp. nov.">
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<emphasis id="F9A7B4092B20D51DFF12FA4B02C97AB5" bold="true" box="[151,498,1466,1492]" italics="true" pageId="1" pageNumber="20">Anoplodactylus perissoporus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E29409722B20D51DFE7FFA4B01B17AB5" box="[506,650,1466,1492]" pageId="1" pageNumber="20" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="53E8749E2B20D51DFD1CFA4B00397AB5" box="[665,770,1466,1492]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="1" pageNumber="20">Figure 1</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="83C93B902B20D51DFF12F9F4060A7873" pageId="1" pageNumber="20" type="materials_examined">
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<paragraph id="CB6C681B2B20D51DFF12F9F4060A7873" blockId="1.[151,1436,1541,1810]" pageId="1" pageNumber="20">
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<typeStatus id="1468D6B92B20D51DFF12F9F40232797E" box="[151,265,1541,1567]" pageId="1" pageNumber="20" type="holotype">
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<emphasis id="F9A7B4092B20D51DFF12F9F40232797E" bold="true" box="[151,265,1541,1567]" pageId="1" pageNumber="20">Holotype</emphasis>
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</typeStatus>
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: S105845.
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<specimenCount id="DDD5A3922B20D51DFE11F9F40299797E" box="[404,418,1541,1567]" pageId="1" pageNumber="20" type="male">1</specimenCount>
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male with eggs. Great Barrier Reef (GBR), Central Section, Rib Reef, reef slope, in coral rubble,
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<specimenCount id="DDD5A3922B20D51DFEB4F9DC025E7926" box="[305,357,1581,1607]" pageId="1" pageNumber="20" type="generic">
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<quantity id="0C2BC5FE2B20D51DFEB4F9DC025A7926" box="[305,353,1581,1607]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="20" unit="m" value="8.0">8 m</quantity>
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.
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</specimenCount>
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<date id="BF6D4EDB2B20D51DFEE9F9DC01787926" box="[364,579,1581,1607]" pageId="1" pageNumber="20" value="1998-09-06">6 September 1998</date>
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.
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<emphasis id="F9A7B4092B20D51DFDCBF9DC01F07926" bold="true" box="[590,715,1581,1607]" pageId="1" pageNumber="20">
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<typeStatus id="1468D6B92B20D51DFDCBF9DC01F07926" box="[590,715,1581,1607]" pageId="1" pageNumber="20" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: S105846.
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<specimenCount id="DDD5A3922B20D51DFCD4F9DC005B7926" box="[849,864,1581,1607]" pageId="1" pageNumber="20" type="female">1</specimenCount>
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female; GBR, Central Section, Pandora Reef,
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<quantity id="0C2BC5FE2B20D51DFAFAF9DC03877913" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="20" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3– 6m</quantity>
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in rubble with algae,
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<date id="BF6D4EDB2B20D51DFE53F9A901407913" box="[470,635,1624,1650]" pageId="1" pageNumber="20" value="1999-07-15">15 July 1999</date>
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. S105847.
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<specimenCount id="DDD5A3922B20D51DFC8DF9A9002C7913" box="[776,791,1624,1650]" pageId="1" pageNumber="20" type="juvenile">1</specimenCount>
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juvenile, instar with 4th legs developing. Coral Sea, Flinders Reef, front reef,
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<quantity id="0C2BC5FE2B20D51DFE41F97102C079FB" box="[452,507,1664,1690]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="20" unit="m" value="12.0">12m</quantity>
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, in
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<taxonomicName id="0CD313982B20D51DFDA3F97101A279F8" box="[550,665,1664,1689]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="1" pageNumber="20" phylum="Chlorophyta" rank="genus">
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<emphasis id="F9A7B4092B20D51DFDA3F97101A279F8" box="[550,665,1664,1689]" italics="true" pageId="1" pageNumber="20">Halimeda</emphasis>
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</taxonomicName>
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.
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<date id="BF6D4EDB2B20D51DFD2CF971000C79FB" box="[681,823,1664,1690]" pageId="1" pageNumber="20" value="1999-07-03">3 July 1999</date>
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. S105848.
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<specimenCount id="DDD5A3922B20D51DFC3EF97100F179FB" box="[955,970,1664,1690]" pageId="1" pageNumber="20" type="female">1</specimenCount>
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female. Coral Sea, Flinders Reef, front reef,
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<quantity id="0C2BC5FE2B20D51DFF56F959023179A3" box="[211,266,1704,1730]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="20" unit="m" value="12.0">12m</quantity>
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, in
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<taxonomicName id="0CD313982B20D51DFEB6F959029C79A0" box="[307,423,1704,1729]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="1" pageNumber="20" phylum="Chlorophyta" rank="genus">
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<emphasis id="F9A7B4092B20D51DFEB6F959029C79A0" box="[307,423,1704,1729]" italics="true" pageId="1" pageNumber="20">Halimeda</emphasis>
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</taxonomicName>
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. S105849.
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<specimenCount id="DDD5A3922B20D51DFDA9F959010079A3" box="[556,571,1704,1730]" pageId="1" pageNumber="20" type="male">1</specimenCount>
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male with eggs,
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<specimenCount id="DDD5A3922B20D51DFC85F959004E79A3" box="[768,885,1704,1730]" pageId="1" pageNumber="20" type="female">3 females</specimenCount>
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; GBR, Central Section, Pandora Reef,
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<quantity id="0C2BC5FE2B20D51DFABEF959064479A3" box="[1339,1407,1704,1730]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="20" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6m</quantity>
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in rubble,
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<date id="BF6D4EDB2B20D51DFF74F92102AC798B" box="[241,407,1744,1770]" pageId="1" pageNumber="20" value="2000-04-19">19 April 2000</date>
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. CA111
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<specimenCount id="DDD5A3922B20D51DFE7BF9210172798B" box="[510,585,1744,1770]" pageId="1" pageNumber="20" type="male">1male</specimenCount>
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. GBR Central Section, Pandora Reef,
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<quantity id="0C2BC5FE2B20D51DFB89F921070A798B" box="[1036,1073,1744,1770]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="20" unit="m" value="8.0">8m</quantity>
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in rubble,
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<date id="BF6D4EDB2B20D51DFB34F92106A3798B" box="[1201,1432,1744,1770]" pageId="1" pageNumber="20" value="1999-11-14">14 November 1999</date>
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. CA141, 184.
