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<document id="8CC73DF49695958AF176DB85D33CE0DC" ID-DOI="10.11865/zs.2023101" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941307" ID-ZooBank="7A57D6D4-279E-4A4F-9D9B-2ADF461B5E28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576969620" checkinUser="felipe" docAuthor="Lin, Yejie, Li, Shuqiang &amp; Pham, Dinh-Sac" docDate="2023" docId="431B4B17FFC3FFDFFF7ED419FD39FE37" docLanguage="en" docName="ZoolSyst.48.1.1-99.pdf" docOrigin="Zoological Systematics 48 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8283969" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Lysiteles huyilinae Lin &amp; Li 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="74" masterDocId="BF22336FFF86FF96FFF6D010FFFAFFC0" masterDocTitle="Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam" masterLastPageNumber="99" masterPageNumber="1" pageNumber="70" updateTime="1712578185271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="980ECED8AF88CF4A0F378E9C03F32D4E">Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam</mods:title>
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<mods:name id="D3ECDDA1381AE955FB10AFCAE37DD92D" type="personal">
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<mods:namePart id="8E7C651ACBFF60335700A764214CC76E">Lin, Yejie</mods:namePart>
<mods:affiliation id="A5B4B9AC5181FB27F30A452F6271A0D7">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="BBC731AFF1BA563A7C15A08C70BD79F1">Li, Shuqiang</mods:namePart>
<mods:affiliation id="2CFE084CD68F7715D4FA6DCFBF1B5750">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="A986EABA2EB463F2014E09FF4C023949">Pham, Dinh-Sac</mods:namePart>
<mods:affiliation id="89C7B3325836BA9240FEFFC5C577B1BD">Vietnam National Museum of Nature (VNMN), Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam Corresponding author, E-mail: phamdinhsac @ vnmn. vast. vn</mods:affiliation>
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<mods:title id="22CC843C9D5135C1F21BDD633E5E36C4">Zoological Systematics</mods:title>
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<mods:date id="55ED435D156126987401F2A7CFA882B3">2023</mods:date>
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<treatment id="431B4B17FFC3FFDFFF7ED419FD39FE37" ID-DOI="http://doi.org/10.5281/zenodo.10941646" ID-Zenodo-Dep="10941646" LSID="urn:lsid:plazi:treatment:431B4B17FFC3FFDFFF7ED419FD39FE37" httpUri="http://treatment.plazi.org/id/431B4B17FFC3FFDFFF7ED419FD39FE37" lastPageId="73" lastPageNumber="74" pageId="69" pageNumber="70">
<subSubSection id="83A8A98AFFC3FFD3FF7ED419FCECFBE2" box="[136,790,1033,1058]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="CB0DFA01FFC3FFD3FF7ED419FCECFBE2" blockId="69.[136,790,1033,1058]" box="[136,790,1033,1058]" pageId="69" pageNumber="70">
<heading id="90454D6DFFC3FFD3FF7ED419FCECFBE2" box="[136,790,1033,1058]" centered="true" fontSize="10" level="2" pageId="69" pageNumber="70" reason="2">
<emphasis id="F9C62613FFC3FFD3FF7ED419FDD9FBE2" bold="true" box="[136,547,1033,1058]" pageId="69" pageNumber="70">
<taxonomicName id="0CB28182FFC3FFD3FF7ED419FE38FBE2" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[136,450,1033,1058]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="huyilinae" status="sp. nov.">
<emphasis id="F9C62613FFC3FFD3FF7ED419FEAFFBE1" bold="true" box="[136,341,1033,1057]" italics="true" pageId="69" pageNumber="70">Lysiteles huyilinae</emphasis>
Lin &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="E2F59B68FFC3FFD3FE3BD41AFDD9FBE2" box="[461,547,1034,1058]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5389E684FFC3FFD3FDB6D41AFD41FBE2" box="[576,699,1034,1058]" captionStart-0="Figure 63" captionStart-1="Figure 64" captionStartId-0="73.[155,218,2012,2034]" captionStartId-1="74.[155,218,2022,2044]" captionTargetBox-0="[189,1398,778,1980]" captionTargetBox-1="[189,1398,663,1985]" captionTargetId-0="figure-176@73.[189,1398,777,1980]" captionTargetId-1="figure-240@74.[189,1398,663,1988]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionText-0="Figure 63. Lysiteles huyilinae Lin &amp; Li, sp. nov., holotype male.A. Left palp, ventral view; B. Same, retrolateral view.Abbreviations: C—conductor; E—embolus; RTA—retrolateral tibial apophysis; VTA—ventral tibial apophysis. Scale bars=0.1mm." captionText-1="Figure 64. Lysiteles huyilinae Lin &amp; Li, sp. nov., paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD—copulatory duct; CO—copulatory opening; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.10941497" figureDoi-1="http://doi.org/10.5281/zenodo.10941499" httpUri-0="https://zenodo.org/record/10941497/files/figure.png" httpUri-1="https://zenodo.org/record/10941499/files/figure.png" pageId="69" pageNumber="70">
Figs 63–
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</figureCitation>
,
<figureCitation id="5389E684FFC3FFD3FD30D41AFCF4FBE2" box="[710,782,1034,1058]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="69" pageNumber="70">69E–F</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption id="9FCDAA89FFC3FFD3FF6DD7ACFE1AF7D1" ID-DOI="http://doi.org/10.5281/zenodo.10941483" ID-Zenodo-Dep="10941483" httpUri="https://zenodo.org/record/10941483/files/figure.png" pageId="69" pageNumber="70" startId="69.[155,218,1980,2002]" targetBox="[189,1398,1134,1933]" targetPageId="69" targetType="figure">
<paragraph id="CB0DFA01FFC3FFD3FF6DD7ACFE1AF7D1" blockId="69.[155,1433,1980,2066]" pageId="69" pageNumber="70">
Figure 59.
<taxonomicName id="0CB28182FFC3FFD3FEF5D7ACFDCAF812" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[259,560,1980,2002]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="nanda" status="sp. nov.">
<emphasis id="F9C62613FFC3FFD3FEF5D7ACFE2EF811" box="[259,468,1980,2001]" italics="true" pageId="69" pageNumber="70">Borboropactus nanda</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="F9C62613FFC3FFD3FDCCD7ACFD7CF811" bold="true" box="[570,646,1980,2001]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="E2F59B68FFC3FFD3FDCCD7ACFD7CF811" box="[570,646,1980,2001]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: A—atrium; CD—copulatory duct; CO—copulatory opening; ET—epigynal tooth; FD—fertilization duct; H—hood; S—spermatheca; Se—septum. Scale bars =0.3 mm.
</paragraph>
</caption>
<subSubSection id="83A8A98AFFC0FFDFFF4BD0FCFD39FE37" lastPageId="73" lastPageNumber="74" pageId="70" pageNumber="71" type="etymology">
<paragraph id="CB0DFA01FFC0FFD0FF4BD0FCFBCFFEC4" blockId="70.[136,1452,236,608]" box="[189,1077,236,261]" pageId="70" pageNumber="71">Etymology. The species is named after Ms. Yilin Hu; noun (name) in genitive case.</paragraph>
<paragraph id="CB0DFA01FFC0FFD0FF4BD11FFD18FE37" blockId="70.[136,1452,236,608]" pageId="70" pageNumber="71">
Diagnosis. The new species is similar to
<taxonomicName id="0CB28182FFC0FFD0FD9BD100FB50FEE7" authority="Tang, Yin, Peng, Ubick &amp; Griswold, 2008" authorityName="Tang, Yin, Peng, Ubick &amp; Griswold" authorityYear="2008" box="[621,1194,271,296]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="F9C62613FFC0FFD0FD9BD100FD1FFEE7" box="[621,741,272,295]" italics="true" pageId="70" pageNumber="71">L. arcuatus</emphasis>
Tang, Yin, Peng, Ubick &amp; Griswold, 2008
</taxonomicName>
and
<taxonomicName id="0CB28182FFC0FFD0FB2BD100FEFAFE8A" authority="Song &amp; Zhao, 1994" authorityName="Song &amp; Zhao" authorityYear="1994" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FB2BD100FAA9FEE7" box="[1245,1363,271,295]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
Song &amp; Zhao, 1994
</taxonomicName>
(see
<bibRefCitation id="AF2387F0FFC0FFD0FEC0D122FE12FE8A" author="Tang" box="[310,488,306,330]" pageId="70" pageNumber="71" pagination="1 - 41" refId="ref34962" refString="Tang, G., Yin, C., Peng, X., Ubick, D., Griswold, C. 2008. The crab spiders of the genus Lysiteles from Yunnan Province, China (Araneae: Thomisidae). Zootaxa, 1742: 1 - 41. doi: 10.11646 / zootaxa. 1742.1.1" type="journal article" year="2008">
Tang
<emphasis id="F9C62613FFC0FFD0FE87D123FE5CFE8A" box="[369,422,306,330]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
, 2008
</bibRefCitation>
) by the stick-shaped distal part of the retrolateral tibial apophysis and anticlockwise curved embolus, and the female shares long, curved copulatory ducts and oval spermathecae. However, the new species can be distinguished from
<taxonomicName id="0CB28182FFC0FFD0FEADD168FE29FE4F" authorityName="Song &amp; Zhao" authorityYear="1994" box="[347,467,375,399]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FEADD168FE29FE4F" box="[347,467,375,399]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
by the cybium lacking a terminal apophysis (
<emphasis id="F9C62613FFC0FFD0FC3ED168FC1FFE4F" box="[968,997,376,399]" italics="true" pageId="70" pageNumber="71">vs.</emphasis>
apophysis present in
<taxonomicName id="0CB28182FFC0FFD0FB25D168FAB1FE4F" authorityName="Song &amp; Zhao" authorityYear="1994" box="[1235,1355,375,399]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FB25D168FAB1FE4F" box="[1235,1355,375,399]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
) and the embolic base with an apophysis (
<emphasis id="F9C62613FFC0FFD0FE10D18BFDFEFE72" box="[486,516,411,434]" italics="true" pageId="70" pageNumber="71">vs.</emphasis>
absent in
<taxonomicName id="0CB28182FFC0FFD0FD9BD18BFD19FE72" authorityName="Song &amp; Zhao" authorityYear="1994" box="[621,739,410,434]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FD9BD18BFD19FE72" box="[621,739,410,434]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
). The female can be distinguished from
<taxonomicName id="0CB28182FFC0FFD0FB7AD18BFAFEFE72" authorityName="Tang, Yin, Peng, Ubick &amp; Griswold" authorityYear="2008" box="[1164,1284,411,434]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="F9C62613FFC0FFD0FB7AD18BFAFEFE72" box="[1164,1284,411,434]" italics="true" pageId="70" pageNumber="71">L. arcuatus</emphasis>
</taxonomicName>
and
