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<document id="45DB7A240D8497D5DF9DDB42FC5E738E" ID-DOI="10.5281/zenodo.157876" ID-GBIF-Dataset="b599bfa1-2dd5-4af3-9f06-39716463af6d" ID-ISSN="1175­5326" ID-Zenodo-Dep="157876" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461012943975" checkinUser="plazi" docAuthor="Bird, Graham" docDate="2004" docId="422787BBFFE0E45A3F676D0EFCA7FD15" docLanguage="en" docName="zt00471.pdf" docOrigin="Zootaxa 471" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Thorkelius glacialis Hansen 1913, comb. nov." docType="treatment" docVersion="10" lastPageNumber="30" masterDocId="BE1EFFC3FFFAE4473E6F6C12FFF2FFB3" masterDocTitle="Tanaidacea (Crustacea) of the Northeast Atlantic: non­filiform species of Anarthruridae Lang from the Atlantic Margin" masterLastPageNumber="44" masterPageNumber="1" pageNumber="27" updateTime="1698567921015" updateUser="plazi">
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<mods:title id="A0D777B6A6D417F4CCCB665CAEF5C26F">Tanaidacea (Crustacea) of the Northeast Atlantic: non­filiform species of Anarthruridae Lang from the Atlantic Margin</mods:title>
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<subSubSection id="82946526FFE0E45D3F676D0EFC1FFE86" box="[264,1005,283,310]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="CA3136ADFFE0E45D3F676D0EFC1FFE86" blockId="26.[264,1005,283,310]" box="[264,1005,283,310]" pageId="26" pageNumber="27">
<heading id="917981C1FFE0E45D3F676D0EFC1FFE86" bold="true" box="[264,1005,283,310]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="F8FAEABFFFE0E45D3F676D0EFC1FFE86" bold="true" box="[264,1005,283,310]" pageId="26" pageNumber="27">
<taxonomicName id="0D8E4D2EFFE0E45D3F676D0EFD49FE85" ID-CoL="56KTK" authority="Hansen, 1913" authorityName="Hansen" authorityYear="1913" box="[264,699,283,310]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="glacialis" status="comb. nov.">
<emphasis id="F8FAEABFFFE0E45D3F676D0EFE0AFE85" bold="true" box="[264,504,284,310]" italics="true" pageId="26" pageNumber="27">Thorkelius glacialis</emphasis>
(
<bibRefCitation id="AE1F4B5CFFE0E45D3C676D09FD43FE86" author="Hansen" box="[520,689,283,309]" pageId="26" pageNumber="43" refString="Hansen, H. J. (1913) Crustacea Malacostraca, II, IV, The order Tanaidacea. Danish Ingolf Expedition, 3 (3), 1 - 145." type="journal article" year="1913">Hansen, 1913</bibRefCitation>
<emphasis id="F8FAEABFFFE0E45D3CDE6D0EFD49FE85" bold="true" box="[689,699,284,310]" italics="true" pageId="26" pageNumber="27">)</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3C957C4FFE0E45D3CAD6D09FCBAFE86" box="[706,840,283,309]" pageId="26" pageNumber="27">comb. nov.</taxonomicNameLabel>
<emphasis id="F8FAEABFFFE0E45D3D3F6D0EFCA8FE85" bold="true" box="[848,858,284,310]" italics="true" pageId="26" pageNumber="27">(</emphasis>
<figureCitation id="52B52A28FFE0E45D3D356D09FC17FE86" box="[858,997,283,309]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="26.[264,367,1739,1763]" captionStartId-1="27.[264,367,1783,1807]" captionTargetBox-0="[412,1180,659,1677]" captionTargetBox-1="[396,1175,719,1782]" captionTargetId-0="figure@26.[388,1198,644,1713]" captionTargetId-1="figure@27.[388,1198,699,1787]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." captionText-1="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri-0="https://zenodo.org/record/157887/files/figure.png" httpUri-1="https://zenodo.org/record/157888/files/figure.png" pageId="26" pageNumber="27">Figs. 1112</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="82946526FFE0E45D3F676D75FBF7FE32" box="[264,1029,359,385]" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="CA3136ADFFE0E45D3F676D75FBF7FE32" blockId="26.[264,1029,359,385]" box="[264,1029,359,385]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="EA9E1183FFE0E45D3F676D75FBF7FE32" box="[264,1029,359,385]" pageId="26" pageNumber="27">