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<specimenCount id="DDD5A3922B20D51DFEB0F90902787873" box="[309,323,1784,1810]" pageId="1" pageNumber="20" type="female">1</specimenCount>
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female,
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<specimenCount id="DDD5A3922B20D51DFE2CF90902C77873" box="[425,508,1784,1810]" pageId="1" pageNumber="20" type="male">1 male</specimenCount>
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. GBR, Central Section, Pandora Reef,
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<quantity id="0C2BC5FE2B20D51DFC46F909073D7873" box="[963,1030,1784,1810]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="20" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6m</quantity>
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in rubble,
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<date id="BF6D4EDB2B20D51DFB01F90906177873" box="[1156,1324,1784,1810]" pageId="1" pageNumber="20" value="2000-04-19">19 April 2000</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="83C93B902B20D51DFF12F8B9077378BB" pageId="1" pageNumber="20" type="diagnosis">
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<paragraph id="CB6C681B2B20D51DFF12F8B902357800" blockId="1.[151,1436,1864,2010]" box="[151,270,1864,1889]" pageId="1" pageNumber="20">
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<emphasis id="F9A7B4092B20D51DFF12F8B902357800" box="[151,270,1864,1889]" italics="true" pageId="1" pageNumber="20">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="CB6C681B2B20D51DFF43F881077378BB" blockId="1.[151,1436,1864,2010]" pageId="1" pageNumber="20">Five to nine cement gland pores on femora of males. Smooth chelae. Tibia 2 the longest leg segment, tibia 1 slightly longer than femur; 13 or more crenulations on the distal margin of major heel spine; eight sole spines; tiny distal lamina. Body glabrous, segmentation lines faint or obliterated.</paragraph>
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</subSubSection>
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<subSubSection id="83C93B902B23D51FFF12FF6901F07DF3" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21" type="description">
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<paragraph id="CB6C681B2B23D51EFF12FF6902197FD0" blockId="2.[151,1436,152,378]" box="[151,290,152,177]" pageId="2" pageNumber="21">
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<emphasis id="F9A7B4092B23D51EFF12FF6902197FD0" box="[151,290,152,177]" italics="true" pageId="2" pageNumber="21">Description</emphasis>
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</paragraph>
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<paragraph id="CB6C681B2B23D51EFF43FF3102F67E1B" blockId="2.[151,1436,152,378]" pageId="2" pageNumber="21">
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Specimens of moderate size for genus, leg span about
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<quantity id="0C2BC5FE2B23D51EFCC5FF3100B37FBB" box="[832,904,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="21" unit="mm" value="7.0">7 mm</quantity>
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. Body and legs extremely slender, very attenuate form. Lateral processes separated about four to six times their diameter, as long as two to four times their anterior diameter, the third and fourth pair shorter. Trunk segmentation lines almost indistinct (
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<figureCitation id="53E8749E2B23D51EFB6BFEE1060B7E4B" box="[1262,1328,272,298]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
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A). Ocular tubercle very low, rounded; eyes large, lightly pigmented. Proboscis with swelling at midpoint (
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<figureCitation id="53E8749E2B23D51EFAB5FEC9064E7E33" box="[1328,1397,312,338]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
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B). Abdomen small, glabrous.
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</paragraph>
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<caption id="9FAC38932B23D51EFF12F9E506247910" httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21" targetBox="[420,1166,437,1530]" targetPageId="2">
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<paragraph id="CB6C681B2B23D51EFF12F9E506247910" blockId="2.[151,1437,1556,1649]" pageId="2" pageNumber="21">
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<emphasis id="F9A7B4092B23D51EFF12F9E50223794D" bold="true" box="[151,280,1556,1580]" pageId="2" pageNumber="21">FIGURE 1.</emphasis>
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<taxonomicName id="0CD313982B23D51EFE9AF9E4016E794D" box="[287,597,1557,1580]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="perissoporus" status="sp. nov.">
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<emphasis id="F9A7B4092B23D51EFE9AF9E4016E794D" box="[287,597,1557,1580]" italics="true" pageId="2" pageNumber="21">Anoplodactylus perissoporus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E29409722B23D51EFDDFF9E401AE794D" box="[602,661,1557,1580]" pageId="2" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
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, from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3mm). E. Third right leg (0.5mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25mm). H. Distal oviger segments (details from D).
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</paragraph>
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</caption>
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<paragraph id="CB6C681B2B23D51EFF43F95407AC7986" blockId="2.[151,1436,1701,2007]" pageId="2" pageNumber="21">
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Chelifores long, slender, diverging in acute angle at base, chelae half the length of scape, palms rectangular, as long as fingers, fingers curved and overlapping at tips, no sign of teeth (
|
||
<figureCitation id="53E8749E2B23D51EFBA8F93C074B7986" box="[1069,1136,1741,1767]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B23D51EFF43F904006478E6" blockId="2.[151,1436,1701,2007]" pageId="2" pageNumber="21">
|
||
Ovigers arise ventrally on first lateral processes, bases seen dorsally near the middle (see
|
||
<figureCitation id="53E8749E2B23D51EFB62F9040612786E" box="[1255,1321,1781,1807]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
|
||
A). Third segment length more than 10 times its width, with short setae; fourth segment longer than fifth, with two ectal and one endal setae; fifth segment with five ectal setae and one endal strong seta, sixth as long as fifth, with three endal setae, one ectal and two distal setae (
|
||
<figureCitation id="53E8749E2B23D51EFD4AF89C002A78E6" box="[719,785,1901,1927]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
|
||
D, H).
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B23D51FFF43F86402D07EAB" blockId="2.[151,1436,1701,2007]" lastBlockId="3.[151,1437,152,658]" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">
|
||
Legs very slender, coxae 2 three times longer than 1 and 3, these two subequal, tibia 2 the longest segment, tibia 1 just slightly longer than femur (
|
||
<figureCitation id="53E8749E2B23D51EFD1AF84C01DB78B6" box="[671,736,1981,2007]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="2" pageNumber="21">Fig. 1</figureCitation>
|
||
E), femur with five to nine low cement gland cups (right leg 1: 9, leg 2: 9, leg 3: 6, leg 4: 5; left leg 1: 8, leg 2: lost, leg 3: 7, leg 4: 6) (
|
||
<figureCitation id="53E8749E2B22D51FFB9FFF6907667FD3" box="[1050,1117,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
|
||
F). Tarsus small, propodus slender, with three heel spines, the most distal the largest, with 13 teeth; eight sole spines, tiny distal lamina. Propodal main claw long, auxiliaries absent (
|
||
<figureCitation id="53E8749E2B22D51FFD37FF1901CD7E63" box="[690,758,232,258]" captionStart="FIGURE 1" captionStartId="2.[151,255,1556,1580]" captionTargetBox="[420,1166,437,1530]" captionTargetId="figure@2.[420,1166,437,1530]" captionTargetPageId="2" captionText="FIGURE 1. Anoplodactylus perissoporus n. sp., from Rib Reef, Central Section of Great Barrier Reef, holotype male. A. Dorsal view (0.5 mm). B. Proboscis (0.1 mm). C. Chelae (0.1 mm). D. Oviger (0.3 mm). E. Third right leg (0.5 mm). F. Femur of third left leg (0.5 mm). G. Distal leg articles (0.25 mm). H. Distal oviger segments (details from D)." httpUri="https://zenodo.org/record/175851/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
|
||
G). Genital pores ventrally on second coxae of third and fourth legs. Female slightly larger than male (
|
||
<quantity id="0C2BC5FE2B22D51FFD2BFEE100397E4B" box="[686,770,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="22" unit="mm" value="2.3">2.3mm</quantity>
|
||
in body length), with slightly longer propodal claw, otherwise same characteristics as male. Genital pores ventrally on second coxae of all legs. One male from Pandora Reef (CA111) has dorsodistal setae on major leg segments and an incipient crenulation of second heel spine of some legs. This specimen has five to nine cement glands as the
|
||
<typeStatus id="1468D6B92B22D51FFC60FE7907757EC3" box="[997,1102,392,418]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
, justifying its designation as
|
||
<taxonomicName id="0CD313982B22D51FFF12FE4102757EA8" box="[151,334,432,457]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="perissoporus" status="sp. nov.">
|
||
<emphasis id="F9A7B4092B22D51FFF12FE4102757EA8" box="[151,334,432,457]" italics="true" pageId="3" pageNumber="22">A. perissoporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E29409722B22D51FFED3FE4102DC7EAB" box="[342,487,432,458]" pageId="3" pageNumber="22" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B22D51FFF43FE2901F07DF3" blockId="3.[151,1437,152,658]" pageId="3" pageNumber="22">
|
||
Measurements
|
||
<typeStatus id="1468D6B92B22D51FFEFFFE2902DE7E93" box="[378,485,472,498]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
: Body length (from tip of last lateral process to anterior margin of cephalic segment):
|
||
<quantity id="0C2BC5FE2B22D51FFF77FDF102737D7B" box="[242,328,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="22" unit="mm" value="2.1">2.1mm</quantity>
|
||
; width (across second pair of lateral processes):
|
||
<quantity id="0C2BC5FE2B22D51FFC33FDF107217D7B" box="[950,1050,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="22" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; second lateral process length:
|
||
<quantity id="0C2BC5FE2B22D51FFF12FDD903C07D23" box="[151,251,552,578]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="22" unit="mm" value="0.45">0.45mm</quantity>
|
||
; chelifore scape length:
|
||
<quantity id="0C2BC5FE2B22D51FFDBCFDD901A67D23" box="[569,669,552,578]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="22" unit="mm" value="0.48">0.48mm</quantity>
|
||
; palm:
|
||
<quantity id="0C2BC5FE2B22D51FFC81FDD900537D23" box="[772,872,552,578]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="22" unit="mm" value="0.14">0.14mm</quantity>
|
||
; chela fingers: ~
|
||
<quantity id="0C2BC5FE2B22D51FFBD7FDD9078C7D23" box="[1106,1207,552,578]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="22" unit="mm" value="0.14">0.14mm</quantity>
|
||
; proboscis length:
|
||
<quantity id="0C2BC5FE2B22D51FFF12FDA103C07D0B" box="[151,251,592,618]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="22" unit="mm" value="0.39">0.39mm</quantity>
|
||
; proboscis width at midpoint:
|
||
<quantity id="0C2BC5FE2B22D51FFDE3FDA101E17D0B" box="[614,730,592,618]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="22" unit="mm" value="0.135">0.135mm</quantity>
|
||
; femur:
|
||
<quantity id="0C2BC5FE2B22D51FFCBBFDA100987D0B" box="[830,931,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="1.46" pageId="3" pageNumber="22" unit="mm" value="1.46">1.46mm</quantity>
|
||
; tibia 1:
|
||
<quantity id="0C2BC5FE2B22D51FFB88FDA107497D0B" box="[1037,1138,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="3" pageNumber="22" unit="mm" value="1.48">1.48mm</quantity>
|
||
; tibia 2:
|
||
<quantity id="0C2BC5FE2B22D51FFB59FDA1067B7D0B" box="[1244,1344,592,618]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="3" pageNumber="22" unit="mm" value="1.76">1.76mm</quantity>
|
||
; propodus:
|
||
<quantity id="0C2BC5FE2B22D51FFF4AFD8902087DF3" box="[207,307,632,658]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="22" unit="mm" value="0.55">0.55mm</quantity>
|
||
; claw:
|
||
<quantity id="0C2BC5FE2B22D51FFE03FD8902E77DF3" box="[390,476,632,658]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="22" unit="mm" value="0.3">0.3mm</quantity>
|
||
; abdomen:
|
||
<quantity id="0C2BC5FE2B22D51FFDE6FD8901FC7DF3" box="[611,711,632,658]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="3" pageNumber="22" unit="mm" value="0.18">0.18mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83C93B902B22D51FFF12FD3900C77C53" pageId="3" pageNumber="22" type="etymology">
|
||
<paragraph id="CB6C681B2B22D51FFF12FD39022F7D80" blockId="3.[151,1436,712,818]" box="[151,276,712,737]" pageId="3" pageNumber="22">
|
||
<emphasis id="F9A7B4092B22D51FFF12FD39022F7D80" box="[151,276,712,737]" italics="true" pageId="3" pageNumber="22">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B22D51FFF43FD0100C77C53" blockId="3.[151,1436,712,818]" pageId="3" pageNumber="22">
|
||
Gr.
|
||
<emphasis id="F9A7B4092B22D51FFF77FD0102697C68" box="[242,338,752,777]" italics="true" pageId="3" pageNumber="22">perissos</emphasis>
|
||
, beyond the regular number, extraordinary, odd number. In reference to the numerous cement gland pores on the femora of males, an unusual feature in
|
||
<taxonomicName id="0CD313982B22D51FFCBAFCE900CE7C50" box="[831,1013,792,817]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9A7B4092B22D51FFCBAFCE900CE7C50" box="[831,1013,792,817]" italics="true" pageId="3" pageNumber="22">Anoplodactylus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83C93B902B22D51FFF12FC9302237C1A" box="[151,280,866,891]" pageId="3" pageNumber="22" type="discussion">
|
||
<paragraph id="CB6C681B2B22D51FFF12FC9302237C1A" blockId="3.[151,1437,866,1287]" box="[151,280,866,891]" pageId="3" pageNumber="22">
|
||
<emphasis id="F9A7B4092B22D51FFF12FC9302237C1A" box="[151,280,866,891]" italics="true" pageId="3" pageNumber="22">Discussion</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="83C93B902B22D518FF43FC74006C7DEA" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22" type="reference_group">
|
||
<paragraph id="CB6C681B2B22D51FFF43FC7406437A66" blockId="3.[151,1437,866,1287]" pageId="3" pageNumber="22">
|
||
The
|
||
<taxonomicName id="0CD313982B22D51FFF79FC7402BC7CFF" ID-CoL="66X4L" box="[252,391,901,926]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B22D51FFF79FC7402BC7CFF" box="[252,391,901,926]" italics="true" pageId="3" pageNumber="22">tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
complex seems to be confined to the Indo-West Pacific region (
|
||
<figureCitation id="53E8749E2B22D51FFBFBFC7407DD7CFE" box="[1150,1254,901,927]" captionStart="FIGURE 2" captionStartId="4.[151,255,1913,1937]" captionTargetBox="[152,1434,947,1896]" captionTargetId="figure@4.[152,1435,947,1897]" captionTargetPageId="4" captionText="FIGURE 2. Distribution map of the species in the A. tenuicorpus complex. Records of single male and females of A. tenuicorpus are shown: Female paratype from Aliguay Island, Philippines (Child 1988 b), one female from Aldabra Atoll (Child 1988 a), one female from Seychelles (Child 1988 a), one female from Calalan Bank off Glass Breakwater, near Magundas, Guam (Child 1991), one female from Madang, Riwa Bay, Port Moresby Bootlet Inlet (Child 1996)." httpUri="https://zenodo.org/record/175852/files/figure.png" pageId="3" pageNumber="22">Figure 2</figureCitation>
|
||
). In the case of specimens identified as
|
||
<taxonomicName id="0CD313982B22D51FFE2BFC5C01637CA7" ID-CoL="66X4L" box="[430,600,941,966]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B22D51FFE2BFC5C01637CA7" box="[430,600,941,966]" italics="true" pageId="3" pageNumber="22">A. tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
only the
|
||
<typeStatus id="1468D6B92B22D51FFD41FC5C01CC7CA6" box="[708,759,941,967]" pageId="3" pageNumber="22">type</typeStatus>
|
||
locality (Siquijor Island, off Mindanao) yielded a male (= the
|
||
<typeStatus id="1468D6B92B22D51FFF46FC2402147C8E" box="[195,303,981,1007]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
). Other records refer only to females (see
|
||
<figureCitation id="53E8749E2B22D51FFCA1FC2400B17C8E" box="[804,906,981,1007]" captionStart="FIGURE 2" captionStartId="4.[151,255,1913,1937]" captionTargetBox="[152,1434,947,1896]" captionTargetId="figure@4.[152,1435,947,1897]" captionTargetPageId="4" captionText="FIGURE 2. Distribution map of the species in the A. tenuicorpus complex. Records of single male and females of A. tenuicorpus are shown: Female paratype from Aliguay Island, Philippines (Child 1988 b), one female from Aldabra Atoll (Child 1988 a), one female from Seychelles (Child 1988 a), one female from Calalan Bank off Glass Breakwater, near Magundas, Guam (Child 1991), one female from Madang, Riwa Bay, Port Moresby Bootlet Inlet (Child 1996)." httpUri="https://zenodo.org/record/175852/files/figure.png" pageId="3" pageNumber="22">Figure 2</figureCitation>
|
||
). Traditionally, most
|
||
<taxonomicName id="0CD313982B22D51FFB0DFC2406057C8F" ID-CoL="84LFK" box="[1160,1342,981,1006]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9A7B4092B22D51FFB0DFC2406057C8F" box="[1160,1342,981,1006]" italics="true" pageId="3" pageNumber="22">Anoplodactylus</emphasis>
|
||
</taxonomicName>
|
||
species can be readily identified only if male specimens are available (but see
|
||
<bibRefCitation id="AF4215EA2B22D51FFC76FC0C06137B76" author="Arango" box="[1011,1320,1021,1047]" pageId="3" pageNumber="24" refString="Arango, C. P. & Maxmen, A. (2006) Proboscis ornamentation as diagnostic character for the Anoplodactylus californicus-digitatus complex (Arthropoda: Pycnogonida) with an example from the Anoplodactylus eroticus female. Zootaxa, 1311, 51 - 64." type="journal article" year="2006">Arango & Maxmen 2006</bibRefCitation>
|
||
), because diagnostic characters are usually related to male secondary sexual characteristics such as form and number of cement gland openings and ovigers features. When Child reported the females with no accompanying males from Eastern Indian Ocean (
|
||
<bibRefCitation id="AF4215EA2B22D51FFE61FB8401407BEE" author="Child" box="[484,635,1141,1167]" pageId="3" pageNumber="24" refString="Child, A. C. (1988 a) Pycnogonida from Aldabra Atoll. Biological Society of Washington Bulletin, 8, 45 - 78." type="journal article" year="1988" yearSuffix="a">Child 1988a</bibRefCitation>