<taxonomicName id="0CB28182FFC0FFD0FAC0D18BFA51FE72" authorityName="Song &amp; Zhao" authorityYear="1994" box="[1334,1451,410,434]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FAC0D18BFA51FE72" box="[1334,1451,410,434]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
by having the atrium longer than wide (
<emphasis id="F9C62613FFC0FFD0FDB7D1ADFDADFE14" box="[577,599,445,468]" italics="true" pageId="70" pageNumber="71">vs</emphasis>
. wider than long in
<taxonomicName id="0CB28182FFC0FFD0FCCDD1ADFC4EFE14" authorityName="Song &amp; Zhao" authorityYear="1994" box="[827,948,444,468]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FCCDD1ADFC4EFE14" box="[827,948,444,468]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
) and the copulatory ducts with two turns (
<emphasis id="F9C62613FFC0FFD0FA78D1ADFA56FE14" box="[1422,1452,445,468]" italics="true" pageId="70" pageNumber="71">vs.</emphasis>
lacking turns in
<taxonomicName id="0CB28182FFC0FFD0FEC3D1F0FE57FE37" authorityName="Tang, Yin, Peng, Ubick &amp; Griswold" authorityYear="2008" box="[309,429,480,503]" class="Arachnida" family="Araneidae" genus="Araneus" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="F9C62613FFC0FFD0FEC3D1F0FE57FE37" box="[309,429,480,503]" italics="true" pageId="70" pageNumber="71">L. arcuatus</emphasis>
</taxonomicName>
and one turn in
<taxonomicName id="0CB28182FFC0FFD0FDADD1F0FD2BFE37" authorityName="Song &amp; Zhao" authorityYear="1994" box="[603,721,479,503]" class="Arachnida" family="Thomisidae" genus="Lysiteles" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dianicus">
<emphasis id="F9C62613FFC0FFD0FDADD1F0FD2BFE37" box="[603,721,479,503]" italics="true" pageId="70" pageNumber="71">L. dianicus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="CB0DFA01FFC0FFD1FF4BD212FEC5FEE7" blockId="70.[136,1452,236,608]" lastBlockId="71.[136,1452,236,677]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
Description. Male (
<typeStatus id="140944A3FFC0FFD0FE7AD212FE15FDDA" box="[396,495,514,538]" pageId="70" pageNumber="71" type="holotype">holotype</typeStatus>
). Total length 3.03; carapace 1.49 long, 1.32 wide, opisthosoma 1.54 long, 1.11 wide. Eye sizes and interdistances: AME 0.08, ALE 0.13, PME 0.06, PLE 0.10, AME–AME 0.14, AME–ALE 0.15, PME–PME 0.19, PME–PLE 0.28, AME–PME 0.15, ALE–PLE 0.25. Chelicerae with two promarginal and one retromarginal teeth. Leg measurements: I 5.82 (1.70, 2.10, 1.25, 0.77), II 6.06 (1.76, 2.20, 1.32, 0.78), III 3.57 (1.22, 1.22, 0.66, 0.47), IV 3.69 (1.25, 1.29, 0.67, 0.48).
</paragraph>
<caption id="9FCDAA89FFC0FFD0FF6DD816FAEFF7DC" ID-DOI="http://doi.org/10.5281/zenodo.10941487" ID-Zenodo-Dep="10941487" box="[155,1301,2054,2076]" httpUri="https://zenodo.org/record/10941487/files/figure.png" pageId="70" pageNumber="71" startId="70.[155,218,2054,2076]" targetBox="[189,1398,658,2019]" targetPageId="70" targetType="figure">
<paragraph id="CB0DFA01FFC0FFD0FF6DD816FAEFF7DC" blockId="70.[155,1301,2054,2076]" box="[155,1301,2054,2076]" pageId="70" pageNumber="71">
Figure 60.
<taxonomicName id="0CB28182FFC0FFD0FEF2D816FDC8F7DC" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[260,562,2054,2076]" class="Arachnida" family="Thomisidae" genus="Borboropactus" kingdom="Animalia" order="Araneae" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="nanda" status="sp. nov.">
<emphasis id="F9C62613FFC0FFD0FEF2D816FE2FF7DB" box="[260,469,2054,2075]" italics="true" pageId="70" pageNumber="71">Borboropactus nanda</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="F9C62613FFC0FFD0FDCAD817FD73F7DC" bold="true" box="[572,649,2055,2076]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="E2F59B68FFC0FFD0FDCAD817FD73F7DC" box="[572,649,2055,2076]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, alive. A. Holotype male; B. Paratype female. Photoed by Fan Gao.
