155 lines
20 KiB
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155 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.913.48650" ID-GBIF-Dataset="81713c0c-9aa1-4a4e-8c5e-ea568c065442" ID-PMC="PMC7044255" ID-Pensoft-Pub="1313-2970-913-1" ID-Pensoft-UUID="2AC899BE84CE53E0B7461D63DA3F0A03" ID-PubMed="32132849" ID-ZooBank="575D3DC1CBF44E99816D0F68F931F927" ModsDocID="1313-2970-913-1" checkinTime="1582137866924" checkinUser="pensoft" docAuthor="Chang, Wan-Jin & Li, Shuqiang" docDate="2020" docId="43548059222E5EF1A49F7B3CBA85CDA9" docLanguage="en" docName="ZooKeys 913: 1-87" docOrigin="ZooKeys 913" docSource="http://dx.doi.org/10.3897/zookeys.913.48650" docTitle="Leclercera jianzuiyu Chang & Li 2020, sp. nov." docType="treatment" docUuid="8294145B-88E3-41DC-9E2F-8B0AB36F6A53" docUuidSource="ZooBank" docVersion="5" id="2AC899BE84CE53E0B7461D63DA3F0A03" lastPageNumber="1" masterDocId="2AC899BE84CE53E0B7461D63DA3F0A03" masterDocTitle="Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="1" updateTime="1732488515709" updateUser="ExternalLinkService">
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<mods:mods id="042AAE934F9E31A5471AD5692D9504BD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="146928189A1F1AA615B924094E8FEAAB">
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<mods:title id="B5EFB82D4F8ED720F96BD749837CA3ED">Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae)</mods:title>
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<mods:name id="80928BAB7C48C1488E4296340183A222" type="personal">
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<mods:namePart id="3F929E336A971ED9E297DF6305A85A54">Chang, Wan-Jin</mods:namePart>
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<mods:namePart id="7DDD0985E7C160A944F752BB93EA0C8F">Li, Shuqiang</mods:namePart>
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<mods:typeOfResource id="24C6EA1FCE29E127C5559EABEB712BB2">text</mods:typeOfResource>
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<mods:titleInfo id="3A85C08DCB19BD733110A4AAA18B26B6">
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<mods:title id="CF48B78F6E797E0CC93CA45853BA1675">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="DD7B0C0310970D75C99C76B713ECD0E8">
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<mods:date id="6C6713CBA7576E5B79CD579F5570961D">2020</mods:date>
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<mods:detail id="D8FDCBB9CA8438EF9C8BC70DABE1A32E" type="volume">
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<mods:number id="EF6C8CE3C74C506484823FF579315802">913</mods:number>
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<mods:start id="E34F9852DDD1B82DCCF333C4491FBADD">1</mods:start>
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<mods:end id="ED28626EDE15197B8CB3DC12CEABAFB0">87</mods:end>
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<mods:url id="4FAF36C29C3938DE9FDF8517F93C849A">http://dx.doi.org/10.3897/zookeys.913.48650</mods:url>
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<mods:classification id="272286891C7033B719227CBB17FFAAD7">journal article</mods:classification>
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<mods:identifier id="5A283A1201C76097162A2369E459AF91" type="DOI">http://dx.doi.org/10.3897/zookeys.913.48650</mods:identifier>
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<mods:identifier id="D06F9D5CAF755130E1B89D5EBFA0C0BF" type="Pensoft-Pub">1313-2970-913-1</mods:identifier>
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<mods:identifier id="300B34A7474D35F6207B1E29BAE27629" type="ZooBank">575D3DC1CBF44E99816D0F68F931F927</mods:identifier>
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<treatment id="43548059222E5EF1A49F7B3CBA85CDA9" ID-GBIF-Taxon="162174150" LSID="urn:lsid:zoobank.org:act:8294145B-88E3-41DC-9E2F-8B0AB36F6A53" httpUri="http://treatment.plazi.org/id/43548059222E5EF1A49F7B3CBA85CDA9" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Psilodercidae" scope_order="Araneae">
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<subSubSection id="DC52801A965A7890E4252B47F8770827" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="4718439FCB621CE2708BF95419EC2BBD" pageId="0" pageNumber="1">