<taxonomicName id="0D8E4D2EFFE0E45D3F676D75FCF2FE32" authority="Hansen, 1913: 102" authorityName="Hansen" authorityPageNumber="102" authorityYear="1913" box="[264,768,359,385]" class="Malacostraca" family="Leptognathiidae" genus="Leptognathia" kingdom="Animalia" order="Tanaidacea" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="F8FAEABFFFE0E45D3F676D75FDE2FE33" box="[264,528,359,384]" italics="true" pageId="26" pageNumber="27">Leptognathia glacialis</emphasis>
:
<treatmentCitation id="4B2F10BCFFE0E45D3C4F6D75FCF2FE32" author="Hansen" box="[544,768,359,385]" page="102" pageId="26" pageNumber="27" year="1913">
<bibRefCitation id="AE1F4B5CFFE0E45D3C4F6D75FD34FE32" author="Hansen" box="[544,710,359,385]" pageId="26" pageNumber="43" refString="Hansen, H. J. (1913) Crustacea Malacostraca, II, IV, The order Tanaidacea. Danish Ingolf Expedition, 3 (3), 1 - 145." type="journal article" year="1913">Hansen, 1913</bibRefCitation>
: 102
</treatmentCitation>
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, plate X figure 3a3b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="82946526FFE0E45D3F676DA4FDD2FD92" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="CA3136ADFFE0E45D3F676DA4FDD2FD92" blockId="26.[264,1323,438,585]" pageId="26" pageNumber="27">
<emphasis id="F8FAEABFFFE0E45D3F676DA4FE72FE63" bold="true" box="[264,384,438,464]" pageId="26" pageNumber="27">Diagnosis</emphasis>
. Fairly stout
<taxonomicName id="0D8E4D2EFFE0E45D3C4D6DA5FD6DFE63" box="[546,671,439,464]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="F8FAEABFFFE0E45D3C4D6DA5FD6DFE63" box="[546,671,439,464]" italics="true" pageId="26" pageNumber="27">Thorkelius</emphasis>
</taxonomicName>
with all pereonites distinctly shorter than broad. Pleotelson short and conical. Cheliped fixed finger conical, 2.5 times longer than broad, with long shallow convexity.
</paragraph>
</subSubSection>
<subSubSection id="82946526FFE0E45A3F576E3CFCC2FE35" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="CA3136ADFFE0E45D3F576E3CFB39FDFA" blockId="26.[264,1323,438,585]" box="[312,1227,558,585]" pageId="26" pageNumber="27">
<emphasis id="F8FAEABFFFE0E45D3F576E3CFDD7FDFB" bold="true" box="[312,549,558,584]" pageId="26" pageNumber="27">Material examined</emphasis>
. BIOICE Stn 2629, one neuter; Stn 2652, one prep. male.
</paragraph>
<caption id="9EF16625FFE0E45D3F676AD9FD42F8F9" httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="26" pageNumber="27" targetBox="[412,1180,659,1677]" targetPageId="26">
<paragraph id="CA3136ADFFE0E45D3F676AD9FD42F8F9" blockId="26.[264,1323,1739,1866]" pageId="26" pageNumber="27">
<emphasis id="F8FAEABFFFE0E45D3F676AD9FE69F950" bold="true" box="[264,411,1739,1763]" pageId="26" pageNumber="27">FIGURE 11.</emphasis>
<taxonomicName id="0D8E4D2EFFE0E45D3FC86AD9FD72F951" box="[423,640,1739,1762]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="F8FAEABFFFE0E45D3FC86AD9FD72F951" box="[423,640,1739,1762]" italics="true" pageId="26" pageNumber="27">Thorkelius glacialis</emphasis>
</taxonomicName>
. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; FG mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1mm (B), 0.67 mm (CD,K), 0.33 mm (EJ).
</paragraph>
</caption>
<paragraph id="CA3136ADFFE1E45C3F576D09FEB8FD95" blockId="27.[264,1324,283,670]" pageId="27" pageNumber="28">
<emphasis id="F8FAEABFFFE1E45C3F576D09FE14FE86" bold="true" box="[312,486,283,309]" pageId="27" pageNumber="28">Redescription</emphasis>
.
<emphasis id="F8FAEABFFFE1E45C3F996D0EFCA7FE86" box="[502,853,284,309]" italics="true" pageId="27" pageNumber="28">Neuter/non­ovigerous female.</emphasis>
Large, body (
<figureCitation id="52B52A28FFE1E45C3D956D0EFBBFFE85" box="[1018,1101,284,310]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
A) fairly stout, 5.1 times longer than broad. Length
<quantity id="0D769B48FFE1E45C3CE26D56FCEDFEED" box="[653,799,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="27" pageNumber="28" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.83.5 mm</quantity>
. Cephalothorax almost round and as long as broad, but tapering rostrally; rostrum with round apex. Pereonites 12 narrower caudally, all pereonites shorter than broad and laterally rounded; pereonites 14 with two dorsorostral setae. Pleon 21% of body length. Pleotelson (
<figureCitation id="52B52A28FFE1E45C3D226DAEFC52FE65" box="[845,928,444,470]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
B) as long as one and a half pleonites, conical but only 0.43 times as long as broad, with lateral, dorsocaudal and terminal setae.
</paragraph>
<paragraph id="CA3136ADFFE1E45C3F576E26FD4BFDC5" blockId="27.[264,1324,283,670]" pageId="27" pageNumber="28">
Antennule (see
<figureCitation id="52B52A28FFE1E45C3F806E26FDCDFDFD" box="[495,575,564,590]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
A) just longer than cephalothorax, similar to that of
<taxonomicName id="0D8E4D2EFFE1E45C3ACF6E26FAD1FDFE" box="[1184,1315,564,589]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="latiremis">
<emphasis id="F8FAEABFFFE1E45C3ACF6E26FAD1FDFE" box="[1184,1315,564,589]" italics="true" pageId="27" pageNumber="28">T. latiremis</emphasis>
</taxonomicName>
. Antenna similar to preparatory male.
</paragraph>
<paragraph id="CA3136ADFFE1E45C3F576E96FB39FD2D" blockId="27.[264,1324,283,670]" box="[312,1227,644,670]" pageId="27" pageNumber="28">Mouthparts as for preparatory male (as far as can be seen in whole specimen).</paragraph>
<caption id="9EF16625FFE1E45C3F676AE5FAFAF8DD" httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="27" pageNumber="28" targetBox="[396,1175,719,1782]" targetPageId="27">
<paragraph id="CA3136ADFFE1E45C3F676AE5FAFAF8DD" blockId="27.[264,1323,1783,1902]" pageId="27" pageNumber="28">
<emphasis id="F8FAEABFFFE1E45C3F676AE5FE68F8BC" bold="true" box="[264,410,1783,1807]" pageId="27" pageNumber="28">FIGURE 12.</emphasis>
<taxonomicName id="0D8E4D2EFFE1E45C3FCC6AEAFD8BF8BC" box="[419,633,1784,1807]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="F8FAEABFFFE1E45C3FCC6AEAFD8BF8BC" box="[419,633,1784,1807]" italics="true" pageId="27" pageNumber="28">Thorkelius glacialis</emphasis>
</taxonomicName>
. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; EG, pereopods 13 respectively; HK, pereopods 46 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm(EL), 0.5 mm (M), 0.33 mm (AD).