</paragraph>
</caption>
<paragraph id="CB0DFA01FFC1FFD1FF4BD122FECCFE72" blockId="71.[136,1452,236,677]" pageId="71" pageNumber="72">
Coloration (
<figureCitation id="5389E684FFC1FFD1FEC8D122FE64FE8A" box="[318,414,306,330]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="71" pageNumber="72">Fig. 69E</figureCitation>
). Carapace yellow, two brown lines across carapace, with sparse, long black setae. Chelicera dark brown. Endites and labium brown, sternum yellow. Legs paler yellow to yellow-brown, covered with black setae. Opisthosoma pentagonal, paler yellow with symmetrical irregular black pattern, covered with sparse, long black setae, ventrum yellow.
</paragraph>
<paragraph id="CB0DFA01FFC1FFD1FF4BD1ACFB8BFDFD" blockId="71.[136,1452,236,677]" pageId="71" pageNumber="72">
Palp (
<figureCitation id="5389E684FFC1FFD1FF0BD1ACFEB4FE14" box="[253,334,444,468]" captionStart="Figure 63" captionStartId="73.[155,218,2012,2034]" captionTargetBox="[189,1398,778,1980]" captionTargetId="figure-176@73.[189,1398,777,1980]" captionTargetPageId="73" captionText="Figure 63. Lysiteles huyilinae Lin &amp; Li, sp. nov., holotype male.A. Left palp, ventral view; B. Same, retrolateral view.Abbreviations: C—conductor; E—embolus; RTA—retrolateral tibial apophysis; VTA—ventral tibial apophysis. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941497" httpUri="https://zenodo.org/record/10941497/files/figure.png" pageId="71" pageNumber="72">Fig. 63</figureCitation>
). Patella almost as long as tibia. Tibia with one dorsal macroseta. Ventral tibial apophysis club shaped, terminus expanded, almost straight. Retrolateral tibial apophysis almost two times longer than ventral tibial apophysis, terminus bifurcated. Dorsal tibial apophysis absent. Tegulum flat, disk shaped. Conductor lamellate. Embolus spiraled, thin, base arising at clock 11:00 position, ratio of length of embolus to length of embolus base 6: 1.
</paragraph>
<paragraph id="CB0DFA01FFC1FFDFFF4BD257FB79FEC5" blockId="71.[136,1452,236,677]" lastBlockId="73.[136,1452,236,504]" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72">Female (IZCAS-Ar44012). Total length 3.66; carapace 1.38 long, 1.25 wide, opisthosoma 2.28 long, 2.06 wide. Eye sizes and interdistances: AME 0.08, ALE 0.13, PME 0.04, PLE 0.10, AME–AME 0.15, AME–ALE 0.15, PME–PME 0.23, PME–PLE 0.30, AME–PME 0.17, ALE–PLE 0.25. Chelicerae as in male. Leg measurements: I 4.42 (1.38, 1.59, 0.86, 0.59), II 4.58 (1.48, 1.58, 0.84, 0.68), III 3.06 (1.05, 1.06, 0.52, 0.43), IV 3.29 (1.12, 1.16, 0.60, 0.41).</paragraph>
<caption id="9FCDAA89FFC1FFD1FF6DD7DAFEA4F7E0" ID-DOI="http://doi.org/10.5281/zenodo.10941489" ID-Zenodo-Dep="10941489" httpUri="https://zenodo.org/record/10941489/files/figure.png" pageId="71" pageNumber="72" startId="71.[155,218,1994,2016]" targetBox="[189,1399,736,1946]" targetPageId="71" targetType="figure">
<paragraph id="CB0DFA01FFC1FFD1FF6DD7DAFEA4F7E0" blockId="71.[155,1434,1994,2080]" pageId="71" pageNumber="72">
Figure 61.