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<taxonomicName id="C9EC0376E4FAAA078902107B8D6A0BCF" ID-CoL="98ZV2" LSID="43548059-222E-5EF1-A49F-7B3CBA85CDA9" authority="Chang & Li, 2020" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera jianzuiyu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jianzuiyu" status="sp. nov.">Leclercera jianzuiyu</taxonomicName>
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<taxonomicNameLabel id="3E1E41917E3D2FCD28DA315E5148ECAE" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="06DFA959E19AE465EA4F98CF45AE36BF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leclercera jianzuiyu sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1">Figs 11</figureCitation>
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<figureCitation id="E7FC59DE9508CB4236E4FA20EF3B3231" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Leclercera jianzuiyu sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure12" httpUri="https://binary.pensoft.net/fig/380954" pageId="0" pageNumber="1">, 12</figureCitation>
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<figureCitation id="13AB7853F5161397191112C6A8151FA5" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Cheliceral retromargin, posterior view. A Leclercera xiaodai sp. nov. B L. zhamensis sp. nov. C L. sanjiao sp. nov. D L. banensis sp. nov. E L. jianzuiyu sp. nov. F L. xiangbabang sp. nov. G L. maochong sp. nov. H L. shanzi sp. nov. I L. yandou sp. nov. J L. suwanensis sp. nov. K L. duandai sp. nov. L L. yanjing sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure55" httpUri="https://binary.pensoft.net/fig/380997" pageId="0" pageNumber="1">, 55E</figureCitation>
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<figureCitation id="6D2CFDA299414C0F240030946B00C3EC" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">, 58</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="360CDAD4A5C75F5C83C941520FDBD5A5" pageId="0" pageNumber="1" type="types">
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<paragraph id="693DC775412E84B925E6FD1C47797B31" pageId="0" pageNumber="1">Types.</paragraph>
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<paragraph id="D0803F296910133C22E3A0DD8FE28519" pageId="0" pageNumber="1">
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<emphasis id="43C8F5C096CD533AE9825455A35FC12C" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: ♂ (IZCAS), Thailand, Prachuap Kiri Khan Province, Hua Hin District, Nong Phlap Subdistrict, Laplae Cave and Kailone Cave,
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<geoCoordinate id="E260A66EE409D73DE62FA972B76DB985" degrees="12" direction="north" minutes="36.2550" orientation="latitude" precision="1" value="12.60425">12°36.2550'N</geoCoordinate>
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,
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<geoCoordinate id="2F29BE00855AA6A5626EC6AF1BEE9F56" degrees="99" direction="east" minutes="43.3410" orientation="longitude" precision="1" value="99.72235">99°43.3410'E</geoCoordinate>
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, elevation ca 175 m, 30.X.2014, H. Zhao, Y. Li and Z. Chen leg.
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<emphasis id="B1BF4801119E3AAF86138F02057EF873" bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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: 1♀ (IZCAS), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="CDA1803FA9E8FE5B9E421F78DDFEDBB9" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="83AB8FB2CE0C302CDAA7A0B0E1CE2B07" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="D11DD51F3741F5E8CF6B8344481AF4DE" pageId="0" pageNumber="1">
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The species name is a noun in apposition derived from the Chinese pinyin
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<normalizedToken id="88D466615587E38CBE155518A4979C04" originalValue="“jiānzuĭyú”">"jianzuĭyu"</normalizedToken>
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(bird wrasse - a type of fish) and refers to the entire structure of bulb which resembles the mouth of a bird wrasse.