</paragraph>
</caption>
<paragraph id="CA3136ADFFE6E45B3F576D0EFD73FDC5" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Cheliped (
<figureCitation id="52B52A28FFE6E45B3FDD6D0EFDF7FE85" box="[434,517,284,310]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
K) surfaces with numerous microtrichia; pseudocoxa large, twice as long as basis; basis conical, with lateral seta; merus subtriangular, with strong ventral seta; carpus 1.4 times longer than broad, with two relatively long dorsal setae and two unequal setae near ventral protuberance; propodus without lateral fold, with group of three anterior spiniform setae (
<figureCitation id="52B52A28FFE6E45B3FA06DAEFDD4FE65" box="[463,550,444,470]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
D); fixed finger cylindrical, with two ventral setae, one seta near dactylus and a close group of three spiniform seta near incisive margin; incisive margin convex, proximal half with low crenulations, distal (
<figureCitation id="52B52A28FFE6E45B3DFF6E1EFC18FD95" box="[912,1002,524,550]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
C) with four subtriangular teeth; terminal spine conical; dactylus curved, with strong anterior spiniform seta and two small spines on incisive margin.
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F576E96FC51FD2D" blockId="28.[264,1324,284,1870]" box="[312,931,644,670]" pageId="28" pageNumber="29">Pereopods and pleopods similar to preparatory male.</paragraph>
<paragraph id="CA3136ADFFE6E45B3F576EBEFEB8FCA5" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Uropod (
<figureCitation id="52B52A28FFE6E45B3FCC6EBEFE0AFD75" box="[419,504,684,710]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
M) exopod shorter than basal article, with three unequal distal setae; endopod 1­articled, 5.5 times longer than broad, with one sensory seta and five terminal setae.
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F576F36FCEDFBCD" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
<emphasis id="F8FAEABFFFE6E45B3F576F36FDFCFC8E" box="[312,526,804,829]" italics="true" pageId="28" pageNumber="29">Preparatory male</emphasis>
. Body similar to neuter but pleon larger, 26% of body length. Length
<quantity id="0D769B48FFE6E45B3F0C6F5EFE33FCD5" box="[355,449,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="28" pageNumber="29" unit="mm" value="3.4">3.4 mm</quantity>
. Antennule (
<figureCitation id="52B52A28FFE6E45B3C366F5EFD5EFCD5" box="[601,684,844,870]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
C) articles broader than in neuter. Antenna (
<figureCitation id="52B52A28FFE6E45B3AD86F5EFAF8FCD5" box="[1207,1290,844,870]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
D) similar to that of
<taxonomicName id="0D8E4D2EFFE6E45B3FBC6F66FDA5FC3E" box="[467,599,884,909]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="latiremis">
<emphasis id="F8FAEABFFFE6E45B3FBC6F66FDA5FC3E" box="[467,599,884,909]" italics="true" pageId="28" pageNumber="29">T. latiremis</emphasis>
</taxonomicName>
. Labrum (
<figureCitation id="52B52A28FFE6E45B3CBC6F66FCD7FC3D" box="[723,805,884,910]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
E) broadly conical. Mandible (
<figureCitation id="52B52A28FFE6E45B3AE16F66FB18FC3D" box="[1166,1258,884,910]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Figs. 11</figureCitation>
FG) with long conical incisor. Maxilla (
<figureCitation id="52B52A28FFE6E45B3CD96F8EFCFEFC05" box="[694,780,924,950]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
H) pear shaped. Maxillule endite (
<figureCitation id="52B52A28FFE6E45B3ADD6F8EFAFAFC05" box="[1202,1288,924,950]" captionStart="FIGURE 11" captionStartId="26.[264,367,1739,1763]" captionTargetBox="[412,1180,659,1677]" captionTargetId="figure@26.[388,1198,644,1713]" captionTargetPageId="26" captionText="FIGURE 11. Thorkelius glacialis. Non­ovigerous female A, dorsal; B, pleotelson; preparatory male C, antennule, lateral view, setae omitted; D, antenna; E, labrum, ventral view; F G mandibles; H, maxillule and maxilla; J, epignath; non­ovigerous female K, cheliped. Scale bar 2 mm (A), 1 mm (B), 0.67 mm (C D, K), 0.33 mm (E J)." httpUri="https://zenodo.org/record/157887/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
H) with one short and six long terminal spiniform setae. Maxilliped basis with seta near palp; palp (
<figureCitation id="52B52A28FFE6E45B3F236FFEFE51FBB5" box="[332,419,1004,1030]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
A) article­2 with three setae; article­3 with four medial setae and one other seta; article­4 with five distal setae, all setae smooth or only minutely pinnate; endite (
<figureCitation id="52B52A28FFE6E45B3A976806FED4FBE5" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
B) rectangular, with strong incurved medial seta. Cheliped as neuter, distal teeth of fixed finger better defined (possibly less abraded?).
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F57689EFC6AFAF5" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Pereopod­1 (
<figureCitation id="52B52A28FFE6E45B3FBF689EFDD4FB15" box="[464,550,1164,1190]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
E) coxa with seta; basis about five times longer than broad, with proximal seta; ischium with one seta; merus 2.6 times longer than broad, with distoventral seta; carpus subrectangular, three times longer than broad, with three distal setae; propodus 1.4 times longer than carpus, with distoventral seta; dactylus and unguis together about as long as propodus, dactylus with accessory seta.
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F576946FE1AFA55" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Pereopod­2 (
<figureCitation id="52B52A28FFE6E45B3FBF6946FDD7FADD" box="[464,549,1364,1390]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
F) similar to pereopod­1 but basis and merus slightly shorter and stouter; carpus twice as long as broad; propodus about 0.8 times length of pereopod­1 article. Pereopod­3 (
<figureCitation id="52B52A28FFE6E45B3FBB69B6FDD8FA0D" box="[468,554,1444,1470]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
G) similar to pereopod­2 but basis and carpus shorter; propodus longer than carpus.
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F5769E6FEB8F965" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Pereopod­4 (
<figureCitation id="52B52A28FFE6E45B3FBB69E6FDDFF9BD" box="[468,557,1524,1550]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
H) basis stout, 2.5 times longer than broad, with two sensory setae; ischium with two setae; merus curved, twice as long as broad, with two spiniform setae bearing bifid tips; carpus subrectangular, twice as long as broad, with three spiniform setae bearing complex tips and a simple seta; propodus as long as carpus, with three long, spiniform, and complex setae; dactylus and unguis 0.7 times length of propodus, unguis short.
</paragraph>
<paragraph id="CA3136ADFFE6E45B3F576AF6FEB8F8FD" blockId="28.[264,1324,284,1870]" pageId="28" pageNumber="29">
Pereopod­5 (
<figureCitation id="52B52A28FFE6E45B3FA16AF6FDD0F94D" box="[462,546,1764,1790]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
J) similar to pereopod­4, ischium with two setae. Pereopod­6 (
<figureCitation id="52B52A28FFE6E45B3A956AF6FED7F895" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="28" pageNumber="29">Fig. 12</figureCitation>
K) similar to pereopod­5, but basis shorter and propodus with four distal spiniform setae.
</paragraph>
<paragraph id="CA3136ADFFE7E45A3F576D0EFD3EFEED" blockId="29.[264,1323,284,678]" pageId="29" pageNumber="30">
Pleopod (
<figureCitation id="52B52A28FFE7E45A3FC76D0EFE09FE85" box="[424,507,284,310]" captionStart="FIGURE 12" captionStartId="27.[264,367,1783,1807]" captionTargetBox="[396,1175,719,1782]" captionTargetId="figure@27.[388,1198,699,1787]" captionTargetPageId="27" captionText="FIGURE 12. Thorkelius glacialis. Preparatory male A, maxilliped palp; B, maxilliped endite; C, cheliped fixed finger, detail; D, anterior setae of cheliped propodus; E G, pereopods 1 3 respectively; H K, pereopods 4 6 respectively, with enlargements of complex spiniform setae; L, pleopod; non­ovigerous female M, uropod. Scale bar 0.67 mm (E L), 0.5 mm (M), 0.33 mm (A D)." httpUri="https://zenodo.org/record/157888/files/figure.png" pageId="29" pageNumber="30">Fig. 12</figureCitation>
L) rami well developed, endopod and exopod with about ten and thirteen finely plumose setae respectively.
</paragraph>
<paragraph id="CA3136ADFFE7E45A3F576D7EFCC2FE35" blockId="29.[264,1323,284,678]" box="[312,816,364,390]" pageId="29" pageNumber="30">Uropod as in neuter/non­ovigerous female.</paragraph>
</subSubSection>
<subSubSection id="82946526FFE7E45A3F576D81FCA7FD15" pageId="29" pageNumber="30" type="discussion">
<paragraph id="CA3136ADFFE7E45A3F576D81FCF5FD95" blockId="29.[264,1323,284,678]" pageId="29" pageNumber="30">
<emphasis id="F8FAEABFFFE7E45A3F576D81FE58FE1E" bold="true" box="[312,426,403,429]" pageId="29" pageNumber="30">Remarks</emphasis>
. This species, not extensively figured by Hansen, can easily be accommodated in the same genus as
<taxonomicName id="0D8E4D2EFFE7E45A3C226DAEFD3DFE66" box="[589,719,444,469]" class="Malacostraca" family="Anarthruridae" genus="Thorkelius" kingdom="Animalia" order="Tanaidacea" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="latiremis">
<emphasis id="F8FAEABFFFE7E45A3C226DAEFD3DFE66" box="[589,719,444,469]" italics="true" pageId="29" pageNumber="30">T. latiremis</emphasis>
</taxonomicName>
, sharing very similar maxilliped, mandible, pereopod and uropod morphology. It is best distinguished by its shorter pereonites and more elongate cheliped dactylus and fixed finger.
</paragraph>
<paragraph id="CA3136ADFFE7E45A3F576E26FCA7FD15" blockId="29.[264,1323,284,678]" pageId="29" pageNumber="30">
Apart from the
<typeStatus id="1535880FFFE7E45A3F826E26FDEDFDFD" box="[493,543,564,590]" pageId="29" pageNumber="30">type</typeStatus>
locality in Fleming Inlet, East
<collectingCountry id="B299763DFFE7E45A3DE46E26FBFAFDFD" box="[907,1032,564,590]" name="Greenland" pageId="29" pageNumber="30">Greenland</collectingCountry>
,
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(
<bibRefCitation id="AE1F4B5CFFE7E45A3A026E26FAF8FDFD" author="Hansen" box="[1133,1290,564,590]" pageId="29" pageNumber="43" refString="Hansen, H. J. (1913) Crustacea Malacostraca, II, IV, The order Tanaidacea. Danish Ingolf Expedition, 3 (3), 1 - 145." type="journal article" year="1913">Hansen 1913</bibRefCitation>
) it has been recorded from two BIOICE stations on the
<collectingCountry id="B299763DFFE7E45A3DFF6E4EFC14FDC5" box="[912,998,604,630]" name="Iceland" pageId="29" pageNumber="30">Iceland</collectingCountry>
Plateau,
<quantity id="0D769B48FFE7E45A3A356E4EFB16FDC5" box="[1114,1252,604,630]" metricMagnitude="2" metricUnit="m" metricValue="7.58" metricValueMax="9.129999999999999" metricValueMin="6.029999999999999" pageId="29" pageNumber="30" unit="m" value="758.0" valueMax="913.0" valueMin="603.0">603913 m</quantity>
, with mean bottom temperature 1o C, and silt substrate.
</paragraph>
</subSubSection>
</treatment>
</document>