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<mods:title id="13F96DE259FEB2F53E8FA17D4D68AE82">Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="E1A95F786998F2FF7AC15B8B25690A6F">Kluge, Nikita J.</mods:namePart>
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<treatment id="4267878DFFAAFFD22687FA09FAA7FB0B" ID-DOI="http://doi.org/10.5281/zenodo.6042285" ID-GBIF-Taxon="134017031" ID-Zenodo-Dep="6042285" LSID="urn:lsid:plazi:treatment:4267878DFFAAFFD22687FA09FAA7FB0B" httpUri="http://treatment.plazi.org/id/4267878DFFAAFFD22687FA09FAA7FB0B" lastPageId="4" lastPageNumber="95" pageId="1" pageNumber="92">
<subSubSection id="82D46510FFAAFFD72687FA09FEA1F9C1" pageId="1" pageNumber="92" type="nomenclature">
<paragraph id="CA71369BFFAAFFD72687FA09FE7DFA20" blockId="1.[151,506,1492,1551]" box="[151,506,1492,1518]" pageId="1" pageNumber="92">
<heading id="913981F7FFAAFFD72687FA09FE7DFA20" bold="true" box="[151,506,1492,1518]" fontSize="11" level="1" pageId="1" pageNumber="92" reason="1">
<emphasis id="F8BAEA89FFAAFFD72687FA09FE7DFA20" bold="true" box="[151,506,1492,1518]" pageId="1" pageNumber="92">
Genus
<taxonomicName id="0DCE4D18FFAAFFD726FEFA09FE7DFA20" ID-CoL="3R5X" ID-ENA="328896" authority="Traver 1938" authorityName="Traver" authorityYear="1938" box="[238,506,1492,1518]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAAFFD726FEFA09FEDCFA20" bold="true" box="[238,347,1492,1518]" italics="true" pageId="1" pageNumber="92">Cloeodes</emphasis>
<bibRefCitation id="AE5F4B6AFFAAFFD72772FA09FE7DFA20" author="Traver" box="[354,506,1492,1518]" pageId="1" pageNumber="92" refString="Traver, J. R. (1938) Mayflies of Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 22, 5 - 42, pls. 1 - 3." type="journal article" year="1938">Traver 1938</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="CA71369BFFAAFFD72687FA2BFEA1F9C1" blockId="1.[151,506,1492,1551]" box="[151,294,1526,1551]" pageId="1" pageNumber="92">(Figs 1–146)</paragraph>
</subSubSection>
<subSubSection id="82D46510FFAAFFD72687F9E4FD26F8F0" pageId="1" pageNumber="92" type="reference_group">
<paragraph id="CA71369BFFAAFFD72687F9E4FDFDF9A0" blockId="1.[151,1071,1593,1646]" pageId="1" pageNumber="92">
<treatmentCitationGroup id="EADE11B5FFAAFFD72687F9E4FDFDF9A0" pageId="1" pageNumber="92">
Genus
<taxonomicName id="0DCE4D18FFAAFFD726CEF9E4FD43F981" ID-CoL="3R5X" authority=": Salles, Gattoliatt &amp; Sartori 2016: 100" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityPageNumber="100" authorityYear="2016" box="[222,708,1593,1615]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAAFFD726CEF9E4FEBBF981" box="[222,316,1593,1615]" italics="true" pageId="1" pageNumber="92">Cloeodes</emphasis>
: Salles, Gattoliatt &amp;
<treatmentCitation id="4B6F108AFFAAFFD72403F9E4FD43F981" author="Sartori" box="[531,708,1593,1615]" page="100" pageId="1" pageNumber="92" year="2016">
<bibRefCitation id="AE5F4B6AFFAAFFD72403F9E4FD13F981" author="Sartori" box="[531,660,1593,1615]" pageId="1" pageNumber="92" refString="Salles, F. F., Gattolliat, J. - L. &amp; Sartori, M. (2016) Phylogenetic analyses of Cloeodes Traver and related genera (Ephemeroptera: Baetidae). Systematic Entomology, 41 (1), 93 - 111. https: // doi. org / 10.1111 / syen. 12144" type="journal article" year="2016">Sartori 2016</bibRefCitation>
: 100
</treatmentCitation>
</taxonomicName>
(diagnosis for larvae and imagoes). Type species:
<taxonomicName id="0DCE4D18FFAAFFD72736F985FDF1F9A0" ID-CoL="5ZC57" authority="Traver 1938" authorityName="Traver" authorityYear="1938" box="[294,630,1624,1646]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="maculipes">
<emphasis id="F8BAEA89FFAAFFD72736F985FE76F9A0" box="[294,497,1624,1646]" italics="true" pageId="1" pageNumber="92">Cloeodes maculipes</emphasis>
<bibRefCitation id="AE5F4B6AFFAAFFD727E7F984FDF1F9A0" author="Traver" box="[503,630,1624,1646]" pageId="1" pageNumber="92" refString="Traver, J. R. (1938) Mayflies of Puerto Rico. Journal of Agriculture of the University of Puerto Rico, 22, 5 - 42, pls. 1 - 3." type="journal article" year="1938">Traver 1938</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="CA71369BFFAAFFD72687F948FD26F8F0" blockId="1.[151,1437,1685,2034]" pageId="1" pageNumber="92">
<emphasis id="F8BAEA89FFAAFFD72687F948FE0AF960" bold="true" box="[151,397,1685,1710]" pageId="1" pageNumber="92">Systematic position.</emphasis>
According to the phylogenetic classification of
<taxonomicName id="0DCE4D18FFAAFFD725F3F948FA1FF960" ID-CoL="6ZG" authority="(Kluge &amp; Novikova 2011)" authorityName="Kluge &amp; Novikova" authorityYear="2011" baseAuthorityName="Kluge &amp; Novikova" baseAuthorityYear="2011" box="[995,1432,1685,1710]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="family">
Baetidae (
<bibRefCitation id="AE5F4B6AFFAAFFD7224CF948FA17F960" author="Kluge" box="[1116,1424,1685,1710]" pageId="1" pageNumber="92" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFAAFFD72687F964FF78F91F" ID-CoL="3R5X" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[151,255,1721,1745]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAAFFD72687F964FF78F91F" box="[151,255,1721,1745]" italics="true" pageId="1" pageNumber="92">Cloeodes</emphasis>
</taxonomicName>
belongs to the following subsequently subordinated taxa: Tetramerotarsata
<bibRefCitation id="AE5F4B6AFFAAFFD72272F964FB6FF91C" author="Kluge" box="[1122,1256,1721,1746]" pageId="1" pageNumber="92" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics (Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne). Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="journal article" year="1997">Kluge 1997</bibRefCitation>
&gt; Liberevenata
<bibRefCitation id="AE5F4B6AFFAAFFD72687F900FE9BF938" author="Kluge" box="[151,284,1757,1782]" pageId="1" pageNumber="92" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics (Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne). Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="journal article" year="1997">Kluge 1997</bibRefCitation>
&gt; Turbanoculata
<bibRefCitation id="AE5F4B6AFFAAFFD727F5F900FDEDF938" author="Kluge" box="[485,618,1757,1782]" pageId="1" pageNumber="92" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics (Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne). Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="journal article" year="1997">Kluge 1997</bibRefCitation>
&gt; Anteropatellata
<bibRefCitation id="AE5F4B6AFFAAFFD7252FF900FC44F938" author="Kluge" box="[831,963,1757,1782]" pageId="1" pageNumber="92" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics (Proc. VIII Int. Conf. on Ephemeroptera and XII Int. Symposium on Plecoptera, August 1995, Lausanne). Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="journal article" year="1997">Kluge 1997</bibRefCitation>
&gt; Baetovectata
<bibRefCitation id="AE5F4B6AFFAAFFD7226DF900FA1FF938" author="Kluge" box="[1149,1432,1757,1782]" pageId="1" pageNumber="92" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
. Patella-tibial suture is typical for Anteropatellata [see discussion of apomorphy (1)]. Gonovectes have structure typical for Baetovectata (
<figureCitation id="52F52A1EFFAAFFD727A1F8F8FD8FF8F0" box="[433,520,1829,1854]" captionStart="FIGURES 32 – 36" captionStartId="11.[151,264,1813,1836]" captionTargetBox="[173,1416,207,1773]" captionTargetId="figure@11.[170,1417,193,1792]" captionTargetPageId="11" captionText="FIGURES 32 – 36. Cloeodes vibratorius sp. n. 32, fore wing of male; 33, subimaginal gonostyli developing under larval cuticle; 34, exuviae of subimaginal genitals; 35, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 36, gonostylus (32, 34 – 35, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888637/files/figure.png" pageId="1" pageNumber="92">Figs 36</figureCitation>
,
<figureCitation id="52F52A1EFFAAFFD72404F8FBFDDBF8F0" box="[532,604,1830,1854]" captionStart="FIGURES 86 – 91" captionStartId="22.[151,264,1689,1712]" captionTargetBox="[179,1404,198,1674]" captionTargetId="figure@22.[170,1417,185,1676]" captionTargetPageId="22" captionText="FIGURES 86 – 91. Cloeodes redactus, male genitals. 86 – 88, form « nigrostriatus »; 89 – 91, form « pallidus ». 86 and 89, subimaginal gonostyli developing under larval cuticle; 87 and 90, exuviae of subimaginal genitals; 88 and 91, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888651/files/figure.png" pageId="1" pageNumber="92">88, 91</figureCitation>
,
<figureCitation id="52F52A1EFFAAFFD72478F8FBFD12F8F0" box="[616,661,1830,1854]" captionStart="FIGURES 116 – 118" captionStartId="29.[151,264,1368,1391]" captionTargetBox="[191,1395,214,1346]" captionTargetId="figure@29.[170,1417,193,1348]" captionTargetPageId="29" captionText="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="1" pageNumber="92">118</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="82D46510FFAAFFD226D7F894FAA7FB0B" lastPageId="4" lastPageNumber="95" pageId="1" pageNumber="92" type="description">
<paragraph id="CA71369BFFAAFFD726D7F894FF72F83C" blockId="1.[151,1437,1685,2034]" pageId="1" pageNumber="92">
<emphasis id="F8BAEA89FFAAFFD726D7F894FDCDF8AC" bold="true" box="[199,586,1865,1890]" pageId="1" pageNumber="92">
Autapomorphies of
<taxonomicName id="0DCE4D18FFAAFFD727A4F894FD9BF8AC" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[436,540,1865,1890]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAAFFD727A4F894FD9BF8AC" bold="true" box="[436,540,1865,1890]" italics="true" pageId="1" pageNumber="92">Cloeodes</emphasis>
</taxonomicName>
. (1)
</emphasis>
On larval tibia, proximal arched row of fine setae is very regular, with setal sockets contiguous; this setal row is short and disconnected with patella-tibial suture (
<figureCitation id="52F52A1EFFAAFFD72265F8B0FB4CF848" box="[1141,1227,1901,1926]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="1" pageNumber="92">Fig. 16</figureCitation>
;
<bibRefCitation id="AE5F4B6AFFAAFFD722C9F8B0FAE4F848" author="Kluge" box="[1241,1379,1901,1926]" pageId="1" pageNumber="92" refString="Kluge, N. J. (1991) [Cuban mayflies of the family Baetidae (Ephemeroptera). 1. Genera Callibaetis, Cloeodes and Paracloeodes]. [Zoologicheskiy Zhurnal], 70 (12), 128 - 135. [in Russian with English summary]" type="journal article" year="1991">Kluge 1991</bibRefCitation>
: Fig. 3:6;
<bibRefCitation id="AE5F4B6AFFAAFFD726DAF84CFE08F864" author="Salles" box="[202,399,1937,1962]" pageId="1" pageNumber="92" refString="Salles, F. F., Massariol, F. C., Angeli, K. B, Lima, M. M., Gattolliat, J. - L. &amp; Sartori, M. S. (2015) Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges. Zootaxa, 4020 (1), 1 - 50. http: // dx. doi. org / 10.11646 / zootaxa. 4020.1.1" type="journal article" year="2015">
Salles
<emphasis id="F8BAEA89FFAAFFD72704F84FFEC2F867" box="[276,325,1937,1961]" italics="true" pageId="1" pageNumber="92">et al</emphasis>
. 2015
</bibRefCitation>
:
<figureCitation id="52F52A1EFFAAFFD7278AF84CFE67F864" box="[410,480,1937,1962]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1969,1992]" captionTargetBox="[191,1397,203,1957]" captionTargetId="figure@7.[191,1407,193,1957]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888629/files/figure.png" pageId="1" pageNumber="92">Figs 7</figureCitation>
d, 9d, 12a, 14a, 16a, 18a, 20a, 22a, 24a, 26a, 28a;
<bibRefCitation id="AE5F4B6AFFAAFFD72203F84CFB5FF864" author="Salles" box="[1043,1240,1937,1962]" pageId="1" pageNumber="92" refString="Salles, F. F., Gattolliat, J. - L. &amp; Sartori, M. (2016) Phylogenetic analyses of Cloeodes Traver and related genera (Ephemeroptera: Baetidae). Systematic Entomology, 41 (1), 93 - 111. https: // doi. org / 10.1111 / syen. 12144" type="journal article" year="2016">
Salles
<emphasis id="F8BAEA89FFAAFFD7224DF84FFB09F867" box="[1117,1166,1937,1961]" italics="true" pageId="1" pageNumber="92">et al</emphasis>
. 2016
</bibRefCitation>
: 6F). The species of doubtful position
<emphasis id="F8BAEA89FFAAFFD7276FF86BFE3CF803" box="[383,443,1974,1997]" italics="true" pageId="1" pageNumber="92">incus</emphasis>
<bibRefCitation id="AE5F4B6AFFAAFFD727D3F868FD70F800" author="Waltz" box="[451,759,1973,1998]" pageId="1" pageNumber="92" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 a) Generic revision of Cloeodes and description of two new genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184." type="journal article" year="1987" yearSuffix="a">Waltz &amp; McCafferty 1987a</bibRefCitation>
[
<taxonomicName id="0DCE4D18FFAAFFD72519F86BFCF0F803" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[777,887,1974,1997]" class="Insecta" family="Baetidae" genus="Bernerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAAFFD72519F86BFCF0F803" box="[777,887,1974,1997]" italics="true" pageId="1" pageNumber="92">Bernerius</emphasis>
</taxonomicName>
] has this character (
<bibRefCitation id="AE5F4B6AFFAAFFD7224DF868FF60F83C" author="Waltz" pageId="1" pageNumber="92" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 a) Generic revision of Cloeodes and description of two new genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184." type="journal article" year="1987" yearSuffix="a">
Waltz &amp; McCafferty 1987a:
<figureCitation id="52F52A1EFFAAFFD72687F804FF60F83C" box="[151,231,2009,2034]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1969,1992]" captionTargetBox="[191,1397,203,1957]" captionTargetId="figure@7.[191,1407,193,1957]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888629/files/figure.png" pageId="1" pageNumber="92">Fig. 11</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="CA71369BFFA9FFD426D7FF45FDC2FE8E" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="93">
In
<taxonomicName id="0DCE4D18FFA9FFD426FBFF45FED4FF7E" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[235,339,152,176]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD426FBFF45FED4FF7E" box="[235,339,152,176]" italics="true" pageId="2" pageNumber="93">Cloeodes</emphasis>
</taxonomicName>
patella-tibial suture retains usual shape, being not shifted distally (
<figureCitation id="52F52A1EFFA9FFD42290FF4AFB5FFF7E" box="[1152,1240,151,176]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="2" pageNumber="93">Fig. 16</figureCitation>
), and is equally developed on all legs on larva (
<figureCitation id="52F52A1EFFA9FFD427E5FF61FDF0FF1A" box="[501,631,188,213]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="2" pageNumber="93">Figs 21–23</figureCitation>
,
<figureCitation id="52F52A1EFFA9FFD42492FF61FD4CFF1B" box="[642,715,188,213]" captionStart="FIGURES 58 – 62" captionStartId="16.[151,264,1522,1545]" captionTargetBox="[151,1417,193,1501]" captionTargetId="figure@16.[151,1436,193,1501]" captionTargetPageId="16" captionText="FIGURES 58 – 62. Cloeodes redactus. 58 – 59, immature last instar female larva; 60 – 62, larval exuviae: 60, fore leg; 61, middle leg and half of metanotum; 62, hind leg." httpUri="https://zenodo.org/record/888641/files/figure.png" pageId="2" pageNumber="93">60–62</figureCitation>
,
<figureCitation id="52F52A1EFFA9FFD424C6FF61FC86FF1A" box="[726,769,188,212]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="2" pageNumber="93">102</figureCitation>
) and on all legs of female subimago and imago; it is absent only on fore legs of male subimago and imago. These features are initial for Anteropatellata and different from
<taxonomicName id="0DCE4D18FFA9FFD42687FED9FEA3FED2" authorityName="Braasch &amp; Soldan" authorityYear="1980" box="[151,292,260,284]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD42687FED9FEA3FED2" box="[151,292,260,284]" italics="true" pageId="2" pageNumber="93">Centroptella</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFA9FFD4274EFED9FE56FED2" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[350,465,260,284]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD4274EFED9FE56FED2" box="[350,465,260,284]" italics="true" pageId="2" pageNumber="93">Chopralla</emphasis>
</taxonomicName>
, whose patella-tibial suture on larval middle and hind legs is greatly shifted distally, and on fore legs is absent in all stages.
</paragraph>
<paragraph id="CA71369BFFA9FFD426D7FE91FE22FE3B" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="93">
Proximal arched row of fine setae is developed on larval tibia in some other taxa, being a plesiomorphy (
<bibRefCitation id="AE5F4B6AFFA9FFD42348FE91FE94FE46" author="Salles" pageId="2" pageNumber="93" refString="Salles, F. F., Gattolliat, J. - L. &amp; Sartori, M. (2016) Phylogenetic analyses of Cloeodes Traver and related genera (Ephemeroptera: Baetidae). Systematic Entomology, 41 (1), 93 - 111. https: // doi. org / 10.1111 / syen. 12144" type="journal article" year="2016">
Salles
<emphasis id="F8BAEA89FFA9FFD42687FEACFF40FE46" box="[151,199,368,392]" italics="true" pageId="2" pageNumber="93">et al</emphasis>
. 2016
</bibRefCitation>
); it can be poorly expressed (e.g., in
<taxonomicName id="0DCE4D18FFA9FFD424BFFEADFCAAFE46" box="[687,813,368,392]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD424BFFEADFCAAFE46" box="[687,813,368,392]" italics="true" pageId="2" pageNumber="93">Indocloeon</emphasis>
</taxonomicName>
—
<bibRefCitation id="AE5F4B6AFFA9FFD42559FEB2FC5AFE46" author="Kluge" box="[841,989,367,392]" pageId="2" pageNumber="93" refString="Kluge, N. J. (2012 a) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2012" yearSuffix="a">Kluge 2012a</bibRefCitation>
:
<figureCitation id="52F52A1EFFA9FFD425F7FEB2FBBEFE46" box="[999,1081,367,392]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="2" pageNumber="93">Fig. 15</figureCitation>
), or dense and regular, but with spaces between setae sockets and stretched distally up to beginning of the patella-tibial suture (e.g., in Cloeoninae—Kluge &amp;
<bibRefCitation id="AE5F4B6AFFA9FFD427B2FE6AFDD1FE1E" author="Novikova" box="[418,598,439,464]" pageId="2" pageNumber="93" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. [Entomologicheskoe Obozreni e = Revue d'Entomologie de l'URSS] 71 (1), 60 - 83. [in Russian with English summary, English translation: Entomological Review, 71 (9), 29 - 54]" type="journal article" year="1992">Novikova 1992</bibRefCitation>
: Figs 1–2). In
<taxonomicName id="0DCE4D18FFA9FFD42516FE65FC14FE1E" authorityName="Braasch &amp; Soldan" authorityYear="1980" box="[774,915,440,464]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD42516FE65FC14FE1E" box="[774,915,440,464]" italics="true" pageId="2" pageNumber="93">Centroptella</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFA9FFD425C0FE65FBC4FE1E" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[976,1091,440,464]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA9FFD425C0FE65FBC4FE1E" box="[976,1091,440,464]" italics="true" pageId="2" pageNumber="93">Chopralla</emphasis>
</taxonomicName>
it is also regular and spaced, being greatly elongated.
</paragraph>
<paragraph id="CA71369BFFA9FFD42687F8EFFEFFF848" pageId="2" pageNumber="93">
<table id="B8CEC43BFFA900292687F8EFFA1BF848" box="[151,1436,1842,1926]" gridcols="4" gridrows="3" pageId="2" pageNumber="93">
<tr id="74FE34D9FFA900292687F8EFFA1BF887" box="[151,1436,1842,1865]" gridrow="0" pageId="2" pageNumber="93">
<th id="372F5DA5FFA900292687F8EFFDA2F887" box="[151,549,1842,1865]" gridcol="0" gridrow="0" pageId="2" pageNumber="93">
<emphasis id="F8BAEA89FFA9FFD42687F8EFFEB8F889" bold="true" box="[151,319,1842,1865]" pageId="2" pageNumber="93">FIGURES 1–5.</emphasis>
<taxonomicName id="0DCE4D18FFA9FFD4275AF8EFFDA2F886" authorityName="Kluge" authorityYear="2017" box="[330,549,1842,1864]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="vibratorius">
<emphasis id="F8BAEA89FFA9FFD4275AF8EFFDA2F886" box="[330,549,1842,1864]" italics="true" pageId="2" pageNumber="93">Cloeodes vibratorius</emphasis>
</taxonomicName>
</th>
<th id="372F5DA5FFA90029243DF8EFFD2FF887" box="[557,680,1842,1865]" gridcol="1" gridrow="0" pageId="2" pageNumber="93">
<emphasis id="F8BAEA89FFA9FFD42420F8EEFDE9F886" bold="true" box="[560,622,1843,1864]" pageId="2" pageNumber="93">sp. n.</emphasis>
1–2,
</th>
<th id="372F5DA5FFA9002924A0F8EFFC03F887" box="[688,900,1842,1865]" gridcol="2" gridrow="0" pageId="2" pageNumber="93">immature last instar</th>
<th id="372F5DA5FFA90029259DF8EFFA1BF887" box="[909,1436,1842,1865]" gridcol="3" gridrow="0" pageId="2" pageNumber="93">male larva; 3, mature last instar female larva; 4,</th>
</tr>
<tr id="74FE34D9FFA900292687F88CFA1BF8A9" box="[151,1436,1873,1895]" gridrow="1" pageId="2" pageNumber="93">
<th id="372F5DA5FFA900292687F88CFDA2F8A9" box="[151,549,1873,1895]" gridcol="0" gridrow="1" pageId="2" pageNumber="93">penultimate instar female larva; 5, last</th>
<td id="372F5DA5FFA90029243DF88CFD2FF8A9" box="[557,680,1873,1895]" gridcol="1" gridrow="1" pageId="2" pageNumber="93">instar larval</td>
<td id="372F5DA5FFA9002924A0F88CFC03F8A9" box="[688,900,1873,1895]" gridcol="2" gridrow="1" pageId="2" pageNumber="93">exuviae of right half</td>
<td id="372F5DA5FFA90029259DF88CFA1BF8A9" box="[909,1436,1873,1895]" gridcol="3" gridrow="1" pageId="2" pageNumber="93">of pronotum and mesonotum. Abbreviations: II–X,</td>
</tr>
<tr id="74FE34D9FFA900292687F8ADFA1BF848" box="[151,1436,1904,1926]" gridrow="2" pageId="2" pageNumber="93" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="372F5DA5FFA900292687F8ADFDA2F848" box="[151,549,1904,1926]" gridcol="0" gridrow="2" pageId="2" pageNumber="93">abdominal terga II–X.</th>
</tr>
</table>
</paragraph>
<paragraph id="CA71369BFFA9FFD526D7F86FFCBFFE8E" blockId="2.[151,1436,1970,2031]" lastBlockId="3.[151,1437,151,1976]" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
<emphasis id="F8BAEA89FFA9FFD426D7F86FFF6FF805" bold="true" box="[199,232,1970,1995]" pageId="2" pageNumber="93">(2)</emphasis>
On larval abdominal sternum, transverse row of fine setae is very regular, with setal sockets contiguous (
<figureCitation id="52F52A1EFFA9FFD4268FF80BFF72F821" box="[159,245,2006,2031]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="2" pageNumber="93">Fig. 20</figureCitation>
:
<bibRefCitation id="AE5F4B6AFFA9FFD42712F80BFDB9F821" author="Waltz" box="[258,574,2006,2031]" pageId="2" pageNumber="93" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 a) Generic revision of Cloeodes and description of two new genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184." type="journal article" year="1987" yearSuffix="a">Waltz &amp; McCafferty 1987a</bibRefCitation>
: Fig. 5; 1987b:
<figureCitation id="52F52A1EFFA9FFD424E5F80BFCCEF821" box="[757,841,2006,2031]" captionStart="FIGURES 37 – 57" captionStartId="15.[151,264,1964,1987]" captionTargetBox="[185,1399,538,1939]" captionTargetId="figure@15.[170,1417,517,1951]" captionTargetPageId="15" captionText="FIGURES 37 – 57. Cloeodes spp., tergalii. 37 – 43, tergalii I – VII of Cloeodes vibratorius sp. n.; 44 – 57, tergalii I – VII of two individuals of Cloeodes redactus. For all three individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888639/files/figure.png" pageId="2" pageNumber="93">Fig. 44</figureCitation>
;
<bibRefCitation id="AE5F4B6AFFA9FFD42547F80BFBA6F821" author="Salles" box="[855,1057,2006,2031]" pageId="2" pageNumber="93" refString="Salles, F. F., Massariol, F. C., Angeli, K. B, Lima, M. M., Gattolliat, J. - L. &amp; Sartori, M. S. (2015) Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges. Zootaxa, 4020 (1), 1 - 50. http: // dx. doi. org / 10.11646 / zootaxa. 4020.1.1" type="journal article" year="2015">
Salles
<emphasis id="F8BAEA89FFA9FFD425B3F805FC52F821" box="[931,981,2007,2031]" italics="true" pageId="2" pageNumber="93">et al</emphasis>
. 2015
</bibRefCitation>
: 10E–F). Setae of these rows are long and simple (not furcate). Pair of these setal rows is present on each of abdominal sterna II–VI (
<bibRefCitation id="AE5F4B6AFFA8FFD52305FF4AFF53FF1B" author="Salles" pageId="3" pageNumber="94" refString="Salles, F. F., Gattolliat, J. - L. &amp; Sartori, M. (2016) Phylogenetic analyses of Cloeodes Traver and related genera (Ephemeroptera: Baetidae). Systematic Entomology, 41 (1), 93 - 111. https: // doi. org / 10.1111 / syen. 12144" type="journal article" year="2016">
Salles
<emphasis id="F8BAEA89FFA8FFD52372FF44FA12FF7E" box="[1378,1429,152,176]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. 2016
</bibRefCitation>
). On sternum II these rows can be either well-developed, or vestigial, consisting of few (3–2) setae, which sometimes are situated not close together.
<bibRefCitation id="AE5F4B6AFFA8FFD52497FF02FC51FF36" author="Waltz" box="[647,982,223,248]" pageId="3" pageNumber="94" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 a) Generic revision of Cloeodes and description of two new genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184." type="journal article" year="1987" yearSuffix="a">Waltz &amp; McCafferty (1987a)</bibRefCitation>
wrote about
<emphasis id="F8BAEA89FFA8FFD52264FF3DFB28FF39" box="[1140,1199,224,247]" italics="true" pageId="3" pageNumber="94">incus</emphasis>
[
<taxonomicName id="0DCE4D18FFA8FFD522D3FF3DFAB6FF39" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[1219,1329,224,247]" class="Insecta" family="Baetidae" genus="Bernerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD522D3FF3DFAB6FF39" box="[1219,1329,224,247]" italics="true" pageId="3" pageNumber="94">Bernerius</emphasis>
</taxonomicName>
], that its sterna have «long, fine setae randomly scattered over surface, without tufts of setae», that was a reason for placing it to a separate genus
<taxonomicName id="0DCE4D18FFA8FFD52799FEF5FCB3FE8E" authority="Waltz &amp; McCafferty 1987 a" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[393,820,295,320]" class="Insecta" family="Baetidae" genus="Bernerius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52799FEF5FE70FEF1" box="[393,503,296,319]" italics="true" pageId="3" pageNumber="94">Bernerius</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD527EEFEFAFCB3FE8E" author="Waltz" box="[510,820,295,320]" pageId="3" pageNumber="94" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 a) Generic revision of Cloeodes and description of two new genera (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 89 (1), 177 - 184." type="journal article" year="1987" yearSuffix="a">Waltz &amp; McCafferty 1987a</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FE90FB67FE3A" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
Transverse rows of long setae on the same places of abdominal sterna are present also in
<taxonomicName id="0DCE4D18FFA8FFD522CDFE91FAEDFEAA" authorityName="Braasch &amp; Soldan" authorityYear="1980" box="[1245,1386,332,356]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD522CDFE91FAEDFEAA" box="[1245,1386,332,356]" italics="true" pageId="3" pageNumber="94">Centroptella</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFA8FFD52687FEADFE8DFE46" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[151,266,368,392]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52687FEADFE8DFE46" box="[151,266,368,392]" italics="true" pageId="3" pageNumber="94">Chopralla</emphasis>
</taxonomicName>
, but their setal sockets are separated, and all setae are bifurcate. Presence of such spaced rows of bifurcate setae is not a reliable apomorphy, because the same rows are present in some non-related taxa, for example in selected Oriental species of
<taxonomicName id="0DCE4D18FFA8FFD5245FFE65FCFAFE1E" authority="Bengtsson 1915" authorityName="Bengtsson" authorityYear="1915" box="[591,893,439,464]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD5245FFE65FD44FE1E" box="[591,707,440,464]" italics="true" pageId="3" pageNumber="94">Procloeon</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD524D9FE65FCFAFE1E" author="Bengtsson" box="[713,893,439,464]" pageId="3" pageNumber="94" refString="Bengtsson, S. (1915) Eine Namensanderung. Entomologisk Tidskrift, 36 (1), 34." type="journal article" year="1915">Bengtsson 1915</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="AE5F4B6AFFA8FFD5259AFE6AFBCCFE1E" author="Salles" box="[906,1099,439,464]" pageId="3" pageNumber="94" refString="Salles, F. F., Gattolliat, J. - L. &amp; Sartori, M. (2016) Phylogenetic analyses of Cloeodes Traver and related genera (Ephemeroptera: Baetidae). Systematic Entomology, 41 (1), 93 - 111. https: // doi. org / 10.1111 / syen. 12144" type="journal article" year="2016">
Salles
<emphasis id="F8BAEA89FFA8FFD525C5FE64FB81FE1E" box="[981,1030,440,464]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. 2016
</bibRefCitation>
and own observation). Unlike this, rows of non-bifurcate setae with contiguous sockets is not found in taxa other than
<taxonomicName id="0DCE4D18FFA8FFD52261FE01FB5EFE3A" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[1137,1241,476,500]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52261FE01FB5EFE3A" box="[1137,1241,476,500]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FDDDFAA9FDF2" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">Setae forming transverse rows on sterna, are very delicate and colorless; under light microscope, their structure is well visible only if prepare a dry slide of empty cuticle (molting exuviae can be used for this purpose).</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FD9AFDAFFDAE" blockId="3.[151,1437,151,1976]" box="[199,552,583,608]" pageId="3" pageNumber="94">
<emphasis id="F8BAEA89FFA8FFD526D7FD9AFDAFFDAE" bold="true" box="[199,552,583,608]" pageId="3" pageNumber="94">
Other characters of
<taxonomicName id="0DCE4D18FFA8FFD527A9FD9AFDA6FDAE" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[441,545,583,608]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD527A9FD9AFDA6FDAE" bold="true" box="[441,545,583,608]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FDB0FDC1FD69" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
Eyes of female imago enlarged and projected above head surface (
<figureCitation id="52F52A1EFFA8FFD525D5FDB1FB98FD4A" box="[965,1055,620,645]" captionStart="FIGURES 26 – 31" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[151,1436,240,1855]" captionTargetId="figure@10.[151,1436,240,1855]" captionTargetPageId="10" captionText="FIGURES 26 – 31. Cloeodes vibratorius sp. n. 26 – 27, female imago; 28 – 31, male imago: 28, femur of fore leg; 29, femur of hind leg; 30, abdominal terga and sterna spread on slide; 31, lateral view. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888635/files/figure.png" pageId="3" pageNumber="94">Figs 27</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD5223DFDB1FBCBFD4A" box="[1069,1100,620,644]" captionStart="FIGURES 82 – 85" captionStartId="21.[151,264,2002,2025]" captionTargetBox="[151,1436,215,1951]" captionTargetId="figure@21.[151,1436,215,1951]" captionTargetPageId="21" captionText="FIGURES 82 – 85. Cloeodes redactus, female imago, form « nigrostriatus »." httpUri="https://zenodo.org/record/888649/files/figure.png" pageId="3" pageNumber="94">83</figureCitation>
;
<bibRefCitation id="AE5F4B6AFFA8FFD5224AFDB1FA1FFD4B" author="Waltz" box="[1114,1432,620,645]" pageId="3" pageNumber="94" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 b) Revision of the genus Cloeodes Traver (Ephemeroptera: Baetidae). Annals of the Entomological Society of America, 80 (2), 191 - 207. https: // doi. org / 10.1093 / aesa / 80.2.191" type="journal article" year="1987" yearSuffix="b">Waltz &amp; McCafferty 1987b</bibRefCitation>
:
<figureCitation id="52F52A1EFFA8FFD52687FD52FF6EFD66" box="[151,233,655,680]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="3" pageNumber="94">Fig. 17</figureCitation>
). The same in some other taxa.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FD69FDFFFCF6" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
Subimaginal tarsi bear pointed microlepides at least on apical (initial 5th) tarsomere; previous tarsomeres are either also covered by pointed microlepides, or covered mainly by blunt microlepides (as in
<bibRefCitation id="AE5F4B6AFFA8FFD522D1FD0AFF53FCDA" author="Kluge" pageId="3" pageNumber="94" refString="Kluge, N. J. &amp; Novikova, E. A. (2014) Systematics of Indobaetis Muller-Liebenau &amp; Morihara 1982, and related implications for some other Baetidae genera (Ephemeroptera). Zootaxa, 3835 (2), 209 - 236. https: // doi. org / 10.11646 / zootaxa. 3835.2.3" type="journal article" year="2014">Kluge &amp; Novikova 2014</bibRefCitation>
:
<figureCitation id="52F52A1EFFA8FFD526CEFD21FEDBFCDA" box="[222,348,764,789]" captionStart-0="FIGURES 6 – 12" captionStart-1="FIGURES 13 – 17" captionStartId-0="7.[151,264,1969,1992]" captionStartId-1="8.[151,264,1918,1941]" captionTargetBox-0="[191,1397,203,1957]" captionTargetBox-1="[191,1395,198,1904]" captionTargetId-0="figure@7.[191,1407,193,1957]" captionTargetId-1="figure@8.[191,1395,193,1908]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." captionText-1="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri-0="https://zenodo.org/record/888629/files/figure.png" httpUri-1="https://zenodo.org/record/888631/files/figure.png" pageId="3" pageNumber="94">Figs 11–17</figureCitation>
) (
<tableCitation id="874C0320FFA8FFD52762FD21FE41FCDA" box="[370,454,764,789]" captionStart="TABLE 1" captionStartId="12.[151,239,152,175]" captionText="TABLE 1. Texture of subimaginal legs and apical spines of imaginal and subimaginal legs." pageId="3" pageNumber="94">Table 1</tableCitation>
). In this respect
<taxonomicName id="0DCE4D18FFA8FFD52494FD21FD6BFCDA" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[644,748,764,788]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52494FD21FD6BFCDA" box="[644,748,764,788]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
differs from Baetofemorata and some other taxa, whose tarsi are entirely covered by blunt microlepides.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FC98FA05FC6B" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
In last larval instar, developing subimaginal gonostyli are folded under larval cuticle in «
<taxonomicName id="0DCE4D18FFA8FFD522C1FC99FAD3FC92" authorityName="Novikova &amp; Kluge" authorityYear="1987" box="[1233,1364,836,860]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD522C1FC99FAD3FC92" box="[1233,1364,836,860]" italics="true" pageId="3" pageNumber="94">Nigrobaetis</emphasis>
</taxonomicName>
-type» pose, with 2nd segment bent by convexity medially, and 3rd segment directed caudally (
<figureCitation id="52F52A1EFFA8FFD52264FCBAFB4CFC4E" box="[1140,1227,871,896]" captionStart="FIGURES 32 – 36" captionStartId="11.[151,264,1813,1836]" captionTargetBox="[173,1416,207,1773]" captionTargetId="figure@11.[170,1417,193,1792]" captionTargetPageId="11" captionText="FIGURES 32 – 36. Cloeodes vibratorius sp. n. 32, fore wing of male; 33, subimaginal gonostyli developing under larval cuticle; 34, exuviae of subimaginal genitals; 35, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 36, gonostylus (32, 34 – 35, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888637/files/figure.png" pageId="3" pageNumber="94">Figs 33</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD522C7FCBAFA98FC4E" box="[1239,1311,871,896]" captionStart="FIGURES 86 – 91" captionStartId="22.[151,264,1689,1712]" captionTargetBox="[179,1404,198,1674]" captionTargetId="figure@22.[170,1417,185,1676]" captionTargetPageId="22" captionText="FIGURES 86 – 91. Cloeodes redactus, male genitals. 86 – 88, form « nigrostriatus »; 89 – 91, form « pallidus ». 86 and 89, subimaginal gonostyli developing under larval cuticle; 87 and 90, exuviae of subimaginal genitals; 88 and 91, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888651/files/figure.png" pageId="3" pageNumber="94">86, 89</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD5233BFCBAFAD0FC4E" box="[1323,1367,871,896]" captionStart="FIGURES 116 – 118" captionStartId="29.[151,264,1368,1391]" captionTargetBox="[191,1395,214,1346]" captionTargetId="figure@29.[170,1417,193,1348]" captionTargetPageId="29" captionText="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="3" pageNumber="94">116</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD52372FCBAFA17FC4E" box="[1378,1424,871,896]" captionStart="FIGURES 136 – 138" captionStartId="33.[151,264,1795,1818]" captionTargetBox="[192,1400,350,1776]" captionTargetId="figure@33.[191,1401,337,1782]" captionTargetPageId="33" captionText="FIGURES 136 – 138. Cloeodes spp. 136, Cloeodes auwe, subimaginal sternum IX with styliger and gonostyli developing under larval cuticle (gonovectes shown by interrupted line; at left half gonovectal and gonostylar muscles shown by interrupted lines, hypodermal pigment shown by dots; median styligeral muscle shown by interrupted lines). 137 – 138, Cloeodes superior: 137, exuviae of subimaginal genitals; 138, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888667/files/figure.png" pageId="3" pageNumber="94">136</figureCitation>
). This pose is the most usual for the plesiomorphon Baetovectata-non-Baetofemorata, to which
<taxonomicName id="0DCE4D18FFA8FFD522A4FC51FA9BFC6A" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[1204,1308,908,932]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD522A4FC51FA9BFC6A" box="[1204,1308,908,932]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
belongs.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FC72FC81FA2A" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
<emphasis id="F8BAEA89FFA8FFD526D7FC72FDCDFC06" bold="true" box="[199,586,943,968]" pageId="3" pageNumber="94">
Variable characters of
<taxonomicName id="0DCE4D18FFA8FFD527CBFC72FDC4FC06" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[475,579,943,968]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD527CBFC72FDC4FC06" bold="true" box="[475,579,943,968]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
.
</emphasis>
MARGINAL INTERCALARIES OF FORE WING. At least in male, fore wing has double marginal intercalaries—i. e. two intercalaries per space between veins (
<figureCitation id="52F52A1EFFA8FFD52293FC09FB5CFC22" box="[1155,1243,980,1005]" captionStart="FIGURES 32 – 36" captionStartId="11.[151,264,1813,1836]" captionTargetBox="[173,1416,207,1773]" captionTargetId="figure@11.[170,1417,193,1792]" captionTargetPageId="11" captionText="FIGURES 32 – 36. Cloeodes vibratorius sp. n. 32, fore wing of male; 33, subimaginal gonostyli developing under larval cuticle; 34, exuviae of subimaginal genitals; 35, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 36, gonostylus (32, 34 – 35, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888637/files/figure.png" pageId="3" pageNumber="94">Figs 32</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD522F8FC09FA92FC22" box="[1256,1301,980,1004]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="3" pageNumber="94">110</figureCitation>
). In female, marginal intercalaries are either double as in male, or secondarily single. Among species examined, females of
<taxonomicName id="0DCE4D18FFA8FFD52687FBC1FE04FBFA" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[151,387,1052,1076]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="vibratorius" status="sp. nov.">
<emphasis id="F8BAEA89FFA8FFD52687FBC1FE04FBFA" box="[151,387,1052,1076]" italics="true" pageId="3" pageNumber="94">Cloeodes vibratorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFA8FFD5279AFBC1FE4EFBFB" box="[394,457,1052,1077]" pageId="3" pageNumber="94" rank="species">
<emphasis id="F8BAEA89FFA8FFD5279AFBC1FE4EFBFB" bold="true" box="[394,457,1052,1077]" pageId="3" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="0DCE4D18FFA8FFD527EFFBC1FCA4FBFA" authority="Salles et al. 2004" authorityName="Salles et al." authorityYear="2004" box="[511,803,1052,1077]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="auwe">
<emphasis id="F8BAEA89FFA8FFD527EFFBC1FDDCFBFA" box="[511,603,1052,1076]" italics="true" pageId="3" pageNumber="94">C. auwe</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD52472FBC1FCA4FBFA" author="Salles" box="[610,803,1052,1077]" pageId="3" pageNumber="94" refString="Salles, F. F., Batista, J. D. &amp; Cabette, H. R. S. (2004) Baetidae (Insecta: Ephemeroptera) de Nova Xavantina, Mato Grosso, Brasil: Novos registros e descricao de uma nova especie de Cloeodes Traver. Biota Neotropica, 4 (2), 1 - 8. https: // doi. org / 10.1590 / s 1676 - 06032004000200004" type="journal article" year="2004">
Salles
<emphasis id="F8BAEA89FFA8FFD524BCFBC0FD5AFBFA" box="[684,733,1052,1076]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. 2004
</bibRefCitation>
</taxonomicName>
have marginal intercalaries double, as in male (
<figureCitation id="52F52A1EFFA8FFD5232BFBC1FA17FBFB" box="[1339,1424,1052,1077]" captionStart="FIGURES 26 – 31" captionStartId="10.[151,264,1951,1974]" captionTargetBox="[151,1436,240,1855]" captionTargetId="figure@10.[151,1436,240,1855]" captionTargetPageId="10" captionText="FIGURES 26 – 31. Cloeodes vibratorius sp. n. 26 – 27, female imago; 28 – 31, male imago: 28, femur of fore leg; 29, femur of hind leg; 30, abdominal terga and sterna spread on slide; 31, lateral view. Abbreviations: I – X, abdominal segments I – X." httpUri="https://zenodo.org/record/888635/files/figure.png" pageId="3" pageNumber="94">Fig. 26</figureCitation>
). Females of
<taxonomicName id="0DCE4D18FFA8FFD5270CFB9DFCA7FB96" authority="Waltz &amp; McCafferty 1987" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[284,800,1087,1112]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="redactus">
<emphasis id="F8BAEA89FFA8FFD5270CFB9DFE6CFB96" box="[284,491,1088,1112]" italics="true" pageId="3" pageNumber="94">Cloeodes redactus</emphasis>
Waltz &amp; McCafferty 1987
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFA8FFD5253CFB9DFC63FB96" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[812,996,1088,1112]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
<emphasis id="F8BAEA89FFA8FFD5253CFB9DFC63FB96" box="[812,996,1088,1112]" italics="true" pageId="3" pageNumber="94">C. nigrohumeris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFA8FFD525FCFBE2FBAAFB96" box="[1004,1069,1087,1112]" pageId="3" pageNumber="94" rank="species">
<emphasis id="F8BAEA89FFA8FFD525FCFBE2FBAAFB96" bold="true" box="[1004,1069,1087,1112]" pageId="3" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="0DCE4D18FFA8FFD5222CFB9DFACEFB96" authority="Kluge 1991" authorityName="Kluge" authorityYear="1991" box="[1084,1353,1087,1112]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="superior">
<emphasis id="F8BAEA89FFA8FFD5222CFB9DFB3BFB99" box="[1084,1212,1088,1111]" italics="true" pageId="3" pageNumber="94">C. superior</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD522D5FBE2FACEFB96" author="Kluge" box="[1221,1353,1087,1112]" pageId="3" pageNumber="94" refString="Kluge, N. J. (1991) [Cuban mayflies of the family Baetidae (Ephemeroptera). 1. Genera Callibaetis, Cloeodes and Paracloeodes]. [Zoologicheskiy Zhurnal], 70 (12), 128 - 135. [in Russian with English summary]" type="journal article" year="1991">Kluge 1991</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFA8FFD52392FB9DFE04FBB2" authority="Kluge 1991" authorityName="Kluge" authorityYear="1991" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="inferior">
<emphasis id="F8BAEA89FFA8FFD52392FB9DFF6AFBB2" italics="true" pageId="3" pageNumber="94">C. inferior</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD526E9FBB9FE04FBB2" author="Kluge" box="[249,387,1124,1149]" pageId="3" pageNumber="94" refString="Kluge, N. J. (1991) [Cuban mayflies of the family Baetidae (Ephemeroptera). 1. Genera Callibaetis, Cloeodes and Paracloeodes]. [Zoologicheskiy Zhurnal], 70 (12), 128 - 135. [in Russian with English summary]" type="journal article" year="1991">Kluge 1991</bibRefCitation>
</taxonomicName>
have marginal intercalaries less developed than in male: between some veins only one intercalary is present, between others intercalaries are absent; sometimes marginal intercalaries are absent on the whole wing. In the cases, when there is one marginal intercalary in a space between veins, this intercalary locates not in the middle of the space, but either anteriad, or posteriad of the middle (
<figureCitation id="52F52A1EFFA8FFD5221AFB12FBE6FB26" box="[1034,1121,1231,1256]" captionStart="FIGURES 82 – 85" captionStartId="21.[151,264,2002,2025]" captionTargetBox="[151,1436,215,1951]" captionTargetId="figure@21.[151,1436,215,1951]" captionTargetPageId="21" captionText="FIGURES 82 – 85. Cloeodes redactus, female imago, form « nigrostriatus »." httpUri="https://zenodo.org/record/888649/files/figure.png" pageId="3" pageNumber="94">Figs 85</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD5227EFB0DFB1DFB26" box="[1134,1178,1232,1256]" captionStart="FIGURES 110 – 115" captionStartId="28.[151,264,1961,1984]" captionTargetBox="[165,1429,193,1948]" captionTargetId="figure@28.[165,1429,193,1948]" captionTargetPageId="28" captionText="FIGURES 110 – 115. Cloeodes nigrohumeris sp. n. 110, male imago and its enlarged wing margin; 111, male imago; 112, fore leg of male imago; 113, middle leg of male imago; 114, female imago and its enlarged wing margin; 115, female imago." httpUri="https://zenodo.org/record/888659/files/figure.png" pageId="3" pageNumber="94">114</figureCitation>
;
<bibRefCitation id="AE5F4B6AFFA8FFD522B7FB12FAB7FB26" author="Kluge" box="[1191,1328,1231,1256]" pageId="3" pageNumber="94" refString="Kluge, N. J. (1991) [Cuban mayflies of the family Baetidae (Ephemeroptera). 1. Genera Callibaetis, Cloeodes and Paracloeodes]. [Zoologicheskiy Zhurnal], 70 (12), 128 - 135. [in Russian with English summary]" type="journal article" year="1991">Kluge 1991</bibRefCitation>
:
<figureCitation id="52F52A1EFFA8FFD5232CFB12FA08FB26" box="[1340,1423,1231,1256]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="3" pageNumber="94">Fig. 13</figureCitation>
). Such eccentric single marginal intercalaries are secondary ones, i. e. originating from double intercalaries characteristic for Baetovectata. In contrast to them, the primary single marginal intercalaries, which are initially characteristic for Liberevenata, are located at the middle of each space. In this respect
<taxonomicName id="0DCE4D18FFA8FFD52268FAE1FB67FA9A" authorityName=": Salles, Gattoliatt &amp; Sartori" authorityYear="2016" box="[1144,1248,1340,1364]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52268FAE1FB67FA9A" box="[1144,1248,1340,1364]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
differs from the Afrotropical taxon
<taxonomicName id="0DCE4D18FFA8FFD52761FABDFD21FAB6" authority="Gillies 1990" authorityName="Gillies" authorityYear="1990" box="[369,678,1375,1400]" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52761FABDFD94FAB6" box="[369,531,1376,1400]" italics="true" pageId="3" pageNumber="94">Potamocloeon</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD5240BFA82FD21FAB6" author="Gillies" box="[539,678,1375,1400]" pageId="3" pageNumber="94" refString="Gillies, M. T. (1990) A new genus for the Afrotropical mayfly, Cloeon dentatum Kimmins (Ephem., Baetidae). Entomologist's Monthly Magazine, 126, 207 - 208." type="journal article" year="1990">Gillies 1990</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="0DCE4D18FFA8FFD524C1FABDFB24FAB6" authority="Lugo-Ortiz &amp; McCafferty 1997" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1997" box="[721,1187,1375,1400]" class="Insecta" family="Baetidae" genus="Maliqua" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD524C1FABDFCB7FAB6" box="[721,816,1376,1400]" italics="true" pageId="3" pageNumber="94">Maliqua</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD52528FA82FB24FAB6" author="Lugo-Ortiz" box="[824,1187,1375,1400]" pageId="3" pageNumber="94" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1997). Maliqua: a new genus of Baetidae (Ephemeroptera) for a species previously assigned to Afroptilum. Entomological News, 108 (5), 367 - 371." type="journal article" year="1997">Lugo-Ortiz &amp; McCafferty 1997</bibRefCitation>
</taxonomicName>
), which some authors united with
<taxonomicName id="0DCE4D18FFA8FFD5273BFA59FC21FA53" authority="(Jacobus &amp; McCafferty &amp; Gattolliat 2006)" authorityName="Jacobus &amp; McCafferty &amp; Gattolliat" authorityYear="2006" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[299,934,1412,1437]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD5273BFA59FE13FA52" box="[299,404,1412,1436]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
(
<bibRefCitation id="AE5F4B6AFFA8FFD527B9FA59FC19FA53" author="Jacobus" box="[425,926,1412,1437]" pageId="3" pageNumber="94" refString="Jacobus, L. M., McCafferty, W. P. &amp; Gattolliat, J. - L. (2006). Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology, 14 (1), 129 - 140." type="journal article" year="2006">Jacobus &amp; McCafferty &amp; Gattolliat 2006</bibRefCitation>
)
</taxonomicName>
: in contrast to
<taxonomicName id="0DCE4D18FFA8FFD52276FA59FB49FA52" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[1126,1230,1412,1436]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52276FA59FB49FA52" box="[1126,1230,1412,1436]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
, whose marginal intercalaries are either double or eccentric,
<taxonomicName id="0DCE4D18FFA8FFD5246DFA75FC98FA0E" authorityName="Gillies" authorityYear="1990" box="[637,799,1448,1472]" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD5246DFA75FC98FA0E" box="[637,799,1448,1472]" italics="true" pageId="3" pageNumber="94">Potamocloeon</emphasis>
</taxonomicName>
has one marginal intercalary in the middle of space, that is characteristic for baetid taxa other than Baetovectata.
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7FA2CFC1EF8CB" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
HIND WING. In various species of
<taxonomicName id="0DCE4D18FFA8FFD5244CFA2DFD43F9C6" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[604,708,1520,1544]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD5244CFA2DFD43F9C6" box="[604,708,1520,1544]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
, hind wings are either developed in both sexes, or absent in both sexes; in the last case larva has no vestiges of hind protoptera (
<figureCitation id="52F52A1EFFA8FFD52573F9C9FC45F9E2" box="[867,962,1556,1581]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="3" pageNumber="94">Figs. 22</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD525DEF9C9FC6BF9E3" box="[974,1004,1556,1581]" captionStart="FIGURES 58 – 62" captionStartId="16.[151,264,1522,1545]" captionTargetBox="[151,1417,193,1501]" captionTargetId="figure@16.[151,1436,193,1501]" captionTargetPageId="16" captionText="FIGURES 58 – 62. Cloeodes redactus. 58 – 59, immature last instar female larva; 60 – 62, larval exuviae: 60, fore leg; 61, middle leg and half of metanotum; 62, hind leg." httpUri="https://zenodo.org/record/888641/files/figure.png" pageId="3" pageNumber="94">61</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD525E9F9C9FBA0F9E2" box="[1017,1063,1556,1580]" captionStart="FIGURES 98 – 103" captionStartId="26.[151,264,1937,1960]" captionTargetBox="[174,1418,193,1924]" captionTargetId="figure@26.[174,1418,193,1925]" captionTargetPageId="26" captionText="FIGURES 98 – 103. Cloeodes nigrohumeris sp. n. 98 – 99, subimaginal exuviae: 98, half of mesonotum; 99, part of meso- and metapleuron with postsubalar sclerite and lateropostnotal crest; 100 – 103, larval exuviae: 100 – 101, abdomen and caudalii; 102, left thoracic pleura, legs and half of metanotum; 103, half of pronotum and mesonotum (99, holotype)." httpUri="https://zenodo.org/record/888655/files/figure.png" pageId="3" pageNumber="94">102</figureCitation>
). All species of
<taxonomicName id="0DCE4D18FFA8FFD522CFF9C9FAC0F9E2" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[1247,1351,1556,1580]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD522CFF9C9FAC0F9E2" box="[1247,1351,1556,1580]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
known from North and Central America, have no hind wings. Among South American species reliably attributed to
<taxonomicName id="0DCE4D18FFA8FFD52687F981FF78F9BA" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[151,255,1628,1652]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52687F981FF78F9BA" box="[151,255,1628,1652]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
(i.e., with known larval characters), hind wings are absent in
<taxonomicName id="0DCE4D18FFA8FFD525A7F981FAF1F9BB" authority="Waltz &amp; McCafferty 1987 b" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[951,1398,1628,1653]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="redactus">
<emphasis id="F8BAEA89FFA8FFD525A7F981FBB0F9BA" box="[951,1079,1628,1652]" italics="true" pageId="3" pageNumber="94">C. redactus</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD5222FF981FAF1F9BB" author="Waltz" box="[1087,1398,1628,1653]" pageId="3" pageNumber="94" refString="Waltz, R. D. &amp; McCafferty, W. P. (1987 b) Revision of the genus Cloeodes Traver (Ephemeroptera: Baetidae). Annals of the Entomological Society of America, 80 (2), 191 - 207. https: // doi. org / 10.1093 / aesa / 80.2.191" type="journal article" year="1987" yearSuffix="b">Waltz &amp; McCafferty 1987b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFA8FFD52392F981FD1FF956" authority="Salles &amp; Batista (in Salles et al.) 2004" authorityName="Salles &amp; Batista (in Salles et al." authorityYear="2004" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="auwe">
<emphasis id="F8BAEA89FFA8FFD52392F981FF55F956" italics="true" pageId="3" pageNumber="94">C. auwe</emphasis>
Salles &amp; Batista (in
<bibRefCitation id="AE5F4B6AFFA8FFD527D5F9A2FD1FF956" author="Salles" box="[453,664,1663,1688]" pageId="3" pageNumber="94" refString="Salles, F. F., Batista, J. D. &amp; Cabette, H. R. S. (2004) Baetidae (Insecta: Ephemeroptera) de Nova Xavantina, Mato Grosso, Brasil: Novos registros e descricao de uma nova especie de Cloeodes Traver. Biota Neotropica, 4 (2), 1 - 8. https: // doi. org / 10.1590 / s 1676 - 06032004000200004" type="journal article" year="2004">
Salles
<emphasis id="F8BAEA89FFA8FFD52401F95CFDC3F956" box="[529,580,1664,1688]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
.) 2004
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFA8FFD524B6F95DFBD6F956" authority="Nieto &amp; Richard 2008" authorityName="Nieto &amp; Richard" authorityYear="2008" box="[678,1105,1663,1688]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="barituensis">
<emphasis id="F8BAEA89FFA8FFD524B6F95DFCC2F956" box="[678,837,1664,1688]" italics="true" pageId="3" pageNumber="94">C. barituensis</emphasis>
<bibRefCitation id="AE5F4B6AFFA8FFD5255EF9A2FBD6F956" author="Nieto" box="[846,1105,1663,1688]" pageId="3" pageNumber="94" refString="Nieto, C. &amp; Richard, B. (2008) The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species. Zootaxa, 1727, 1 - 21." type="journal article" year="2008">Nieto &amp; Richard 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFA8FFD5224EF95DFB6BF956" authorityName="Massariol, Lima, Pinheiro, Quieroz, Concalves &amp; Salles" authorityYear="2013" box="[1118,1260,1664,1688]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="maracatu">
<emphasis id="F8BAEA89FFA8FFD5224EF95DFB6BF956" box="[1118,1260,1664,1688]" italics="true" pageId="3" pageNumber="94">C. maracatu</emphasis>
</taxonomicName>
Lima
<emphasis id="F8BAEA89FFA8FFD5232AF95CFAEAF956" box="[1338,1389,1664,1688]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. (in
<bibRefCitation id="AE5F4B6AFFA8FFD52687F979FE08F972" author="Massariol" box="[151,399,1700,1725]" pageId="3" pageNumber="94" refString="Massariol, F. C., Lima, L. R. C., Pinheiro, U. D. S., Quieroz, L. L., Oliveira, L. G. &amp; Salles, F. F. (2013) Two-winged Cloeodes in Brazil: New species, stage description, and key to South American species. Journal of Insect Science (Tucson), 13 (Article 60), 1 - 20." type="journal article" year="2013">
Massariol
<emphasis id="F8BAEA89FFA8FFD5271CF978FEBBF972" box="[268,316,1700,1724]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
.) 2013
</bibRefCitation>
,
<taxonomicName id="0DCE4D18FFA8FFD5278AF979FC57F972" authority="Queiroz et al. (in Massariol et al.) 2013" authorityName="Queiroz et al. (in Massariol et al." authorityYear="2013" box="[410,976,1700,1725]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="spaceki">
<emphasis id="F8BAEA89FFA8FFD5278AF979FD89F972" box="[410,526,1700,1724]" italics="true" pageId="3" pageNumber="94">C. spaceki</emphasis>
Queiroz
<emphasis id="F8BAEA89FFA8FFD52466F978FD21F972" box="[630,678,1700,1724]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. (in
<bibRefCitation id="AE5F4B6AFFA8FFD524CAF979FC57F972" author="Massariol" box="[730,976,1700,1725]" pageId="3" pageNumber="94" refString="Massariol, F. C., Lima, L. R. C., Pinheiro, U. D. S., Quieroz, L. L., Oliveira, L. G. &amp; Salles, F. F. (2013) Two-winged Cloeodes in Brazil: New species, stage description, and key to South American species. Journal of Insect Science (Tucson), 13 (Article 60), 1 - 20." type="journal article" year="2013">
Massariol
<emphasis id="F8BAEA89FFA8FFD52540F978FC06F972" box="[848,897,1700,1724]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
.) 2013
</bibRefCitation>
</taxonomicName>
and two new species described here—
<taxonomicName id="0DCE4D18FFA8FFD52392F979FE93F92E" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="vibratorius" status="sp. nov.">
<emphasis id="F8BAEA89FFA8FFD52392F979FE93F92E" italics="true" pageId="3" pageNumber="94">C. vibratorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFA8FFD5270CF91AFEDAF92E" box="[284,349,1735,1760]" pageId="3" pageNumber="94" rank="species">
<emphasis id="F8BAEA89FFA8FFD5270CF91AFEDAF92E" bold="true" box="[284,349,1735,1760]" pageId="3" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="0DCE4D18FFA8FFD52786F915FDC8F92E" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[406,591,1736,1760]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
<emphasis id="F8BAEA89FFA8FFD52786F915FDC8F92E" box="[406,591,1736,1760]" italics="true" pageId="3" pageNumber="94">C. nigrohumeris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFA8FFD52447F91AFD1FF92E" box="[599,664,1735,1760]" pageId="3" pageNumber="94" rank="species">
<emphasis id="F8BAEA89FFA8FFD52447F91AFD1FF92E" bold="true" box="[599,664,1735,1760]" pageId="3" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
(key for larvae of these two-winged species is given below). Other known South American species have hind wings (
<bibRefCitation id="AE5F4B6AFFA8FFD524D6F931FC0AF8CB" author="Salles" box="[710,909,1772,1797]" pageId="3" pageNumber="94" refString="Salles, F. F., Massariol, F. C., Angeli, K. B, Lima, M. M., Gattolliat, J. - L. &amp; Sartori, M. S. (2015) Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges. Zootaxa, 4020 (1), 1 - 50. http: // dx. doi. org / 10.11646 / zootaxa. 4020.1.1" type="journal article" year="2015">
Salles
<emphasis id="F8BAEA89FFA8FFD52501F930FCC4F8CA" box="[785,835,1772,1796]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
. 2015
</bibRefCitation>
).
</paragraph>
<paragraph id="CA71369BFFA8FFD526D7F8CDFF61F876" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="94">
MAXILLA. General shape of maxilla is uniform in all
<taxonomicName id="0DCE4D18FFA8FFD52544F8CDFC3BF8E6" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[852,956,1808,1832]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52544F8CDFC3BF8E6" box="[852,956,1808,1832]" italics="true" pageId="3" pageNumber="94">Cloeodes</emphasis>
</taxonomicName>
: it has short biting edge and thick distal dentiseta adjacent to canines (
<figureCitation id="52F52A1EFFA8FFD527F8F8E9FDAAF882" box="[488,557,1844,1869]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1969,1992]" captionTargetBox="[191,1397,203,1957]" captionTargetId="figure@7.[191,1407,193,1957]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888629/files/figure.png" pageId="3" pageNumber="94">Fig. 7</figureCitation>
); however, shape of the distal dentiseta varies among species: in most species its apex is bent toward canines (
<figureCitation id="52F52A1EFFA8FFD527EBF88AFDC5F8BE" box="[507,578,1879,1904]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1969,1992]" captionTargetBox="[191,1397,203,1957]" captionTargetId="figure@7.[191,1407,193,1957]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888629/files/figure.png" pageId="3" pageNumber="94">Figs 8</figureCitation>
,
<figureCitation id="52F52A1EFFA8FFD5245FF885FDFDF8BE" box="[591,634,1880,1904]" captionStart="FIGURES 139 – 144" captionStartId="34.[151,264,1660,1683]" captionTargetBox="[151,1436,447,1639]" captionTargetId="figure@34.[151,1436,447,1639]" captionTargetPageId="34" captionText="FIGURES 139 – 144. Cuban species of Cloeodes. 139 – 141 and 143 – 144, Cloeodes superior; 142, Cloeodes inferior. 139 – 140, left and right mandibles; 141, left mandible of another specimen; 142, apex of maxilla; 143 – 144, subimaginal exuviae: 143, half of mesonotum; 144, postsubalar sclerite and lateropostnotal crest. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888669/files/figure.png" pageId="3" pageNumber="94">142</figureCitation>
) (that can be called «
<taxonomicName id="0DCE4D18FFA8FFD52579F885FC28F8A1" authorityName="Leach" authorityYear="1815" box="[873,943,1880,1903]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52579F885FC28F8A1" box="[873,943,1880,1903]" italics="true" pageId="3" pageNumber="94">Baetis</emphasis>
</taxonomicName>
-type»), while in
<taxonomicName id="0DCE4D18FFA8FFD5227FF885FB4DF8BE" box="[1135,1226,1880,1904]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="auwe">
<emphasis id="F8BAEA89FFA8FFD5227FF885FB4DF8BE" box="[1135,1226,1880,1904]" italics="true" pageId="3" pageNumber="94">C. auwe</emphasis>
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFA8FFD522C8F885FACBF8BE" box="[1240,1356,1880,1904]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="spaceki">
<emphasis id="F8BAEA89FFA8FFD522C8F885FACBF8BE" box="[1240,1356,1880,1904]" italics="true" pageId="3" pageNumber="94">C. spaceki</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFA8FFD52392F885FEAAF85A" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
<emphasis id="F8BAEA89FFA8FFD52392F885FEAAF85A" italics="true" pageId="3" pageNumber="94">C. nigrohumeris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFA8FFD52724F8A1FEF3F85B" box="[308,372,1916,1941]" pageId="3" pageNumber="94" rank="species">
<emphasis id="F8BAEA89FFA8FFD52724F8A1FEF3F85B" bold="true" box="[308,372,1916,1941]" pageId="3" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
distal dentiseta is bent in the same direction as canines (
<figureCitation id="52F52A1EFFA8FFD525E0F8A1FBC8F85A" box="[1008,1103,1916,1941]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="3" pageNumber="94">Fig. 109</figureCitation>
) (that can be called «
<taxonomicName id="0DCE4D18FFA8FFD52352F8A1FA14F85A" authorityName="Leach" authorityYear="1815" box="[1346,1427,1916,1940]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFA8FFD52352F8A1FA14F85A" box="[1346,1427,1916,1940]" italics="true" pageId="3" pageNumber="94">Cloeon</emphasis>
</taxonomicName>
- type»).
</paragraph>
<paragraph id="CA71369BFFAFFFD226D7FF45FE43FE3B" blockId="4.[151,1436,151,1221]" pageId="4" pageNumber="95">
RESPIRATORY MOVEMENTS OF LARVA. In most
<taxonomicName id="0DCE4D18FFAFFFD22526FF4AFC1FFF7E" baseAuthorityName="Kluge &amp; Novikova" baseAuthorityYear="2011" box="[822,920,151,176]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
ability/inability for respiratory movements by tergalii is a character of high-level taxa; particularly, all representatives of the large world-wide taxon Baetungulata
<bibRefCitation id="AE5F4B6AFFAFFFD22687FF02FE29FF36" author="Kluge" box="[151,430,223,248]" pageId="4" pageNumber="95" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
are unable for respiratory vibrations by tergalii (that can be called «
<taxonomicName id="0DCE4D18FFAFFFD222A1FF3DFB70FF39" authorityName="Leach" authorityYear="1815" box="[1201,1271,224,247]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD222A1FF3DFB70FF39" box="[1201,1271,224,247]" italics="true" pageId="4" pageNumber="95">Baetis</emphasis>
</taxonomicName>
-type») and all representatives of the large Arctogean taxon Cloeonini (sensu
<bibRefCitation id="AE5F4B6AFFAFFFD22560FED9FC7DFED3" author="Kluge" box="[880,1018,260,285]" pageId="4" pageNumber="95" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
) are able for rhythmical respiratory movements by tergalii (that can be called «
<taxonomicName id="0DCE4D18FFAFFFD2248BFEF5FD6BFE8E" authorityName="Leach" authorityYear="1815" box="[667,748,296,320]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD2248BFEF5FD6BFE8E" box="[667,748,296,320]" italics="true" pageId="4" pageNumber="95">Cloeon</emphasis>
</taxonomicName>
-type»). Ability/inability for respiratory movements can be revealed if place alive larva to stagnant water with low oxygen concentration. In
<taxonomicName id="0DCE4D18FFAFFFD2220DFE91FB02FEAA" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[1053,1157,332,356]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD2220DFE91FB02FEAA" box="[1053,1157,332,356]" italics="true" pageId="4" pageNumber="95">Cloeodes</emphasis>
</taxonomicName>
this character is speciesspecific: among species examined,
<taxonomicName id="0DCE4D18FFAFFFD2243BFEADFD2BFE49" authorityName="Kluge" authorityYear="1991" box="[555,684,368,391]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="superior">
<emphasis id="F8BAEA89FFAFFFD2243BFEADFD2BFE49" box="[555,684,368,391]" italics="true" pageId="4" pageNumber="95">C. superior</emphasis>
</taxonomicName>
,
<taxonomicName id="0DCE4D18FFAFFFD224ACFEADFCB4FE46" authorityName="Kluge" authorityYear="1991" box="[700,819,368,392]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="inferior">
<emphasis id="F8BAEA89FFAFFFD224ACFEADFCB4FE46" box="[700,819,368,392]" italics="true" pageId="4" pageNumber="95">C. inferior</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFAFFFD2257EFEADFC76FE46" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[878,1009,368,392]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="redactus">
<emphasis id="F8BAEA89FFAFFFD2257EFEADFC76FE46" box="[878,1009,368,392]" italics="true" pageId="4" pageNumber="95">C. redactus</emphasis>
</taxonomicName>
do not make respiratory movements, while tergalii of
<taxonomicName id="0DCE4D18FFAFFFD2275EFE49FD83FE62" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[334,516,404,428]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="nigrohumeris" status="sp. nov.">
<emphasis id="F8BAEA89FFAFFFD2275EFE49FD83FE62" box="[334,516,404,428]" italics="true" pageId="4" pageNumber="95">C. nigrohumeris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFAFFFD2241AFE49FDCEFE63" box="[522,585,404,429]" pageId="4" pageNumber="95" rank="species">
<emphasis id="F8BAEA89FFAFFFD2241AFE49FDCEFE63" bold="true" box="[522,585,404,429]" pageId="4" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
are able to make fast synchronous vibrations with small amplitude. Larva of
<taxonomicName id="0DCE4D18FFAFFFD22687FE65FEB0FE1E" authority="Kluge, 2017" authorityName="Kluge" authorityYear="2017" box="[151,311,440,464]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="vibratorius" status="sp. nov.">
<emphasis id="F8BAEA89FFAFFFD22687FE65FEB0FE1E" box="[151,311,440,464]" italics="true" pageId="4" pageNumber="95">C. vibratorius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E38957F2FFAFFFD22751FE6AFE04FE1E" box="[321,387,439,464]" pageId="4" pageNumber="95" rank="species">
<emphasis id="F8BAEA89FFAFFFD22751FE6AFE04FE1E" bold="true" box="[321,387,439,464]" pageId="4" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
makes unusual respiratory movements by the whole body and usually does not vibrate by tergalii, but is able to do it.
</paragraph>
<paragraph id="CA71369BFFAFFFD226D7FDDDFBA9FD03" blockId="4.[151,1436,151,1221]" pageId="4" pageNumber="95">
STRUCTURE OF TERGALII. In the two-winged species of
<taxonomicName id="0DCE4D18FFAFFFD22568FDDDFC67FDD6" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[888,992,512,536]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD22568FDDDFC67FDD6" box="[888,992,512,536]" italics="true" pageId="4" pageNumber="95">Cloeodes</emphasis>
</taxonomicName>
examined by me, tergalii have speciesspecific shape, but uniform structure: ribs are present only on proximal parts of costal and anal margins; ribs are smooth, without denticles; distal margin of tergalius is free of rib and slightly notched, with small seta in each notch (
<figureCitation id="52F52A1EFFAFFFD226F7FDB1FEEBFD4B" box="[231,364,620,645]" captionStart="FIGURES 37 – 57" captionStartId="15.[151,264,1964,1987]" captionTargetBox="[185,1399,538,1939]" captionTargetId="figure@15.[170,1417,517,1951]" captionTargetPageId="15" captionText="FIGURES 37 – 57. Cloeodes spp., tergalii. 37 – 43, tergalii I – VII of Cloeodes vibratorius sp. n.; 44 – 57, tergalii I – VII of two individuals of Cloeodes redactus. For all three individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888639/files/figure.png" pageId="4" pageNumber="95">Figs 37–57</figureCitation>
,
<figureCitation id="52F52A1EFFAFFFD2276AFDB1FE67FD4A" box="[378,480,620,644]" captionStart="FIGURES 119 – 132" captionStartId="31.[151,264,1977,2000]" captionTargetBox="[200,1368,511,1956]" captionTargetId="figure@31.[197,1404,507,1970]" captionTargetPageId="31" captionText="FIGURES 119 – 132. Cloeodes spp., tergalii. 119 – 125, tergalii I – VII of Cloeodes nigrohumeris sp. n.; 126 – 132, tergalii I – VII of Cloeodes auwe. For both individuals vertical line corresponds to length of abdominal segment IV." httpUri="https://zenodo.org/record/888663/files/figure.png" pageId="4" pageNumber="95">119–132</figureCitation>
). Among fore-winged species, tergalii are more diverse: in some species tergalius, besides marginal ribs, has a median rib running close to median trachea (
<bibRefCitation id="AE5F4B6AFFAFFFD225C1FD52FB1DFD66" author="Salles" box="[977,1178,655,680]" pageId="4" pageNumber="95" refString="Salles, F. F., Massariol, F. C., Angeli, K. B, Lima, M. M., Gattolliat, J. - L. &amp; Sartori, M. S. (2015) Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges. Zootaxa, 4020 (1), 1 - 50. http: // dx. doi. org / 10.11646 / zootaxa. 4020.1.1" type="journal article" year="2015">
Salles
<emphasis id="F8BAEA89FFAFFFD2220CFD4CFBC9FD66" box="[1052,1102,656,680]" italics="true" pageId="4" pageNumber="95">et al</emphasis>
. 2015
</bibRefCitation>
). Similar median rib is present in all examined species of
<taxonomicName id="0DCE4D18FFAFFFD22408FD69FD22FD02" authorityName="Braasch &amp; Soldan" authorityYear="1980" box="[536,677,692,716]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD22408FD69FD22FD02" box="[536,677,692,716]" italics="true" pageId="4" pageNumber="95">Centroptella</emphasis>
</taxonomicName>
and
<taxonomicName id="0DCE4D18FFAFFFD224CBFD69FCC9FD02" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[731,846,692,716]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD224CBFD69FCC9FD02" box="[731,846,692,716]" italics="true" pageId="4" pageNumber="95">Chopralla</emphasis>
</taxonomicName>
(unpublished data).
</paragraph>
<paragraph id="CA71369BFFAFFFD226D7FD05FAA7FB0B" blockId="4.[151,1436,151,1221]" pageId="4" pageNumber="95">
APODEME OF LARVAL PARAPROCTS. Pair of paraprocts represent a ventral sclerite (sternite) of abdominal segment X, which is the last segment of the insect body (
<bibRefCitation id="AE5F4B6AFFAFFFD225D4FD21FBD4FCDA" author="Kluge" box="[964,1107,764,789]" pageId="4" pageNumber="95" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 456 pp. https: // doi. org / 10.1007 / 978 - 94 - 007 - 0872 - 3" type="book" year="2004">Kluge 2004</bibRefCitation>
). Paired ventral longitudinal muscles, being serially homologous in abdominal segments I–IX, stretch from anterior margin of each sternite to anterior margin of the next sternite; in abdominal segment IX these muscles stretch from anterior margin of sternite IX to anterior margins of paraprocts; pair of these muscles convergent posteriorly and are attached on a common unpaired base of paraprocts. This intersegmental ventral muscle is figured on longitudinal section in male imaginal genitals (
<bibRefCitation id="AE5F4B6AFFAFFFD226EFFC72FD8CFC06" author="Kluge" box="[255,523,943,968]" pageId="4" pageNumber="95" refString="Kluge, N. J. (2012 b) Systematics of Rhithrocloeoninae with new species from Uganda. Russian Entomological Journal, 21 (1), 1 - 13." type="journal article" year="2012" yearSuffix="b">
Kluge (2012b:
<figureCitation id="52F52A1EFFAFFFD227BEFC72FD85FC06" box="[430,514,943,968]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="4" pageNumber="95">Fig. 18</figureCitation>
)
</bibRefCitation>
; paired anterior insertions of these muscles on anterior margin of sternite IX of male imago are shown here on
<figureCitation id="52F52A1EFFAFFFD227E3FC09FDCFFC23" box="[499,584,980,1005]" captionStart="FIGURES 32 – 36" captionStartId="11.[151,264,1813,1836]" captionTargetBox="[173,1416,207,1773]" captionTargetId="figure@11.[170,1417,193,1792]" captionTargetPageId="11" captionText="FIGURES 32 – 36. Cloeodes vibratorius sp. n. 32, fore wing of male; 33, subimaginal gonostyli developing under larval cuticle; 34, exuviae of subimaginal genitals; 35, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines); 36, gonostylus (32, 34 – 35, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888637/files/figure.png" pageId="4" pageNumber="95">Figs 35</figureCitation>
,
<figureCitation id="52F52A1EFFAFFFD22444FC09FD1DFC22" box="[596,666,980,1004]" captionStart="FIGURES 86 – 91" captionStartId="22.[151,264,1689,1712]" captionTargetBox="[179,1404,198,1674]" captionTargetId="figure@22.[170,1417,185,1676]" captionTargetPageId="22" captionText="FIGURES 86 – 91. Cloeodes redactus, male genitals. 86 – 88, form « nigrostriatus »; 89 – 91, form « pallidus ». 86 and 89, subimaginal gonostyli developing under larval cuticle; 87 and 90, exuviae of subimaginal genitals; 88 and 91, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888651/files/figure.png" pageId="4" pageNumber="95">88, 91</figureCitation>
and
<figureCitation id="52F52A1EFFAFFFD224C0FC09FD7EFC22" box="[720,761,980,1004]" captionStart="FIGURES 116 – 118" captionStartId="29.[151,264,1368,1391]" captionTargetBox="[191,1395,214,1346]" captionTargetId="figure@29.[170,1417,193,1348]" captionTargetPageId="29" captionText="FIGURES 116 – 118. Cloeodes nigrohumeris sp. n. 116, subimaginal gonostyli extracted from mature larva; 117, exuviae of subimaginal genitals; 118, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines) (117 – 118, holotype). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger." httpUri="https://zenodo.org/record/888661/files/figure.png" pageId="4" pageNumber="95">118</figureCitation>
as “m.IX-X”. In larva, pair of these muscles is also attached to a common unpaired base of paraprocts. In most mayflies paraprocts have no significant apodeme at place of this muscle attachment, but in some species of
<taxonomicName id="0DCE4D18FFAFFFD2248BFBC1FC84FBFA" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[667,771,1052,1076]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD2248BFBC1FC84FBFA" box="[667,771,1052,1076]" italics="true" pageId="4" pageNumber="95">Cloeodes</emphasis>
</taxonomicName>
such apodeme is present. Among species examined,
<taxonomicName id="0DCE4D18FFAFFFD22392FBC1FE93FB96" authorityName="Kluge" authorityYear="2017" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="vibratorius">
<emphasis id="F8BAEA89FFAFFFD22392FBC1FE93FB96" italics="true" pageId="4" pageNumber="95">C. vibratorius</emphasis>
</taxonomicName>
has a long, thick, pigmented apodeme (
<figureCitation id="52F52A1EFFAFFFD224CFFBE2FCB0FB96" box="[735,823,1087,1112]" captionStart="FIGURES 13 – 17" captionStartId="8.[151,264,1918,1941]" captionTargetBox="[191,1395,198,1904]" captionTargetId="figure@8.[191,1395,193,1908]" captionTargetPageId="8" captionText="FIGURES 13 – 17. Cloeodes vibratorius sp. n. 13, labial palp; 14, labium (left half in ventral view with muscles shown by interrupted lines; right half in dorsal view); 15, paraprocts; 16, tibia of fore leg, anterior view (seta and suture on opposite side shown by interrupted lines; only two setae of proximal arched row are shown); 17, tarsus of hind leg, view from outer side (13, holotype)." httpUri="https://zenodo.org/record/888631/files/figure.png" pageId="4" pageNumber="95">Figs 15</figureCitation>
,
<figureCitation id="52F52A1EFFAFFFD22555FB9DFCE4FB96" box="[837,867,1088,1112]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="4" pageNumber="95">19</figureCitation>
),
<taxonomicName id="0DCE4D18FFAFFFD2256BFB9DFC7BFB96" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[891,1020,1088,1112]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="redactus">
<emphasis id="F8BAEA89FFAFFFD2256BFB9DFC7BFB96" box="[891,1020,1088,1112]" italics="true" pageId="4" pageNumber="95">C. redactus</emphasis>
</taxonomicName>
has thin and flexible apodeme (
<figureCitation id="52F52A1EFFAFFFD2237FFBE2FF31FBB3" captionStart="FIGURES 63 – 68" captionStartId="18.[151,264,1938,1961]" captionTargetBox="[170,1417,223,1879]" captionTargetId="figure@18.[170,1417,200,1900]" captionTargetPageId="18" captionText="FIGURES 63 – 68. Cloeodes redactus, larva. 63 – 64, left and right mandibles with developing mandibles of next instar inside (form « nigrostriatus »); 65 – 66, left and right mandibles (form « pallidus »); 67, paraprocts; 68, tarsus of hind leg, view from outer side." httpUri="https://zenodo.org/record/888643/files/figure.png" pageId="4" pageNumber="95">Fig. 67</figureCitation>
), and
<taxonomicName id="0DCE4D18FFAFFFD226E9FBB9FE28FBB2" authorityName="Kluge" authorityYear="2017" box="[249,431,1124,1148]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="nigrohumeris">
<emphasis id="F8BAEA89FFAFFFD226E9FBB9FE28FBB2" box="[249,431,1124,1148]" italics="true" pageId="4" pageNumber="95">C. nigrohumeris</emphasis>
</taxonomicName>
lacks significant apodeme (
<figureCitation id="52F52A1EFFAFFFD224F8FBB9FCCDFBB2" box="[744,842,1124,1149]" captionStart="FIGURES 104 – 109" captionStartId="27.[151,264,1666,1689]" captionTargetBox="[195,1394,195,1620]" captionTargetId="figure@27.[195,1400,183,1654]" captionTargetPageId="27" captionText="FIGURES 104 – 109. Cloeodes nigrohumeris sp. n., larva. 104 – 105, left and right mandibles; 106, labrum (apical setae not shown); 107, paraprocts; 108, labium (left half in ventral view; right half in dorsal view); 109, apex of maxilla (104 – 106, 108 – 109, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/888657/files/figure.png" pageId="4" pageNumber="95">Fig. 107</figureCitation>
). Possibly, strong development of this apodeme in
<taxonomicName id="0DCE4D18FFAFFFD22392FBB9FE93FB6E" authorityName="Kluge" authorityYear="2017" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="vibratorius">
<emphasis id="F8BAEA89FFAFFFD22392FBB9FE93FB6E" italics="true" pageId="4" pageNumber="95">C. vibratorius</emphasis>
</taxonomicName>
is connected with its peculiar abdominal respiratory movements. Similar apodemes are figured on paraprocts of some four-winged species of
<taxonomicName id="0DCE4D18FFAFFFD2246AFB71FD65FB0A" baseAuthorityName="Jacobus &amp; McCafferty &amp; Gattolliat" baseAuthorityYear="2006" box="[634,738,1196,1220]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="F8BAEA89FFAFFFD2246AFB71FD65FB0A" box="[634,738,1196,1220]" italics="true" pageId="4" pageNumber="95">Cloeodes</emphasis>
</taxonomicName>
(
<bibRefCitation id="AE5F4B6AFFAFFFD224E1FB71FC30FB0B" author="Salles" box="[753,951,1196,1221]" pageId="4" pageNumber="95" refString="Salles, F. F., Massariol, F. C., Angeli, K. B, Lima, M. M., Gattolliat, J. - L. &amp; Sartori, M. S. (2015) Revealing the diversity of Cloeodes Traver, 1938 (Ephemeroptera: Baetidae) in the Neotropics: description of eleven new species from Brazilian mountain ranges. Zootaxa, 4020 (1), 1 - 50. http: // dx. doi. org / 10.11646 / zootaxa. 4020.1.1" type="journal article" year="2015">
Salles
<emphasis id="F8BAEA89FFAFFFD2252CFB70FCEAFB0A" box="[828,877,1196,1220]" italics="true" pageId="4" pageNumber="95">et al</emphasis>
. 2015
</bibRefCitation>
:
<figureCitation id="52F52A1EFFAFFFD225D2FB71FB91FB0B" box="[962,1046,1196,1221]" captionStart="FIGURES 18 – 25" captionStartId="9.[151,264,1930,1953]" captionTargetBox="[151,1435,193,1876]" captionTargetId="figure@9.[151,1436,193,1885]" captionTargetPageId="9" captionText="FIGURES 18 – 25. Cloeodes vibratorius sp. n. 18 – 23, larval exuviae: 18, tergum IX; 19, paraprocts; 20, portion of sternum IV with setal row; 21, fore leg; 22, middle leg and half of metanotum; 23, hind leg; 24 – 25, subimaginal exuviae: 24, left half of mesonotum; 25, postsubalar sclerite and lateropostnotal crest (21 – 23, holotype)." httpUri="https://zenodo.org/record/888633/files/figure.png" pageId="4" pageNumber="95">Figs 22</figureCitation>
f, 24f and some others).
</paragraph>
</subSubSection>
</treatment>
</document>
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