treatments-xml/data/43/13/67/43136755FA255D4387FC0163974AEAFF.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.893.47059" ID-GBIF-Dataset="98b77e11-c729-4d68-b823-60276bae22d5" ID-PMC="PMC6901613" ID-Pensoft-Pub="1313-2970-893-125" ID-Pensoft-UUID="F7454724568F5B6F841A9B7DA9F8172C" ID-PubMed="31844403" ID-ZooBank="8EED21A97B7B405D8D2F1691405F4F75" ModsDocID="1313-2970-893-125" checkinTime="1575325050564" checkinUser="pensoft" docAuthor="Duran, Daniel P. &amp; Roman, Stephen J." docDate="2019" docId="43136755FA255D4387FC0163974AEAFF" docLanguage="en" docName="ZooKeys 893: 125-134" docOrigin="ZooKeys 893" docSource="http://dx.doi.org/10.3897/zookeys.893.47059" docTitle="Amblycheila katzi Roman &amp; Duran, sp. nov." docType="treatment" docUuid="F2A07537-21DD-4371-A52F-2BAB80E8EF84" docUuidSource="ZooBank" docVersion="5" id="F7454724568F5B6F841A9B7DA9F8172C" lastPageNumber="125" masterDocId="F7454724568F5B6F841A9B7DA9F8172C" masterDocTitle="A new petrophilous tiger beetle from the Trans-Pecos region of Texas and revised key to the genus Amblycheila (Coleoptera, Carabidae, Cicindelinae)" masterLastPageNumber="134" masterPageNumber="125" pageNumber="125" updateTime="1668168153744" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new petrophilous tiger beetle from the Trans-Pecos region of Texas and revised key to the genus Amblycheila (Coleoptera, Carabidae, Cicindelinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Duran, Daniel P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Roman, Stephen J.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>893</mods:number>
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<mods:start>125</mods:start>
<mods:end>134</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.893.47059</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.893.47059</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-893-125</mods:identifier>
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<treatment ID-GBIF-Taxon="160612164" LSID="urn:lsid:zoobank.org:act:F2A07537-21DD-4371-A52F-2BAB80E8EF84" httpUri="http://treatment.plazi.org/id/43136755FA255D4387FC0163974AEAFF" lastPageNumber="125" pageId="0" pageNumber="125">
<subSubSection pageId="0" pageNumber="125" type="nomenclature">
<paragraph pageId="0" pageNumber="125">
<taxonomicName LSID="43136755-FA25-5D43-87FC-0163974AEAFF" authority="Roman &amp; Duran" class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">Amblycheila katzi Roman &amp; Duran</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="125">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Amblycheila katzi Duran &amp; Roman, sp. nov. dorsal habitus (female)." figureDoi="10.3897/zookeys.892.47059.figure1" httpUri="https://binary.pensoft.net/fig/361370" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure1">Figs 1</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amblycheila katzi Duran &amp; Roman, sp. nov. frontal habitus (female)." figureDoi="10.3897/zookeys.892.47059.figure2" httpUri="https://binary.pensoft.net/fig/361371" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure2">2</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat for Amblycheila katzi sp. nov. A steep-sided limestone road cut B rock ledges with grooves and cracks C rock ledge with calcareous soil and third instar A. katzi larval burrow D closeup of third instar A. katzi larval burrow opening." figureDoi="10.3897/zookeys.892.47059.figure3" httpUri="https://binary.pensoft.net/fig/361372" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure3">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="materials_examined">
<paragraph pageId="0" pageNumber="125">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Holotype</emphasis>
: 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas / Val Verde Co / 5mi E Langtry / 15-VI-2017 / leg. D. Katz (USNM). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Paratypes</emphasis>
: 2 ♂♂, 2 ♀♀, same label data as holotype. 1♀, USA: Texas, 5mi W Langtry / 12-VII-2010 D. Katz (DEKC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas / Val Verde Co / Langtry / 18-VI-2013 / leg. D. Sundberg (DSC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / Langtry / 14-VI-2014 / leg. D. Sundberg (DSC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / Langtry / 28-XI-2014 / leg. D. Sundberg (DSC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / Langtry / 06-VII-2013 / leg. D. Sundberg (DSC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / Langtry / **-VII-2014 / leg. D. Sundberg (DSC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.2mi W. Langtry / 21-VI-2014 / leg. D. Duran (DPDC). 5 ♂♂, 3 ♀♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 4mi W Langtry / 18-VI-2015 / leg. S. Roman (SJRC). 1♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Terrell Co / 13mi W Langtry / 18-VI-2015 / leg. S. Roman (SJRC). 3 ♂♂, 4 ♀♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi W Langtry / 01-VI-2017 / leg. S. Roman (SJRC). 2 ♂♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi W Langtry / 01-VI-2017 / leg. S. Roman (SJRC). 2 ♂♂, 2 ♀♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi W Langtry / 01-VII-2019 / leg. S. Roman (SJRC). 1 ♂, 2 ♀♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi W Langtry / 01-VII-2019 / leg. D. Katz (DKC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi W Langtry / 01-VII-2019 / leg. D. Katz (DKC). 4♂, 5 ♀♀, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 3.5mi NW Langtry / 23-V-2016 / leg. D. Brzoska (DWBC). 1 ♂, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 12.5mi NW Langtry / 23-V-2016 / leg. Brzoska (DWBC). 1 ♂ USA: Texas / Brewster Co / Heath Canyon / 22/IX/2019 / leg. J. Chong/S. Roman (SJRC). 6 ♂♂, 5 ♀♀, 
<emphasis italics="true" pageId="0" pageNumber="125">ex ovum</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Terrell Co / 13mi W Langtry /
</paragraph>
<paragraph pageId="0" pageNumber="125">
18-VI-15 / leg. S. Roman (SJRC). 24 ♂♂, 24 ♀♀, 
<emphasis italics="true" pageId="0" pageNumber="125">ex ovum</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="125">USA</emphasis>
: Texas, / Val Verde Co / 13mi W Langtry / 18-VI-15 / leg. S. Roman (SJRC): 1 ♀ (USNM), 1 ♂, 1 ♀ (TAMUIC), 1 ♂, 1 ♀ (AMNH), 1 ♂, 1 ♀ (CMNH), 1 ♂, 1 ♀ (FSCA).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="diagnosis">
<paragraph pageId="0" pageNumber="125">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="125">
This species can be distinguished from all other 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila</emphasis>
</taxonomicName>
by the combination of trapezoidal pronotum, almond-shaped elytra with sharply defined dorsal carinae, smooth and polished elytra with sparse irregular rows of setigerous punctures, the lack of a spine-line projection on the male hind trochanter, and small size for the genus (total body length 23-28 mm). No other geographically proximate congeners can be confused with this species. 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila schwarzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila schwarzi</emphasis>
</taxonomicName>
is most similar in body shape and size, but its dorsal elytral texture is not smooth or polished, it possesses a greater number of shallow setigerous punctures that are arranged more regularly into rows between the dorsal carinae, and the two species are separated by 1300 km, with congeners occurring in the intervening areas.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="description">
<paragraph pageId="0" pageNumber="125">Description.</paragraph>
<paragraph pageId="0" pageNumber="125">
Small-sized 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila</emphasis>
</taxonomicName>
. Total body length 23.35-27.75 mm, mean ♀ 26.0 mm, mean ♂ 26.0 mm. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Head</emphasis>
(
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amblycheila katzi Duran &amp; Roman, sp. nov. frontal habitus (female)." figureDoi="10.3897/zookeys.892.47059.figure2" httpUri="https://binary.pensoft.net/fig/361371" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure2">Fig. 2</figureCitation>
): slightly narrower than anterior margin of pronotum, black throughout, with dark reddish reflections under strong light, moderately polished, 2-8 supraorbital setae next to each eye. Eyes round, proportionately small, not protruding beyond the genae when viewed from above. Frons clearly delimited from clypeus by a visible suture, gradually blending into vertex. Frons surface mostly smooth, lacking striae, but broadly wrinkled with shallow depressions. Vertex smooth, flat, glabrous. Genae smooth, with 0-6 setae present on each side. Clypeus slightly convex, with 2 setae. Labrum black, convex, smooth with 6-12 setae (typically 8-10) set in deep punctae, length 3.0-3.8 mm, width 1.1-1.8 mm, bidentate. Mandibles black, medium-sized, each with 2-12 setae along outside margin. Maxillary palpi dark reddish brown to black, color consistent in each segment. Labial palpi as above. Antennae long, reaching apical third of elytron, slightly longer in male than female; scape dark testaceous to black, with 2 or more subapical setae, sometimes with additional non-apical setae; pedicel dark reddish brown to black, with 2 or more subapical setae; flagellum dark reddish brown, with ring of long apical setae, covered with fine pubescence throughout.
</paragraph>
<caption doi="10.3897/zookeys.892.47059.figure1" httpUri="https://binary.pensoft.net/fig/361370" pageId="0" pageNumber="125" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 1.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila katzi</emphasis>
</taxonomicName>
Duran &amp; Roman, sp. nov. dorsal habitus (female).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.892.47059.figure2" httpUri="https://binary.pensoft.net/fig/361371" pageId="0" pageNumber="125" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 2.</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila katzi</emphasis>
</taxonomicName>
Duran &amp; Roman, sp. nov. frontal habitus (female).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Thorax</emphasis>
: Pronotum 5.60-6.85 mm in width (mean = 6.30), black, slightly polished, nearly smooth, with fine shallow rugosity, especially near margins; trapezoidal in shape with nearly straight sides, widest near apex, 5.80-7.60 mm (mean = 6.95); base narrow, 4.00-5.30 (mean = 4.70), setae sparse (1-7 per side) and present along lateral third of dorsal surface; disc with thin weakly impressed median line and deeply impressed transverse line; notopleural sutures clearly defined, not visible from dorsal view; proepisternum black, dull, glabrous or with few sparse long setae mostly concentrated near the coxae; meso- and metasternum glabrous or with a few setigerous punctures, especially near coxae. Elytra amygdaloid, 13.60-16.40 mm length (mean = 15.15), shape similar in both sexes; sutural spine absent, micro-serrations absent; elytral dorsal surface smooth and slightly polished, with sparse and irregularly placed setigerous punctae present between dorsal carinae; additional small shallow setigerous punctures are present in lateral areas, especially between dorsal carinae and epipleura.
</paragraph>
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Legs</emphasis>
: Pro-, meso- and metacoxae dark reddish brown to black, with sparse setae; trochanters dark reddish brown to black, pro- and mesotrochanter with a single long seta, sometimes with additional shorter setae; metatrochanter glabrous, apex blunt, lacking a produced apical tip; femora black, surface with multiple rows of erect brown setae; tibiae dark reddish brown to black, surface with erect brown setae similar to femora, with additional dense brown setae present at the apices near the tibial spines; tarsi dark reddish brown to black, bearing fine erect setae on lateral and ventral areas, tarsal claw long, simple.
</paragraph>
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">Abdomen</emphasis>
: Ventrites black, erect setae present and mostly restricted to a distinct row running parallel to the suture between ventrites, abdominal setae on ventrites 1-4 sparser in female that male; terminal abdominal ventrite with scattered setae near apical half in males, no setae present in females except for a dense fringe of setae emerging from the tip of the apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="etymology">
<paragraph pageId="0" pageNumber="125">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="125">
This new 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila</emphasis>
</taxonomicName>
is named after David Katz, the discoverer of this remarkable petrophilous insect. We propose the common name of Trans-Pecos Giant Tiger Beetle.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="distribution">
<paragraph pageId="0" pageNumber="125">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="125">
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila katzi</emphasis>
</taxonomicName>
is currently known only from western Texas in the Trans-Pecos region of the Chihuahuan Desert. All known occurrences are from steep-sided canyons in Val Verde and Terrell Counties, where natural or man-made forces have exposed bedrock, primarily Cretaceous limestone (
<bibRefCitation DOI="https://doi.org/10.3133/pp594K" author="Freeman, VL" journalOrPublisher="Unites States Government Printing Office, Washington DC" pageId="0" pageNumber="125" refId="B2" refString="Freeman, VL, 1968. Geology of the Comstock-Indian Wells area, Val Verde, Terrell, and Brewster Counties, Texas. Shorter contributions to general geology - Geological Survey Professional Paper 594-K. Unites States Government Printing Office, Washington DC" title="Geology of the Comstock-Indian Wells area, Val Verde, Terrell, and Brewster Counties, Texas. Shorter contributions to general geology - Geological Survey Professional Paper 594 - K." url="https://doi.org/10.3133/pp594K" year="1968">Freeman 1968</bibRefCitation>
) (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat for Amblycheila katzi sp. nov. A steep-sided limestone road cut B rock ledges with grooves and cracks C rock ledge with calcareous soil and third instar A. katzi larval burrow D closeup of third instar A. katzi larval burrow opening." figureDoi="10.3897/zookeys.892.47059.figure3" httpUri="https://binary.pensoft.net/fig/361372" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure3">Fig. 3A</figureCitation>
). This species appears to prefer vertical surfaces; the authors did not find any specimens on the ground, or in pitfall traps placed at the base of rock walls. All individuals observed by the authors were present 1-5m above the ground in rock crevices, grooves and ledges (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat for Amblycheila katzi sp. nov. A steep-sided limestone road cut B rock ledges with grooves and cracks C rock ledge with calcareous soil and third instar A. katzi larval burrow D closeup of third instar A. katzi larval burrow opening." figureDoi="10.3897/zookeys.892.47059.figure3" httpUri="https://binary.pensoft.net/fig/361372" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure3">Fig. 3B</figureCitation>
). Adult beetles were found in the same areas where 
<taxonomicName class="Arachnida" family="Theridiidae" genus="Latrodectus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Latrodectus" order="Araneae" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Latrodectus</emphasis>
</taxonomicName>
spiders were abundant. The second author discovered 2nd and 3rd instar larvae in fine calcareous clays within grooves and crevices in vertical limestone walls (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat for Amblycheila katzi sp. nov. A steep-sided limestone road cut B rock ledges with grooves and cracks C rock ledge with calcareous soil and third instar A. katzi larval burrow D closeup of third instar A. katzi larval burrow opening." figureDoi="10.3897/zookeys.892.47059.figure3" httpUri="https://binary.pensoft.net/fig/361372" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure3">Fig. 3C, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.892.47059.figure3" httpUri="https://binary.pensoft.net/fig/361372" pageId="0" pageNumber="125" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 3.</emphasis>
Habitat for 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila katzi</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
steep-sided limestone road cut 
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
rock ledges with grooves and cracks 
<emphasis bold="true" pageId="0" pageNumber="125">C</emphasis>
rock ledge with calcareous soil and third instar 
<taxonomicName lsidName="A. katzi" pageId="0" pageNumber="125" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">A. katzi</emphasis>
</taxonomicName>
larval burrow 
<emphasis bold="true" pageId="0" pageNumber="125">D</emphasis>
closeup of third instar 
<taxonomicName lsidName="A. katzi" pageId="0" pageNumber="125" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">A. katzi</emphasis>
</taxonomicName>
larval burrow opening.
</paragraph>
</caption>
<subSection pageId="0" pageNumber="125" type="key to the genus amblycheila">
<paragraph pageId="0" pageNumber="125">
Key to the genus 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="125">
<table pageId="0" pageNumber="125">
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
One or two distinct carinae (pleats) running the length of each elytron (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinae (pleats) present on Amblycheila elytra. A two distinct carinae (A. nyx) B three distinct carinae (A. halffteri)." figureDoi="10.3897/zookeys.892.47059.figure4" httpUri="https://binary.pensoft.net/fig/361373" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure4">Fig. 4A</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Three distinct carinae (pleats) running the length of each elytron (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Carinae (pleats) present on Amblycheila elytra. A two distinct carinae (A. nyx) B three distinct carinae (A. halffteri)." figureDoi="10.3897/zookeys.892.47059.figure4" httpUri="https://binary.pensoft.net/fig/361373" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure4">Fig. 4B</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Elytral texture dull; indistinct shallow punctae present. Elytra tapering towards apex. Length 19-26 mm. Southeastern Arizona, northern Mexico</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. baroni" pageId="0" pageNumber="125" rank="species" species="baroni">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. baroni</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Elytral texture polished; indistinct shallow punctae present along with many small pinpoint punctures. Elytral apex bulbous. Length 26-31 mm. Northern Mexico, possibly southwestern Texas</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. nyx" pageId="0" pageNumber="125" rank="species" species="nyx">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. nyx</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Elytra densely covered with pinpoint punctures. Without rows of larger punctae between dorsal carinae (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Elytral texture. A Dense pinpoint punctures present, without rows of larger punctae (A. cylindriformis) B with larger punctae loosely arranged into rows (A. hoversoni)." figureDoi="10.3897/zookeys.892.47059.figure5" httpUri="https://binary.pensoft.net/fig/361374" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure5">Fig. 5A</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Punctae between dorsal carinae loosely arranged into rows (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Elytral texture. A Dense pinpoint punctures present, without rows of larger punctae (A. cylindriformis) B with larger punctae loosely arranged into rows (A. hoversoni)." figureDoi="10.3897/zookeys.892.47059.figure5" httpUri="https://binary.pensoft.net/fig/361374" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure5">Fig. 5B</figureCitation>
), not densely covering surface. May also possess smaller pinpoint punctures throughout
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Elytra uniformly and densely punctate, surface texture dull to matte. Elytra dark red to dark brown. Length 29-40 mm. Great Plains region from South Dakota to Texas</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. cylindriformis" pageId="0" pageNumber="125" rank="species" species="cylindriformis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. cylindriformis</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Elytral surface polished, bearing small shallow punctures with smooth edges. Elytra black. Length 29-33 mm. Central Mexico</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. halffteri" pageId="0" pageNumber="125" rank="species" species="halffteri">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. halffteri</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Pronotum rounded (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronotal shape A rounded B trapezoidal." figureDoi="10.3897/zookeys.892.47059.figure6" httpUri="https://binary.pensoft.net/fig/361375" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure6">Fig. 6A</figureCitation>
). Elytra reddish brown to black. Hind trochanter in male with elongated apex, spine-line. Length 25-37 mm
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Pronotum trapezoidal, with nearly straight sides (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronotal shape A rounded B trapezoidal." figureDoi="10.3897/zookeys.892.47059.figure6" httpUri="https://binary.pensoft.net/fig/361375" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure6">Fig. 6B</figureCitation>
). Hind trochanter in male similar to pro- and mesotrochanter, blunt. Elytra black. Length 21-28 mm
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="125">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Dorsum of each elytron with one well-defined longitudinal row of punctae, with a smaller number of additional irregular punctae; many small shallow pinpoint punctures present. Length 25-31 mm. Arizona to west Texas</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. picolominii" pageId="0" pageNumber="125" rank="species" species="picolominii">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. picolominii</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">Dorsum of each elytron with deeply impressed punctae, arranged in multiple longitudinal rows or irregularly; shallow pinpoint punctures visible. Length 30-37 mm. South Texas</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. hoversoni" pageId="0" pageNumber="125" rank="species" species="hoversoni">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. hoversoni</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Elytral texture smooth and polished between sparse irregular setigerous punctae. Elytra nearly flat in lateral view, not bulging (
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lateral habitus. A Amblycheila katzi, sp. nov. male with elytra gradually tapering towards apex B A. schwarzi. male with elytral bulge and steep decline towards apex." figureDoi="10.3897/zookeys.892.47059.figure7" httpUri="https://binary.pensoft.net/fig/361376" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure7">Fig. 7A</figureCitation>
). Length 22-28 mm. Trans-Pecos region of West Texas
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. katzi" pageId="0" pageNumber="125" rank="species" species="katzi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. katzi</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="125">
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
Elytral texture dull, not shining; with one or more longitudinal rows of shallow setigerous punctae. Elytra with bulge near apex, especially in male (
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lateral habitus. A Amblycheila katzi, sp. nov. male with elytra gradually tapering towards apex B A. schwarzi. male with elytral bulge and steep decline towards apex." figureDoi="10.3897/zookeys.892.47059.figure7" httpUri="https://binary.pensoft.net/fig/361376" pageId="0" pageNumber="125" tableDoi="10.3897/zookeys.892.47059.figure7">Fig. 7B</figureCitation>
). Length 21-27 mm. Mojave Desert region of southeast California to southwest Utah and northwest Arizona
</td>
<td colspan="1" pageId="0" pageNumber="125" rowspan="1">
<taxonomicName lsidName="A. schwarzi" pageId="0" pageNumber="125" rank="species" species="schwarzi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="125">A. schwarzi</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zookeys.892.47059.figure4" httpUri="https://binary.pensoft.net/fig/361373" pageId="0" pageNumber="125" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 4.</emphasis>
Carinae (pleats) present on 
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila</emphasis>
</taxonomicName>
elytra. 
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
two distinct carinae (
<taxonomicName lsidName="A. nyx" pageId="0" pageNumber="125" rank="species" species="nyx">
<emphasis italics="true" pageId="0" pageNumber="125">A. nyx</emphasis>
</taxonomicName>
) 
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
three distinct carinae (
<taxonomicName lsidName="A. halffteri" pageId="0" pageNumber="125" rank="species" species="halffteri">
<emphasis italics="true" pageId="0" pageNumber="125">A. halffteri</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.892.47059.figure5" httpUri="https://binary.pensoft.net/fig/361374" pageId="0" pageNumber="125" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 5.</emphasis>
Elytral texture. 
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
Dense pinpoint punctures present, without rows of larger punctae (
<taxonomicName lsidName="A. cylindriformis" pageId="0" pageNumber="125" rank="species" species="cylindriformis">
<emphasis italics="true" pageId="0" pageNumber="125">A. cylindriformis</emphasis>
</taxonomicName>
) 
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
with larger punctae loosely arranged into rows (
<taxonomicName lsidName="A. hoversoni" pageId="0" pageNumber="125" rank="species" species="hoversoni">
<emphasis italics="true" pageId="0" pageNumber="125">A. hoversoni</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.892.47059.figure6" httpUri="https://binary.pensoft.net/fig/361375" pageId="0" pageNumber="125" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 6.</emphasis>
Pronotal shape 
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
rounded 
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
trapezoidal.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.892.47059.figure7" httpUri="https://binary.pensoft.net/fig/361376" pageId="0" pageNumber="125" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="125">
<emphasis bold="true" pageId="0" pageNumber="125">Figure 7.</emphasis>
Lateral habitus. 
<emphasis bold="true" pageId="0" pageNumber="125">A</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Amblycheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblycheila katzi" order="Coleoptera" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="katzi">
<emphasis italics="true" pageId="0" pageNumber="125">Amblycheila katzi</emphasis>
</taxonomicName>
, sp. nov. male with elytra gradually tapering towards apex 
<emphasis bold="true" pageId="0" pageNumber="125">B</emphasis>
<taxonomicName lsidName="A. schwarzi" pageId="0" pageNumber="125" rank="species" species="schwarzi">
<emphasis italics="true" pageId="0" pageNumber="125">A. schwarzi</emphasis>
</taxonomicName>
. male with elytral bulge and steep decline towards apex.
</paragraph>
</caption>
</subSection>
</subSubSection>
</treatment>
</document>