|
||
), he relied only on their thin appearance and the serrated heel spine. Now, with at least another well-definable species (from
|
||
<collectingCountry id="B3C4288B2B22D51FFCBFFB6C00A87BD6" box="[826,915,1181,1207]" name="Yemen" pageId="3" pageNumber="22">Socotra</collectingCountry>
|
||
Island) known to occur in the northeastern part of the Indian Ocean, some doubt is cast upon whether the females recorded from Aldabra and the
|
||
<collectingCountry id="B3C4288B2B22D51FFAE3FB3403D37A66" name="Seychelles" pageId="3" pageNumber="22">Seychelles</collectingCountry>
|
||
should be conspecific with that first described based on a male from the
|
||
<collectingCountry id="B3C4288B2B22D51FFBC5FB1C07FF7A66" box="[1088,1220,1261,1287]" name="Philippines" pageId="3" pageNumber="22">Philippines</collectingCountry>
|
||
(
|
||
<bibRefCitation id="AF4215EA2B22D51FFB56FB1C06577A66" author="Child" box="[1235,1388,1261,1287]" pageId="3" pageNumber="24" refString="Child, A. C. (1988 b) Pycnogonida of the Western Pacific islands III. Recent-Smithsonian Philippine Expeditions. Smithsonian Contributions to Zoology, 468, 1 - 32." type="journal article" year="1988" yearSuffix="b">Child 1988b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="9FAC38932B22D51FFF12FACF06647A37" ID-Table-UUID="9FAC38932B22D51FFF12FACF06647A37" box="[151,1375,1342,1366]" httpUri="http://table.plazi.org/id/9FAC38932B22D51FFF12FACF06647A37" pageId="3" pageNumber="22" targetBox="[156,1428,1399,2020]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="CB6C681B2B22D51FFF12FACF06647A37" blockId="3.[151,1375,1342,1366]" box="[151,1375,1342,1366]" pageId="3" pageNumber="22">
|
||
<emphasis id="F9A7B4092B22D51FFF12FACF02337A37" bold="true" box="[151,264,1342,1366]" pageId="3" pageNumber="22">TABLE 1.</emphasis>
|
||
Main characters distinguishing the four attenuated
|
||
<taxonomicName id="0CD313982B22D51FFCB0FACE00E07A37" ID-CoL="84LFK" box="[821,987,1343,1366]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9A7B4092B22D51FFCB0FACE00E07A37" box="[821,987,1343,1366]" italics="true" pageId="3" pageNumber="22">Anoplodactylus</emphasis>
|
||
</taxonomicName>
|
||
species in the
|
||
<taxonomicName id="0CD313982B22D51FFBFDFACE07CD7A37" ID-CoL="66X4L" box="[1144,1270,1343,1366]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B22D51FFBFDFACE07CD7A37" box="[1144,1270,1343,1366]" italics="true" pageId="3" pageNumber="22">tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CB6C681B2B22D51FFF19FA8606757885" pageId="3" pageNumber="22">
|
||
<table id="B9D39ABB2B222AE3FF19FA8606AF7885" box="[156,1428,1399,2020]" gridcols="4" gridrows="16" pageId="3" pageNumber="22">
|
||
<tr id="75E36A592B222AE3FF19FA8606AF7AEC" box="[156,1428,1399,1421]" gridrow="0" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19FA8601A27AEC" box="[156,665,1399,1421]" gridcol="0" gridrow="0" pageId="3" pageNumber="22">
|
||
Characters
|
||
<taxonomicName id="0CD313982B22D51FFE52FA8901687AEC" ID-CoL="5V93C" box="[471,595,1400,1421]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="perissoporus" status="sp. nov.">
|
||
<emphasis id="F9A7B4092B22D51FFE52FA8901687AEC" box="[471,595,1400,1421]" italics="true" pageId="3" pageNumber="22">perissoporus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E29409722B22D51FFDDCFA8601BC7AED" box="[601,647,1399,1420]" pageId="3" pageNumber="22" rank="species">n.sp.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="363203252B222AE3FD36FA8600BB7AEC" box="[691,896,1399,1421]" gridcol="1" gridrow="0" pageId="3" pageNumber="22">
|
||
<taxonomicName id="0CD313982B22D51FFD41FA89004B7AED" ID-CoL="66X4L" authority="Child" authorityName="Child" box="[708,880,1399,1421]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B22D51FFD41FA89000E7AEC" box="[708,821,1400,1421]" italics="true" pageId="3" pageNumber="22">tenuicorpus</emphasis>
|
||
Child
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="363203252B222AE3FC2EFA8607AB7AEC" box="[939,1168,1399,1421]" gridcol="2" gridrow="0" pageId="3" pageNumber="22">
|
||
<taxonomicName id="0CD313982B22D51FFC40FA89074D7AED" ID-CoL="5V8RT" authority="Stock" authorityName="Stock" box="[965,1142,1399,1421]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="exaggeratus">
|
||
<emphasis id="F9A7B4092B22D51FFC40FA8907017AEC" box="[965,1082,1400,1421]" italics="true" pageId="3" pageNumber="22">exaggeratus</emphasis>
|
||
Stock
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="363203252B222AE3FB2BFA8606AF7AEC" box="[1198,1428,1399,1421]" gridcol="3" gridrow="0" pageId="3" pageNumber="22">Socotra species</th>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19FA6E06AF7AD5" box="[156,1428,1439,1460]" gridrow="1" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19FA6E01A27AD5" box="[156,665,1439,1460]" gridcol="0" gridrow="1" pageId="3" pageNumber="22">Chelae length 1/2 of scape</th>
|
||
<td id="363203252B222AE3FD36FA6E00BB7AD5" box="[691,896,1439,1460]" gridcol="1" gridrow="1" pageId="3" pageNumber="22">1/3 of scape</td>
|
||
<td id="363203252B222AE3FC2EFA6E07AB7AD5" box="[939,1168,1439,1460]" gridcol="2" gridrow="1" pageId="3" pageNumber="22">1/2 of scape</td>
|
||
<td id="363203252B222AE3FB2BFA6E06AF7AD5" box="[1198,1428,1439,1460]" gridcol="3" gridrow="1" pageId="3" pageNumber="22">1/2 of scape</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19FA3606AF7ABD" box="[156,1428,1479,1500]" gridrow="2" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19FA3601A27ABD" box="[156,665,1479,1500]" gridcol="0" gridrow="2" pageId="3" pageNumber="22">Chelae teeth number Not present</th>
|
||
<td id="363203252B222AE3FD36FA3600BB7ABD" box="[691,896,1479,1500]" gridcol="1" gridrow="2" pageId="3" pageNumber="22">3+3</td>
|
||
<td id="363203252B222AE3FC2EFA3607AB7ABD" box="[939,1168,1479,1500]" gridcol="2" gridrow="2" pageId="3" pageNumber="22">5+3</td>
|
||
<td id="363203252B222AE3FB2BFA3606AF7ABD" box="[1198,1428,1479,1500]" gridcol="3" gridrow="2" pageId="3" pageNumber="22">4–5 + 5</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19FA1E06AF7965" box="[156,1428,1519,1540]" gridrow="3" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19FA1E01A27965" box="[156,665,1519,1540]" gridcol="0" gridrow="3" pageId="3" pageNumber="22">Teeth shape on chela N/A</th>
|
||
<td id="363203252B222AE3FD36FA1E00BB7965" box="[691,896,1519,1540]" gridcol="1" gridrow="3" pageId="3" pageNumber="22">triangles</td>
|
||
<td id="363203252B222AE3FC2EFA1E07AB7965" box="[939,1168,1519,1540]" gridcol="2" gridrow="3" pageId="3" pageNumber="22">needles</td>
|
||
<td id="363203252B222AE3FB2BFA1E06AF7965" box="[1198,1428,1519,1540]" gridcol="3" gridrow="3" pageId="3" pageNumber="22">needles</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F9E606AF794D" box="[156,1428,1559,1580]" gridrow="4" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F9E601A2794D" box="[156,665,1559,1580]" gridcol="0" gridrow="4" pageId="3" pageNumber="22">Chelae palm 2 x largest diameter</th>
|
||
<td id="363203252B222AE3FD36F9E600BB794D" box="[691,896,1559,1580]" gridcol="1" gridrow="4" pageId="3" pageNumber="22">2 x largest diameter</td>
|
||
<td id="363203252B222AE3FC2EF9E607AB794D" box="[939,1168,1559,1580]" gridcol="2" gridrow="4" pageId="3" pageNumber="22">3 x largest diameter</td>
|
||
<td id="363203252B222AE3FB2BF9E606AF794D" box="[1198,1428,1559,1580]" gridcol="3" gridrow="4" pageId="3" pageNumber="22">2 x largest diameter</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F9CE06AF7935" box="[156,1428,1599,1620]" gridrow="5" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F9CE01A27935" box="[156,665,1599,1620]" gridcol="0" gridrow="5" pageId="3" pageNumber="22">Propodal cutting lamina tiny, distal</th>
|
||
<td id="363203252B222AE3FD36F9CE00BB7935" box="[691,896,1599,1620]" gridcol="1" gridrow="5" pageId="3" pageNumber="22">tiny, distal</td>
|
||
<td id="363203252B222AE3FC2EF9CE07AB7935" box="[939,1168,1599,1620]" gridcol="2" gridrow="5" pageId="3" pageNumber="22">absent</td>
|
||
<td id="363203252B222AE3FB2BF9CE06AF7935" box="[1198,1428,1599,1620]" gridcol="3" gridrow="5" pageId="3" pageNumber="22">absent</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F99606AF791D" box="[156,1428,1639,1660]" gridrow="6" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F99601A2791D" box="[156,665,1639,1660]" gridcol="0" gridrow="6" pageId="3" pageNumber="22">Propodal sole spines 8</th>
|
||
<td id="363203252B222AE3FD36F99600BB791D" box="[691,896,1639,1660]" gridcol="1" gridrow="6" pageId="3" pageNumber="22">~ 5</td>
|
||
<td id="363203252B222AE3FC2EF99607AB791D" box="[939,1168,1639,1660]" gridcol="2" gridrow="6" pageId="3" pageNumber="22">~ 7</td>
|
||
<td id="363203252B222AE3FB2BF99606AF791D" box="[1198,1428,1639,1660]" gridcol="3" gridrow="6" pageId="3" pageNumber="22">~ 9</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F97E06AF79C5" box="[156,1428,1679,1700]" gridrow="7" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F97E01A279C5" box="[156,665,1679,1700]" gridcol="0" gridrow="7" pageId="3" pageNumber="22">Heel spine crenulations +13+</th>
|
||
<td id="363203252B222AE3FD36F97E00BB79C5" box="[691,896,1679,1700]" gridcol="1" gridrow="7" pageId="3" pageNumber="22">7–8</td>
|
||
<td id="363203252B222AE3FC2EF97E07AB79C5" box="[939,1168,1679,1700]" gridcol="2" gridrow="7" pageId="3" pageNumber="22">12</td>
|
||
<td id="363203252B222AE3FB2BF97E06AF79C5" box="[1198,1428,1679,1700]" gridcol="3" gridrow="7" pageId="3" pageNumber="22">9</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F94606AF79AD" box="[156,1428,1719,1740]" gridrow="8" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F94601A279AD" box="[156,665,1719,1740]" gridcol="0" gridrow="8" pageId="3" pageNumber="22">Oviger articles 6</th>
|
||
<td id="363203252B222AE3FD36F94600BB79AD" box="[691,896,1719,1740]" gridcol="1" gridrow="8" pageId="3" pageNumber="22">6</td>
|
||
<td id="363203252B222AE3FC2EF94607AB79AD" box="[939,1168,1719,1740]" gridcol="2" gridrow="8" pageId="3" pageNumber="22">7 (pseudoarticulation)</td>
|
||
<td id="363203252B222AE3FB2BF94606AF79AD" box="[1198,1428,1719,1740]" gridcol="3" gridrow="8" pageId="3" pageNumber="22">6</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F92E06AF7995" box="[156,1428,1759,1780]" gridrow="9" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F92E01A27995" box="[156,665,1759,1780]" gridcol="0" gridrow="9" pageId="3" pageNumber="22">Relative lengths leg segments Tibia 2>tibia 1>femur</th>