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</paragraph>
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</subSubSection>
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<subSubSection id="B31C44460E573DBCD25276E29E1A21DD" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="C8DCC814C68EFA9E1468BD9F286EA1D6" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="3721F53154C6D8E91F2EE5A20A4D9AE7" pageId="0" pageNumber="1">
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Diagnostic features of males and females are discussed in the diagnosis of
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<taxonomicName id="9101C5D63A61AFAF4BEFCDA658B9DFCE" lsidName="L. xiangbabang" pageId="0" pageNumber="1" rank="species" species="xiangbabang">
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<emphasis id="C87C48C3F90DF4A3C501297A93CD8966" italics="true" pageId="0" pageNumber="1">L. xiangbabang</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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<caption id="3F9DF2091A02C5AA7499D165EAB76EE2" doi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
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<paragraph id="7494C380C860F494698315C25EE95445" pageId="0" pageNumber="1">
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<emphasis id="C8298C221277F1475919DF98CE042DCD" bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
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<taxonomicName id="54A09BA949A28C996C7D10044C598F37" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera jianzuiyu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jianzuiyu">
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<emphasis id="3384086EBADDE3CC0B92F85DA2CD65C7" italics="true" pageId="0" pageNumber="1">Leclercera jianzuiyu</emphasis>
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</taxonomicName>
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sp. nov., male holotype and female paratype.
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<emphasis id="40B28EDEBE0DED7BF69E0D20801DB69A" bold="true" pageId="0" pageNumber="1">A</emphasis>
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Endogyne, dorsal view
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<emphasis id="07052DAC19F778EC87A6487152F8ACCD" bold="true" pageId="0" pageNumber="1">B</emphasis>
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female epigastric area, ventral view
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<emphasis id="356A398A9723042320B5A93828364B23" bold="true" pageId="0" pageNumber="1">C</emphasis>
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male habitus, dorsal view
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<emphasis id="F3275B59D24B34A6055A60194C07DCA9" bold="true" pageId="0" pageNumber="1">D</emphasis>
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female habitus, dorsal view
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<emphasis id="17988F3FBCC871519A1AB298FDAD4127" bold="true" pageId="0" pageNumber="1">E</emphasis>
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female habitus, ventral view. Abbreviation: SP = spermatheca.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="62FE8576FA00E854492CF50B99EF06BD" pageId="0" pageNumber="1" type="description">
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<paragraph id="DF6439DD34979A2FF04DBBD20DA91AC2" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="0F6BA2F09C6BE02BB2454CA2971B3D61" pageId="0" pageNumber="1">
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<emphasis id="A4480B00238680F50DCEEED755CA83CF" bold="true" pageId="0" pageNumber="1">Male</emphasis>
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(Holotype). Total length 2.88; carapace 0.63 long, 0.78 wide; abdomen 2.25 long, 0.63 wide. Carapace round and pale yellow, with three longitudinal dark brown bands, median band two times wider than lateral bands (Fig.
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<figureCitation id="AF4560D6DD934A59CAF7D8218A0A1D36" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leclercera jianzuiyu sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1">11C</figureCitation>
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). Chelicerae brown (Fig.
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<figureCitation id="A67C8F221A7C5346A75418F7AE963E8F" captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Cheliceral retromargin, posterior view. A Leclercera xiaodai sp. nov. B L. zhamensis sp. nov. C L. sanjiao sp. nov. D L. banensis sp. nov. E L. jianzuiyu sp. nov. F L. xiangbabang sp. nov. G L. maochong sp. nov. H L. shanzi sp. nov. I L. yandou sp. nov. J L. suwanensis sp. nov. K L. duandai sp. nov. L L. yanjing sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure55" httpUri="https://binary.pensoft.net/fig/380997" pageId="0" pageNumber="1">55E</figureCitation>
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). Clypeus dark brown medially, pale yellow laterally. Endites pale yellow. Labium brown, circular dark brown spot basally. Sternum pale yellow, anterior with dark brown patches laterally, posterior with dark brown band medially. Abdomen elongated, edges of anterior and posterior dark brown, with complex scattered dark brown pattern, antero-ventrally pale yellow with transverse brown band medially, black patches from edges of anterior to posterior. Legs uniformly brown; measurements: I 13.06 (3.75, 0.31, 4.00, 3.60, 1.40), II 8.63 (2.50, 0.25, 2.60, 2.34, 0.94), III missing, IV 9.20 (2.80, 0.20, 2.66, 2.60, 0.94). Palp (Fig.
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<figureCitation id="1FACB44D723A7D8DA8691AAAC7C39215" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Leclercera jianzuiyu sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure12" httpUri="https://binary.pensoft.net/fig/380954" pageId="0" pageNumber="1">12A-D</figureCitation>
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): femur slender, five times longer than patella; patella not swollen; tibia swollen, 1.5 times shorter and two times wider than femur, with retrolateral apophysis anteriorly bearing a spine almost as long as tegulum; cymbium dark brown, 3.5 times shorter than femur; bulb thinly spatulate, pale brown, with embolus arising distally; embolus elongated with slightly bent, blunt tip, half the width of tegulum (Fig.
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<figureCitation id="D75EE6A1FD068B16A33BF7065D31C714" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Leclercera jianzuiyu sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, RA = retrolateral apophysis, SN = spine." figureDoi="10.3897/zookeys.913.48650.figure12" httpUri="https://binary.pensoft.net/fig/380954" pageId="0" pageNumber="1">12B</figureCitation>
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).
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</paragraph>
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<paragraph id="75BD0ADF60721E74D6FC2FD39416E731" pageId="0" pageNumber="1">
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<emphasis id="85C925A12F134B0B701F12A6C96EE66F" bold="true" pageId="0" pageNumber="1">Female</emphasis>
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(Paratype). General features and coloration similar to those of male (Fig.
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<figureCitation id="3ED27A3955C9ED42686763DCDCBC7B92" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leclercera jianzuiyu sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1">11E, F</figureCitation>
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). Measurements: total length 2.00; carapace 0.60 long, 0.70 wide; abdomen 1.40 long, 1.25 wide. Leg measurements: I-II missing, III 4.45 (1.25, 0.20, 1.30, 1.20, 0.50), IV 7.20 (2.19, 0.25, 2.13, 1.88, 0.75). Epigastric area (Fig.
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<figureCitation id="1366BBDAF6CCDF687FAFC1AC152EAEA5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leclercera jianzuiyu sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1">11B</figureCitation>
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): a pair of dark brown horizontal patches laterally. Endogyne (Fig.
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<figureCitation id="741D904620094441F87F8E952C3BB3C2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leclercera jianzuiyu sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure11" httpUri="https://binary.pensoft.net/fig/380953" pageId="0" pageNumber="1">11A</figureCitation>
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): a pair of circular spermathecae with thin strips laterally, wavy patterns in spermathecae, lateral stripes two times longer than circular spermathecae.
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</paragraph>
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</subSubSection>
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<subSubSection id="D7E7A5473F7D8C7A5AB318BA921A9861" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="E6CCA4B2158D905CBDBB21D771AC01C9" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="974564B82A95138B0347E1EA52AEBB73" pageId="0" pageNumber="1">
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Known only from the type locality (Fig.
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<figureCitation id="2997CCCA7A274CB3FA631CB27A511C6B" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">58</figureCitation>
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).
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</paragraph>
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<caption id="3CBDD2CE2D9B4A39637F543039F8BEA4" doi="10.3897/zookeys.913.48650.figure12" httpUri="https://binary.pensoft.net/fig/380954" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
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<paragraph id="2FE1094A1BB497589A49E8AC53955B0D" pageId="0" pageNumber="1">
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<emphasis id="65A6BFC12CD4C9067CFD2BC1FA5E3EC4" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
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<taxonomicName id="EF1BB0C8378386A74FDEC657E6859159" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera jianzuiyu" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jianzuiyu">
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<emphasis id="8186D19330B44C45526FC2CA3F40886F" italics="true" pageId="0" pageNumber="1">Leclercera jianzuiyu</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="1B1624927272AE2BF99DBD586F57C123" bold="true" pageId="0" pageNumber="1">A</emphasis>
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Palp, ventral view
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<emphasis id="183F686C5084581861B73DB47CD806F8" bold="true" pageId="0" pageNumber="1">B</emphasis>
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bulb, ventral view
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<emphasis id="B3010C0F8560EC77D2AEB52C3D1A55A1" bold="true" pageId="0" pageNumber="1">C</emphasis>
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palp, prolateral view
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<emphasis id="96F144918DEEC4378C302924E561C90D" bold="true" pageId="0" pageNumber="1">D</emphasis>
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palp, retrolateral view. Abbreviations: EM = embolus, RA = retrolateral apophysis, SN = spine.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |