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<document id="93BA4C35756B0D67AB8F2665BAF087D8" ID-DOI="10.11865/zs.2023101" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941307" ID-ZooBank="7A57D6D4-279E-4A4F-9D9B-2ADF461B5E28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576969620" checkinUser="felipe" docAuthor="Lin, Yejie, Li, Shuqiang &amp; Pham, Dinh-Sac" docDate="2023" docId="431B4B17FFC2FFD3FF7ED6CBFD39FC92" docLanguage="en" docName="ZoolSyst.48.1.1-99.pdf" docOrigin="Zoological Systematics 48 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8283969" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Cozyptila haocongi Lin &amp; Li 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="70" masterDocId="BF22336FFF86FF96FFF6D010FFFAFFC0" masterDocTitle="Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam" masterLastPageNumber="99" masterPageNumber="1" pageNumber="69" updateTime="1712578185271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="4499B4D07EDFD9E6BDFDE41056C90568">
<mods:title id="AF8797BB2678BA4C511CEAD563B39835">Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam</mods:title>
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<mods:name id="4F03D5B3B490EE886112B74592C7FE9F" type="personal">
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<mods:namePart id="214C6D0DAD6536A4C69DF1855E38866C">Lin, Yejie</mods:namePart>
<mods:affiliation id="7D48E978F12AC90ECA68325BC553910F">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="3F1EF9087A4A8862CA1EBCA11449BE12">Li, Shuqiang</mods:namePart>
<mods:affiliation id="7C4DF3EA53B6E8FE20CA952661D356BB">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="2BEF05D3092EFD1341F55B3763622CFF">Pham, Dinh-Sac</mods:namePart>
<mods:affiliation id="BAEF2F78637B11A44D872B5DB5E2B8F5">Vietnam National Museum of Nature (VNMN), Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam Corresponding author, E-mail: phamdinhsac @ vnmn. vast. vn</mods:affiliation>
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<mods:title id="A35E2D9AACFD1AE4FBA5AEB072268531">Zoological Systematics</mods:title>
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<mods:part id="242E4FD89977954B03834CB977609C10">
<mods:date id="F3505D7AFCDBA80DCEB1C6AB9F4553B1">2023</mods:date>
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<mods:identifier id="37085897192640F21EB5BF8D5895E5F5" type="ISSN">2095-6827</mods:identifier>
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<treatment id="431B4B17FFC2FFD3FF7ED6CBFD39FC92" ID-DOI="http://doi.org/10.5281/zenodo.10941630" ID-Zenodo-Dep="10941630" LSID="urn:lsid:plazi:treatment:431B4B17FFC2FFD3FF7ED6CBFD39FC92" httpUri="http://treatment.plazi.org/id/431B4B17FFC2FFD3FF7ED6CBFD39FC92" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<subSubSection id="83A8A98AFFC2FFD2FF7ED6CBFCD8F933" box="[136,802,1755,1779]" pageId="68" pageNumber="69" type="nomenclature">
<paragraph id="CB0DFA01FFC2FFD2FF7ED6CBFCD8F933" blockId="68.[136,802,1755,1779]" box="[136,802,1755,1779]" pageId="68" pageNumber="69">
<heading id="90454D6DFFC2FFD2FF7ED6CBFCD8F933" box="[136,802,1755,1779]" centered="true" fontSize="10" level="2" pageId="68" pageNumber="69" reason="2">
<emphasis id="F9C62613FFC2FFD2FF7ED6CBFDD0F933" bold="true" box="[136,554,1755,1779]" pageId="68" pageNumber="69">
<taxonomicName id="0CB28182FFC2FFD2FF7ED6CBFE33F933" authority="Lin &amp; Li" authorityName="Lin &amp; Li" authorityYear="2023" box="[136,457,1755,1779]" class="Arachnida" family="Thomisidae" genus="Cozyptila" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="haocongi" status="sp. nov.">
<emphasis id="F9C62613FFC2FFD2FF7ED6CBFEA1F933" bold="true" box="[136,347,1755,1779]" italics="true" pageId="68" pageNumber="69">Cozyptila haocongi</emphasis>
Lin &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="E2F59B68FFC2FFD2FE22D6CBFDD0F933" box="[468,554,1755,1779]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5389E684FFC2FFD2FDB0D6CBFD3BF933" box="[582,705,1755,1779]" captionStart-0="Figure 61" captionStart-1="Figure 62" captionStartId-0="71.[155,218,1994,2016]" captionStartId-1="72.[155,218,2026,2048]" captionTargetBox-0="[189,1399,736,1946]" captionTargetBox-1="[189,1399,252,1991]" captionTargetId-0="figure-249@71.[189,1399,731,1948]" captionTargetId-1="figure-51@72.[189,1399,252,1991]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="Figure 61. Cozyptila haocongi Lin &amp; Li, sp. nov., holotype male.A. Left palp, ventral view; B. Same, retrolateral view.Abbreviations: E—embolus; ITA—intermediate tibial apophysis; RTC—retrolateral tibial claw; Tu—tutaculum; VTA—ventral tibial apophysis. Scale bars=0.1mm." captionText-1="Figure 62. Cozyptila haocongi Lin &amp; Li, sp. nov., paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD—copulatory duct; CO—copulatory opening; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.10941489" figureDoi-1="http://doi.org/10.5281/zenodo.10941493" httpUri-0="https://zenodo.org/record/10941489/files/figure.png" httpUri-1="https://zenodo.org/record/10941493/files/figure.png" pageId="68" pageNumber="69">Figs 61–62</figureCitation>
,
<figureCitation id="5389E684FFC2FFD2FD3AD6CBFCE0F933" box="[716,794,1755,1779]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="68" pageNumber="69">69C–D</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="83A8A98AFFC2FFD3FF4BD703FD39FC92" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="etymology">
<paragraph id="CB0DFA01FFC2FFD2FF4BD703FB89F8EA" blockId="68.[136,1452,1811,2077]" box="[189,1139,1811,1835]" pageId="68" pageNumber="69">Etymology. The species is named after Mr. Haocong Yang; noun (name) in genitive case.</paragraph>
<paragraph id="CB0DFA01FFC2FFD2FF4BD725FE09F853" blockId="68.[136,1452,1811,2077]" pageId="68" pageNumber="69">Diagnosis. Males of the new species can be distinguished from all congeners by having the ventral tibial apophysis strongly curved and the dorsal tibial apophysis lacking a hook. The female can be distinguished from all congeners by the long and curved copulatory ducts.</paragraph>
<paragraph id="CB0DFA01FFC2FFD2FF4BD78DFC4FF7DD" blockId="68.[136,1452,1811,2077]" pageId="68" pageNumber="69">
Description. Male (
<typeStatus id="140944A3FFC2FFD2FE7AD78DFE15F875" box="[396,495,1949,1973]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
). Total length 3.11; carapace 1.68 long, 1.51 wide, opisthosoma 1.43 long, 1.33 wide. Eye sizes and interdistances: AME 0.06, ALE 0.13, PME 0.05, PLE 0.09, AME–AME 0.20, AME–ALE 0.12, PME–PME 0.18, PME–PLE 0.29, AME–PME 0.19, ALE–PLE 0.19. Leg measurements: I 4.88 (1.45, 1.75, 0.93, 0.75), II 4.92 (1.50, 1.78, 0.98, 0.66), III 3.39 (1.10, 1.20, 0.56, 0.53), IV 3.41 (1.12, 1.19, 0.58, 0.52).
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD0FCFC44FE8A" blockId="69.[136,1452,236,850]" pageId="69" pageNumber="70">
Coloration (
<figureCitation id="5389E684FFC3FFD3FEC8D0FCFE65FEC4" box="[318,415,236,260]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="69" pageNumber="70">Fig. 69C</figureCitation>
). Carapace dark brown, covered with clavate setae. Chelicerae, endites and labium yellow-brown. Sternum brown. Legs yellow-brown with brown irregular patterns. Opisthosoma pentagonal, yellow-brown, covered with clavate setae, with irregular black and white spots. Spinnerets yellow-brown.
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD144FC4FFDFC" blockId="69.[136,1452,236,850]" pageId="69" pageNumber="70">
Palp (
<figureCitation id="5389E684FFC3FFD3FF08D144FEAAFEAC" box="[254,336,340,364]" captionStart="Figure 61" captionStartId="71.[155,218,1994,2016]" captionTargetBox="[189,1399,736,1946]" captionTargetId="figure-249@71.[189,1399,731,1948]" captionTargetPageId="71" captionText="Figure 61. Cozyptila haocongi Lin &amp; Li, sp. nov., holotype male.A. Left palp, ventral view; B. Same, retrolateral view.Abbreviations: E—embolus; ITA—intermediate tibial apophysis; RTC—retrolateral tibial claw; Tu—tutaculum; VTA—ventral tibial apophysis. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941489" httpUri="https://zenodo.org/record/10941489/files/figure.png" pageId="69" pageNumber="70">Fig. 61</figureCitation>
). Patella almost as long as tibia, with one dorsal macroseta. Tibia with three apophyses, ventral tibial apophysis club-shaped, terminus expanded, strongly curved, intermediate apophysis shorter than ventral one. Retrolateral tibial claw as long as ventral tibial apophysis, terminally expanded and flat. Tegulum flat, disk-shaped. Embolus coiled. Female (
<specimenCount id="DDB43188FFC3FFD3FF10D1ACFE88FE15" box="[230,370,444,469]" count="1" pageId="69" pageNumber="70" type="generic" typeStatus="paratype">one paratype</specimenCount>
Ar43998). Total length 3.87; carapace 1.77 long, 1.66 wide, opisthosoma 2.10 long, 2.11 wide. Eye sizes and interdistances: AME 0.06, ALE 0.13, PME 0.05, PLE 0.10, AME–AME 0.23, AME–ALE 0.14, PME–PME 0.20, PME–PLE 0.33, AME–PME 0.23, ALE–PLE 0.21. Leg measurements: I 4.98 (1.47, 1.82, 0.96, 0.73), II 4.87 (1.53, 1.67, 0.93, 0.74), III 3.39 (1.10, 1.22, 0.53, 0.54), IV 3.40 (1.13, 1.12, 0.65, 0.50).
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD257FCC8FD9F" blockId="69.[136,1452,236,850]" box="[189,818,583,607]" pageId="69" pageNumber="70">
Coloration (
<figureCitation id="5389E684FFC3FFD3FEC8D257FE58FD9F" box="[318,418,583,607]" captionStart="Figure 69" captionStartId="80.[155,218,1991,2013]" captionTargetBox="[189,1398,939,1941]" captionTargetId="figure-241@80.[189,1398,938,1942]" captionTargetPageId="80" captionText="Figure 69. Habitus, dorsal view, male (A, C, E, G, I) and female (B, D, F, H). A–B. Borboropactus nanda Lin &amp; Li, sp. nov.; C–D. Cozyptila haocongi Lin &amp; Li, sp. nov.; E–F. Lysiteles huyilinae Lin &amp; Li, sp. nov.; G–H. Massuria daizong Lin &amp; Li, sp. nov.; I. Zametopina wanliae Lin &amp; Li, sp. nov. Scale bars: A–B, H=1.0mm; C–G, I=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.10941519" httpUri="https://zenodo.org/record/10941519/files/figure.png" pageId="69" pageNumber="70">Fig. 69D</figureCitation>
). Similar to that of male except paler.
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD27AFEDBFD07" blockId="69.[136,1452,236,850]" pageId="69" pageNumber="70">
Epigyne (
<figureCitation id="5389E684FFC3FFD3FED3D27AFE8FFD42" box="[293,373,618,642]" captionStart="Figure 62" captionStartId="72.[155,218,2026,2048]" captionTargetBox="[189,1399,252,1991]" captionTargetId="figure-51@72.[189,1399,252,1991]" captionTargetPageId="72" captionText="Figure 62. Cozyptila haocongi Lin &amp; Li, sp. nov., paratype female. A. Epigyne, ventral view; B. Vulva, dorsal view. Abbreviations: CD—copulatory duct; CO—copulatory opening; FD—fertilization duct; S—spermatheca. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.10941493" httpUri="https://zenodo.org/record/10941493/files/figure.png" pageId="69" pageNumber="70">Fig. 62</figureCitation>
). Epigynal plate oval. Atrium wide. Copulatory openings inconspicuous. Copulatory ducts long, with two turns. Spermathecae oval. Fertilization ducts as long as spermathecae, directed toward clock 2:30 position from spermathecae.
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD2C2FE0AFCEF" blockId="69.[136,1452,236,850]" pageId="69" pageNumber="70">
<materialsCitation id="7BDAF05CFFC3FFD3FF4BD2C2FE16FCEF" collectingDate="2007-06-13" collectorName="Dinh-Sac Pham" country="Vietnam" county="Material" elevation="1084" latitude="21.4587" location="Disturbed Forest" longLatPrecision="7" longitude="105.6419" municipality="Tam Dao National Park" pageId="69" pageNumber="70" specimenCount="21" specimenCount-female="10" specimenCount-male="11" stateProvince="Vinh Phuc" typeStatus="holotype">
<collectingCounty id="226C828DFFC3FFD3FF4BD2C2FEEDFD2A" box="[189,279,722,746]" pageId="69" pageNumber="70">Material</collectingCounty>
examined.
<typeStatus id="140944A3FFC3FFD3FE61D2C2FE00FD2A" box="[407,506,722,746]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
<specimenCount id="DDB43188FFC3FFD3FDF5D2C2FDEDFD29" box="[515,535,722,745]" count="1" pageId="69" pageNumber="70" type="male">♂</specimenCount>
(IZCAS-Ar43978),
<collectingCountry id="B3A5BA91FFC3FFD3FD03D2C2FCA9FD2A" box="[757,851,722,746]" name="Vietnam" pageId="69" pageNumber="70">Vietnam</collectingCountry>
:
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,
<collectingMunicipality id="2B69607BFFC3FFD3FC29D2C2FB1EFD2A" box="[991,1252,722,746]" pageId="69" pageNumber="70">Tam Dao National Park</collectingMunicipality>
,
<location id="CE6DACDAFFC3FFD3FB06D2C2FA5DFD29" LSID="urn:lsid:plazi:treatment:431B4B17FFC2FFD3FF7ED6CBFD39FC92:CE6DACDAFFC3FFD3FB06D2C2FA5DFD29" box="[1264,1447,722,746]" country="Vietnam" county="Material" latitude="21.4587" longLatPrecision="7" longitude="105.6419" municipality="Tam Dao National Park" name="Disturbed Forest" pageId="69" pageNumber="70" stateProvince="Vinh Phuc">Disturbed Forest</location>
,
<location id="CE6DACDAFFC3FFD3FF7ED2E4FF26FCCC" LSID="urn:lsid:plazi:treatment:431B4B17FFC2FFD3FF7ED6CBFD39FC92:CE6DACDAFFC3FFD3FF7ED2E4FF26FCCC" box="[136,220,756,780]" country="Vietnam" county="Material" latitude="21.4587" longLatPrecision="7" longitude="105.6419" municipality="Tam Dao National Park" name="Sieving" pageId="69" pageNumber="70" stateProvince="Vinh Phuc">Sieving</location>
, (
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,
<geoCoordinate id="AE869CC6FFC3FFD3FE86D2E4FE08FCCC" box="[368,498,756,780]" degrees="105.6419" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="105.6419">105.6419°E</geoCoordinate>
, elev.
<quantity id="0C4A57E4FFC3FFD3FDC5D2E4FD79FCCD" box="[563,643,756,781]" metricMagnitude="3" metricUnit="m" metricValue="1.084" pageId="69" pageNumber="70" unit="m" value="1084.0">
<elevation id="409F1D32FFC3FFD3FDC5D2E4FD79FCCD" box="[563,643,756,781]" metricMagnitude="3" metricUnit="m" metricValue="1.084" pageId="69" pageNumber="70" unit="m" value="1084.0">1084 m</elevation>
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),
<date id="BF0CDCC1FFC3FFD3FD63D2E5FCDFFCCD" box="[661,805,757,781]" pageId="69" pageNumber="70" value="2007-06-13">
<collectingDate id="AF482529FFC3FFD3FD63D2E5FCDFFCCD" box="[661,805,757,781]" pageId="69" pageNumber="70" value="2007-06-13">13 June 2007</collectingDate>
</date>
,
<collectorName id="66479FD7FFC3FFD3FCD8D2E4FC2CFCCC" box="[814,982,756,780]" pageId="69" pageNumber="70">Dinh-Sac Pham</collectorName>
leg.
<typeStatus id="140944A3FFC3FFD3FBFFD2E5FB8EFCCD" box="[1033,1140,757,781]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="DDB43188FFC3FFD3FB8BD2E5FB55FCCD" box="[1149,1199,757,781]" count="10" pageId="69" pageNumber="70" type="male">10♂</specimenCount>
<specimenCount id="DDB43188FFC3FFD3FB59D2E5FB20FCCD" box="[1199,1242,757,781]" count="10" pageId="69" pageNumber="70" type="female">10♀</specimenCount>
(IZCAS-Ar43979– Ar43998), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="CB0DFA01FFC3FFD3FF4BD32AFD39FC92" blockId="69.[136,1452,236,850]" box="[189,707,826,850]" pageId="69" pageNumber="70">
Distribution. Known only from the
<typeStatus id="140944A3FFC3FFD3FDCFD32AFD9DFC92" box="[569,615,826,850]" pageId="69" pageNumber="70">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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