|
||
<td id="363203252B222AE3FD36F92E00BB7995" box="[691,896,1759,1780]" gridcol="1" gridrow="9" pageId="3" pageNumber="22">femur>tibia 2>tibia 1</td>
|
||
<td id="363203252B222AE3FC2EF92E07AB7995" box="[939,1168,1759,1780]" gridcol="2" gridrow="9" pageId="3" pageNumber="22">tibia 2> tibia 1> femur</td>
|
||
<td id="363203252B222AE3FB2BF92E06AF7995" box="[1198,1428,1759,1780]" gridcol="3" gridrow="9" pageId="3" pageNumber="22">tibia 2> femur> tibia 1</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F8F606AF787D" box="[156,1428,1799,1820]" gridrow="10" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F8F601A2787D" box="[156,665,1799,1820]" gridcol="0" gridrow="10" pageId="3" pageNumber="22">cement gland(s) 5–9 cups-mounds</th>
|
||
<td id="363203252B222AE3FD36F8F600BB787D" box="[691,896,1799,1820]" gridcol="1" gridrow="10" pageId="3" pageNumber="22">2 mounds</td>
|
||
<td id="363203252B222AE3FC2EF8F607AB787D" box="[939,1168,1799,1820]" gridcol="2" gridrow="10" pageId="3" pageNumber="22">one mound</td>
|
||
<td id="363203252B222AE3FB2BF8F606AF787D" box="[1198,1428,1799,1820]" gridcol="3" gridrow="10" pageId="3" pageNumber="22">one slit</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F8DE06AF7825" box="[156,1428,1839,1860]" gridrow="11" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F8DE01A27825" box="[156,665,1839,1860]" gridcol="0" gridrow="11" pageId="3" pageNumber="22">Proboscis length 0.39</th>
|
||
<td id="363203252B222AE3FD36F8DE00BB7825" box="[691,896,1839,1860]" gridcol="1" gridrow="11" pageId="3" pageNumber="22">0.54</td>
|
||
<td id="363203252B222AE3FC2EF8DE07AB7825" box="[939,1168,1839,1860]" gridcol="2" gridrow="11" pageId="3" pageNumber="22">0.43</td>
|
||
<td id="363203252B222AE3FB2BF8DE06AF7825" box="[1198,1428,1839,1860]" gridcol="3" gridrow="11" pageId="3" pageNumber="22">0.4</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F8A606AF780D" box="[156,1428,1879,1900]" gridrow="12" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F8A601A2780D" box="[156,665,1879,1900]" gridcol="0" gridrow="12" pageId="3" pageNumber="22">Proboscis width (inflation) 0.135</th>
|
||
<td id="363203252B222AE3FD36F8A600BB780D" box="[691,896,1879,1900]" gridcol="1" gridrow="12" pageId="3" pageNumber="22">-</td>
|
||
<td id="363203252B222AE3FC2EF8A607AB780D" box="[939,1168,1879,1900]" gridcol="2" gridrow="12" pageId="3" pageNumber="22">0.13</td>
|
||
<td id="363203252B222AE3FB2BF8A606AF780D" box="[1198,1428,1879,1900]" gridcol="3" gridrow="12" pageId="3" pageNumber="22">0.16</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F88E06AF78F5" box="[156,1428,1919,1940]" gridrow="13" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F88E01A278F5" box="[156,665,1919,1940]" gridcol="0" gridrow="13" pageId="3" pageNumber="22">Body length 2.1</th>
|
||
<td id="363203252B222AE3FD36F88E00BB78F5" box="[691,896,1919,1940]" gridcol="1" gridrow="13" pageId="3" pageNumber="22">1.71</td>
|
||
<td id="363203252B222AE3FC2EF88E07AB78F5" box="[939,1168,1919,1940]" gridcol="2" gridrow="13" pageId="3" pageNumber="22">1.69 (by addition)</td>
|
||
<td id="363203252B222AE3FB2BF88E06AF78F5" box="[1198,1428,1919,1940]" gridcol="3" gridrow="13" pageId="3" pageNumber="22">2.17</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F85606AF78DD" box="[156,1428,1959,1980]" gridrow="14" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F85601A278DD" box="[156,665,1959,1980]" gridcol="0" gridrow="14" pageId="3" pageNumber="22">Body width (2nd segment) 1.4</th>
|
||
<td id="363203252B222AE3FD36F85600BB78DD" box="[691,896,1959,1980]" gridcol="1" gridrow="14" pageId="3" pageNumber="22">0.89(first)</td>
|
||
<td id="363203252B222AE3FC2EF85607AB78DD" box="[939,1168,1959,1980]" gridcol="2" gridrow="14" pageId="3" pageNumber="22">0.77</td>
|
||
<td id="363203252B222AE3FB2BF85606AF78DD" box="[1198,1428,1959,1980]" gridcol="3" gridrow="14" pageId="3" pageNumber="22">1.13</td>
|
||
</tr>
|
||
<tr id="75E36A592B222AE3FF19F83E06AF7885" box="[156,1428,1999,2020]" gridrow="15" pageId="3" pageNumber="22">
|
||
<th id="363203252B222AE3FF19F83E01A27885" box="[156,665,1999,2020]" gridcol="0" gridrow="15" pageId="3" pageNumber="22">Body segmentation lines nearly obliterated</th>
|
||
<td id="363203252B222AE3FD36F83E00BB7885" box="[691,896,1999,2020]" gridcol="1" gridrow="15" pageId="3" pageNumber="22">3 distinct</td>
|
||
<td id="363203252B222AE3FC2EF83E07AB7885" box="[939,1168,1999,2020]" gridcol="2" gridrow="15" pageId="3" pageNumber="22">3 distinct</td>
|
||
<td id="363203252B222AE3FB2BF83E06AF7885" box="[1198,1428,1999,2020]" gridcol="3" gridrow="15" pageId="3" pageNumber="22">3 distinct</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B25D518FF43FF6900F97E16" blockId="4.[151,1437,152,651]" pageId="4" pageNumber="23">
|
||
<taxonomicName id="0CD313982B25D518FF43FF6902477FD0" ID-CoL="84LFK" box="[198,380,152,177]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9A7B4092B25D518FF43FF6902477FD0" box="[198,380,152,177]" italics="true" pageId="4" pageNumber="23">Anoplodactylus</emphasis>
|
||
</taxonomicName>
|
||
is a relatively morphologically homogeneous genus, and species are usually described based on one or two distinct characters, or more or less unique combinations of characters, mostly of the males. We present a summary of main differences among the four species included in the complex (