<taxonomicName id="0CB28182FFC1FFD1FEF4D7DAFDE2F820" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[258,536,1994,2016]" class="Arachnida" family="Thomisidae" genus="Cozyptila" kingdom="Animalia" order="Araneae" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="haocongi" status="sp. nov.">
<emphasis id="F9C62613FFC1FFD1FEF4D7DAFE46F81F" box="[258,444,1994,2015]" italics="true" pageId="71" pageNumber="72">Cozyptila haocongi</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="F9C62613FFC1FFD1FDD6D7DBFD96F820" bold="true" box="[544,620,1995,2016]" pageId="71" pageNumber="72">
<taxonomicNameLabel id="E2F59B68FFC1FFD1FDD6D7DBFD96F820" box="[544,620,1995,2016]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A. Left palp, ventral view; B. Same, retrolateral view. Abbreviations: E—embolus; ITA—intermediate tibial apophysis; RTC—retrolateral tibial claw; Tu—tutaculum; VTA—ventral tibial apophysis. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="9FCDAA89FFCEFFDEFF6DD7FAFB3CF7E0" ID-DOI="http://doi.org/10.5281/zenodo.10941493" ID-Zenodo-Dep="10941493" httpUri="https://zenodo.org/record/10941493/files/figure.png" pageId="72" pageNumber="73" startId="72.[155,218,2026,2048]" targetBox="[189,1399,252,1991]" targetPageId="72" targetType="figure">
<paragraph id="CB0DFA01FFCEFFDEFF6DD7FAFB3CF7E0" blockId="72.[155,1433,2026,2080]" pageId="72" pageNumber="73">
Figure 62.
<taxonomicName id="0CB28182FFCEFFDEFEF0D7FAFDD8F7C0" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[262,546,2026,2048]" class="Arachnida" family="Thomisidae" genus="Cozyptila" kingdom="Animalia" order="Araneae" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="haocongi" status="sp. nov.">
<emphasis id="F9C62613FFCEFFDEFEF0D7FAFE3BF83F" box="[262,449,2026,2047]" italics="true" pageId="72" pageNumber="73">Cozyptila haocongi</emphasis>
Lin &amp; Li
</taxonomicName>
,
<emphasis id="F9C62613FFCEFFDEFDDAD7FAFD80F83F" bold="true" box="[556,634,2026,2047]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="E2F59B68FFCEFFDEFDDAD7FAFD80F83F" box="[556,634,2026,2047]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD—copulatory duct; CO—copulatory opening; FD—fertilization duct; S—spermatheca. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="CB0DFA01FFCFFFDFFF4BD11FFCD4FEE7" blockId="73.[136,1452,236,504]" box="[189,814,271,295]" pageId="73" pageNumber="74">
Coloration (
<figureCitation id="5389E684FFCFFFDFFEC8D11FFE64FEE7" box="[318,414,271,295]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="73" pageNumber="74">Fig. 69F</figureCitation>
). Similar to that of male except paler.
</paragraph>
<paragraph id="CB0DFA01FFCFFFDFFF4BD122FCE1FE4F" blockId="73.[136,1452,236,504]" pageId="73" pageNumber="74">
Epigyne (
<figureCitation id="5389E684FFCFFFDFFEDED122FE80FE8A" box="[296,378,306,330]" captionStart="Figure 64" captionStartId="74.[155,218,2022,2044]" captionTargetBox="[189,1398,663,1985]" captionTargetId="figure-240@74.[189,1398,663,1988]" captionTargetPageId="74" captionText="Figure 64. Lysiteles huyilinae Lin &amp; Li, sp. nov., paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD—copulatory duct; CO—copulatory opening; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941499" httpUri="https://zenodo.org/record/10941499/files/figure.png" pageId="73" pageNumber="74">Fig. 64</figureCitation>
). Epigynal plate longer than wide. Atrium wide. Copulatory opening large, almost 0.5 of epigyne length. Copulatory ducts long, with four turns. Spermathecae oval, two times longer than fertilization ducts. Fertilization ducts directed toward clock 3:00 position from spermathecae.
</paragraph>
<paragraph id="CB0DFA01FFCFFFDFFF4BD18AFB2AFE14" blockId="73.[136,1452,236,504]" pageId="73" pageNumber="74">
<materialsCitation id="7BDAF05CFFCFFFDFFF4BD18AFB36FE14" collectingDate="2005-05-20" collectorName="Dinh-Sac Pham" country="Vietnam" county="Material" location="Natural Forest" municipality="Tam Dao National Park" pageId="73" pageNumber="74" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Vinh Phuc" typeStatus="holotype">
<collectingCounty id="226C828DFFCFFFDFFF4BD18AFEEDFE72" box="[189,279,410,434]" pageId="73" pageNumber="74">Material</collectingCounty>
examined.
<typeStatus id="140944A3FFCFFFDFFE60D18AFE03FE72" box="[406,505,410,434]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
<specimenCount id="DDB43188FFCFFFDFFDF7D18AFDEFFE71" box="[513,533,410,433]" count="1" pageId="73" pageNumber="74" type="male">♂</specimenCount>
(IZCAS-Ar43999),
<collectingCountry id="B3A5BA91FFCFFFDFFD07D18AFCB5FE72" box="[753,847,410,434]" name="Vietnam" pageId="73" pageNumber="74">Vietnam</collectingCountry>
:
<collectingRegion id="097634E3FFCFFFDFFCACD18AFC34FE72" box="[858,974,410,434]" country="Vietnam" name="Vinh Phuc" pageId="73" pageNumber="74">Vinh Phuc</collectingRegion>
,
<collectingMunicipality id="2B69607BFFCFFFDFFC2FD18AFB26FE72" box="[985,1244,410,434]" pageId="73" pageNumber="74">Tam Dao National Park</collectingMunicipality>
,
<location id="CE6DACDAFFCFFFDFFB1ED18AFA7FFE71" LSID="urn:lsid:plazi:treatment:431B4B17FFC3FFDFFF7ED419FD39FE37:CE6DACDAFFCFFFDFFB1ED18AFA7FFE71" box="[1256,1413,410,434]" country="Vietnam" county="Material" municipality="Tam Dao National Park" name="Natural Forest" pageId="73" pageNumber="74" stateProvince="Vinh Phuc">Natural Forest</location>
,
<date id="BF0CDCC1FFCFFFDFFA67D18AFF02FE14" pageId="73" pageNumber="74" value="2005-05-20">
<collectingDate id="AF482529FFCFFFDFFA67D18AFF02FE14" pageId="73" pageNumber="74" value="2005-05-20">20 May 2005</collectingDate>
</date>
,
<collectorName id="66479FD7FFCFFFDFFEF4D1ACFE51FE14" box="[258,427,444,468]" pageId="73" pageNumber="74">Dinh-Sac Pham</collectorName>
leg.
<typeStatus id="140944A3FFCFFFDFFE29D1ADFDB0FE15" box="[479,586,445,469]" pageId="73" pageNumber="74" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="DDB43188FFCFFFDFFDA2D1ACFD8DFE14" box="[596,631,444,468]" count="9" pageId="73" pageNumber="74" type="male">9♂</specimenCount>
<specimenCount id="DDB43188FFCFFFDFFD8ED1ACFD6DFE14" box="[632,663,444,468]" count="4" pageId="73" pageNumber="74" type="female">4♀</specimenCount>
(IZCAS-Ar44000–Ar44012), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="CB0DFA01FFCFFFDFFF4BD1CFFD39FE37" blockId="73.[136,1452,236,504]" box="[189,707,479,504]" pageId="73" pageNumber="74">
Distribution. Known only from the
<typeStatus id="140944A3FFCFFFDFFDCED1F0FD9CFE38" box="[568,614,480,504]" pageId="73" pageNumber="74">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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