|
||
<tableCitation id="86515DA02B25D518FAB4FF1606B27E60" box="[1329,1417,231,257]" captionStart="TABLE 1" captionStartId="3.[151,239,1342,1366]" captionTargetBox="[156,1428,1399,2020]" captionTargetPageId="3" captionText="TABLE 1. Main characters distinguishing the four attenuated Anoplodactylus species in the tenuicorpus complex." httpUri="http://table.plazi.org/id/9FAC38932B22D51FFF12FACF06647A37" pageId="4" pageNumber="23" tableUuid="9FAC38932B22D51FFF12FACF06647A37">Table 1</tableCitation>
|
||
), which are also characters used by
|
||
<bibRefCitation id="AF4215EA2B25D518FDA9FEFF01F37E49" author="Stock" box="[556,712,270,296]" pageId="4" pageNumber="24" refString="Stock, J. H. (1994) Indo-West Pacific Pycnogonida collected by some major oceanographic expeditions. Beaufortia, 44, 17 - 77." type="journal article" year="1994">Stock (1994)</bibRefCitation>
|
||
. These differences might seem minor when defining separate entities or species, but they reflect the nature of variation among
|
||
<taxonomicName id="0CD313982B25D518FC24FEC7076C7E2E" ID-CoL="84LFK" box="[929,1111,310,335]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F9A7B4092B25D518FC24FEC7076C7E2E" box="[929,1111,310,335]" italics="true" pageId="4" pageNumber="23">Anoplodactylus</emphasis>
|
||
</taxonomicName>
|
||
records in general, and the difficulties in defining species boundaries within this speciose genus.
|
||
</paragraph>
|
||
<paragraph id="CB6C681B2B25D518FF43FE75006C7DEA" blockId="4.[151,1437,152,651]" pageId="4" pageNumber="23">
|
||
The
|
||
<taxonomicName id="0CD313982B25D518FF79FE7402BC7EFF" ID-CoL="66X4L" box="[252,391,389,414]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B25D518FF79FE7402BC7EFF" box="[252,391,389,414]" italics="true" pageId="4" pageNumber="23">tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
complex seems related to
|
||
<taxonomicName id="0CD313982B25D518FD3AFE7400E47EFF" ID-CoL="66X3F" box="[703,991,389,414]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="pectinus">
|
||
<emphasis id="F9A7B4092B25D518FD3AFE7400E47EFF" box="[703,991,389,414]" italics="true" pageId="4" pageNumber="23">Anoplodactylus pectinus</emphasis>
|
||
</taxonomicName>
|
||
, a widely distributed pantropical species (
|
||
<bibRefCitation id="AF4215EA2B25D518FF51FE5D02B57EA7" author="Hedgpeth" box="[212,398,428,454]" pageId="4" pageNumber="24" refString="Hedgpeth, J. W. (1948) The Pycnogonida of the Western North Atlantic and the Caribbean. Proceedings of the US National Museum, 98, 157 - 342." type="journal article" year="1948">Hedgpeth 1948</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="AF4215EA2B25D518FE1EFE5D01187EA7" author="Stock" box="[411,547,428,454]" pageId="4" pageNumber="24" refString="Stock, J. H. (1975) Pycnogonids from the continental shelf, slope and deep sea of the tropical Atlantic and East Pacific. Bulletin of Marine Science, 24, 957 - 1092." type="journal article" year="1975">Stock 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="AF4215EA2B25D518FDB5FE5D01E87EA7" author="Bamber" box="[560,723,428,454]" pageId="4" pageNumber="24" refString="Bamber, R. N. (1998) Pycnogonids (Arthropoda: Pycnogonida) from the " Grigore Antipa " National Museum's 1991 Indonesian Expeditions. Travaux du Museum National d'Histoire Naturelle " Grigore Antipa ", 50, 27 - 33." type="journal article" year="1998">Bamber 1998</bibRefCitation>
|
||
; Arango 2003;
|
||
<bibRefCitation id="AF4215EA2B25D518FC0FFE5D07347EA7" author="Child" box="[906,1039,428,454]" pageId="4" pageNumber="24" refString="Child, A. C. (2004) Some Pycnogonida from the Western Caribbean with descriptions of three new species. Bulletin of Marine Science, 74, 143 - 161." type="journal article" year="2004">Child 2004</bibRefCitation>
|
||
and other references therein) that shares morphological traits with the complex such as prolonged chelifores (but not the proboscis) and pectination of the largest spine on the propodal heel. However,
|
||
<taxonomicName id="0CD313982B25D518FCA8FE0A008B7D75" ID-CoL="66X3F" box="[813,944,507,532]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="pectinus">
|
||
<emphasis id="F9A7B4092B25D518FCA8FE0A008B7D75" box="[813,944,507,532]" italics="true" pageId="4" pageNumber="23">A. pectinus</emphasis>
|
||
</taxonomicName>
|
||
is not as slender and lateral processes are not as long as in species of the
|
||
<taxonomicName id="0CD313982B25D518FE7BFDD301B27D5A" ID-CoL="66X4L" box="[510,649,546,571]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B25D518FE7BFDD301B27D5A" box="[510,649,546,571]" italics="true" pageId="4" pageNumber="23">tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
complex. Further material and detailed studies including additional sources of data are needed to understand the distribution patterns and affinities among the species and populations of such rare sea spiders as in the
|
||
<taxonomicName id="0CD313982B25D518FDDCFD8001DF7DEB" ID-CoL="66X4L" box="[601,740,625,650]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="tenuicorpus">
|
||
<emphasis id="F9A7B4092B25D518FDDCFD8001DF7DEB" box="[601,740,625,650]" italics="true" pageId="4" pageNumber="23">tenuicorpus</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |