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<document id="40C4E7779E01F36FA3DC57576BACF429" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FFE29879FF63DA7977D2FDD8" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) humerorotunda Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="52" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="50" updateTime="1700188109802" updateUser="ExternalLinkService">
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<mods:title id="1DCE2F0F4057F05DE598C045A2D878CA">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
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<mods:namePart id="3BB85041E189E3DEC36F68B4D608AFA5">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="D710CD1EFE61AADB5608266CCB1535DB">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
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<mods:namePart id="CC19E981018C031A1CB4D785BA7DC325">Silva, Valéria Juliete Da</mods:namePart>
<mods:nameIdentifier id="58DDABE74C8121279FFA23902A30AEE1" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
<mods:affiliation id="0A7E9B8C8ED2B3CE08B8298E43BEB469">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
<mods:nameIdentifier id="565325BDD0D7E5E7E5F86CA05A627129" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="D1BE088D0925E0072FA4BFEE71345406">Fernandes, José Antônio Marin</mods:namePart>
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<mods:affiliation id="D773C9B7BA016AE813D7FF75FC82052E">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
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<mods:title id="C61C20B27AF56294663C0A89821667A9">Zootaxa</mods:title>
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<mods:date id="40EB15073D9D525A80C163C2876D2EFF">2023</mods:date>
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<mods:number id="D0796BC3D89570DD750CE3A24067DF9D">2023-11-14</mods:number>
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<mods:number id="055999ED8035B2EE96C84944B3CAF4A4">5372</mods:number>
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<treatment id="414B87B5FFE29879FF63DA7977D2FDD8" LSID="urn:lsid:plazi:treatment:414B87B5FFE29879FF63DA7977D2FDD8" httpUri="http://treatment.plazi.org/id/414B87B5FFE29879FF63DA7977D2FDD8" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
<subSubSection id="81F86528FFE2987BFF63DA797778FB98" box="[151,619,1086,1113]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="C95D36A3FFE2987BFF63DA797778FB98" blockId="49.[151,619,1086,1149]" box="[151,619,1086,1113]" pageId="49" pageNumber="50">
<heading id="921581CFFFE2987BFF63DA797778FB98" bold="true" box="[151,619,1086,1113]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
<emphasis id="FB96EAB1FFE2987BFF63DA797778FB98" bold="true" box="[151,619,1086,1113]" pageId="49" pageNumber="50">
<taxonomicName id="0EE24D20FFE2987BFF63DA797731FB98" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,546,1086,1113]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="humerorotunda" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFE2987BFF63DA797731FB98" bold="true" box="[151,546,1086,1113]" italics="true" pageId="49" pageNumber="50">Edessa (Edessa) humerorotunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A557CAFFE2987BFDDDDA787778FB98" box="[553,619,1087,1113]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE2987BFF63DA247480FBBC" box="[151,403,1123,1149]" pageId="49" pageNumber="50" type="description">
<paragraph id="C95D36A3FFE2987BFF63DA247480FBBC" blockId="49.[151,619,1086,1149]" box="[151,403,1123,1149]" pageId="49" pageNumber="50">
(
<figureCitation id="51D92A26FFE2987BFF6BDA2475EDFBBC" box="[159,254,1123,1149]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="49" pageNumber="50">Figs. 22</figureCitation>
,
<figureCitation id="51D92A26FFE2987BFEFDDA247471FBBC" box="[265,354,1123,1149]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">61 AB</figureCitation>
,
<figureCitation id="51D92A26FFE2987BFE99DA247499FBBC" box="[365,394,1123,1149]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="49" pageNumber="50">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE2987BFF63DAE3741EFB23" pageId="49" pageNumber="50" type="etymology">
<paragraph id="C95D36A3FFE2987BFF63DAE3741EFB23" blockId="49.[151,1437,1188,2043]" pageId="49" pageNumber="50">
<emphasis id="FB96EAB1FFE2987BFF63DAE3740BFB7F" bold="true" box="[151,280,1188,1214]" pageId="49" pageNumber="50">Etymology</emphasis>
. The name refers to the rounded shape of the humeral angles (L.
<emphasis id="FB96EAB1FFE2987BFBDDDAE3719AFB7F" box="[1065,1161,1188,1214]" italics="true" pageId="49" pageNumber="50">humerus</emphasis>
, shoulder; L.
<emphasis id="FB96EAB1FFE2987BFAC7DAE3708FFB7F" box="[1331,1436,1188,1214]" italics="true" pageId="49" pageNumber="50">rotundus,</emphasis>
spherical).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE2987BFF33DAAB744BFAEB" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="C95D36A3FFE2987BFF33DAAB744BFAEB" blockId="49.[151,1437,1188,2043]" pageId="49" pageNumber="50">
<emphasis id="FB96EAB1FFE2987BFF33DAAB74A5FAC7" bold="true" box="[199,438,1260,1286]" pageId="49" pageNumber="50">Material examined.</emphasis>
<materialsCitation id="798A3CFEFFE2987BFE35DAAB744BFAEB" collectingDate="1970-05-07" collectionCode="INPA" collectorName="I. S. Gorayeb" country="Brazil" location="Amazonas" municipality="Ceplac" pageId="49" pageNumber="50" specimenCount="1" typeStatus="holotype">
<typeStatus id="16598801FFE2987BFE35DAAB773AFAC7" box="[449,553,1260,1286]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
male.
<emphasis id="FB96EAB1FFE2987BFD8BDAAB7691FAC7" bold="true" box="[639,898,1260,1286]" pageId="49" pageNumber="50">
<collectingCountry id="B1F57633FFE2987BFD8BDAAB77F9FAC7" box="[639,746,1260,1286]" name="Brazil" pageId="49" pageNumber="50">BRAZIL</collectingCountry>
,
<location id="CC3D6078FFE2987BFD08DAAB766EFAC7" LSID="urn:lsid:plazi:treatment:414B87B5FFE29879FF63DA7977D2FDD8:CC3D6078FFE2987BFD08DAAB766EFAC7" box="[764,893,1260,1286]" country="Brazil" municipality="Ceplac" name="Amazonas" pageId="49" pageNumber="50">Amazonas</location>
:
</emphasis>
<collectingMunicipality id="2939ACD9FFE2987BFC7ADAAB76CCFAC7" box="[910,991,1260,1286]" pageId="49" pageNumber="50">Ceplac</collectingMunicipality>
, Rod. AM-01 km 30,
<date id="BD5C1063FFE2987BFB18DAAB704AFAC7" box="[1260,1369,1260,1286]" pageId="49" pageNumber="50" value="1970-05-07">
<collectingDate id="AD18E98BFFE2987BFB18DAAB704AFAC7" box="[1260,1369,1260,1286]" pageId="49" pageNumber="50" value="1970-05-07">7-V-1970</collectingDate>
</date>
,
<collectorName id="64175375FFE2987BFA9DDAAA75E4FAEB" pageId="49" pageNumber="50">I. S. Gorayeb</collectorName>
(
<collectionCode id="AFF3AE66FFE2987BFEF2DB57745FFAEB" box="[262,332,1296,1322]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="49" pageNumber="50">INPA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE2987BFF33DB73745BFA56" pageId="49" pageNumber="50" type="description">
<paragraph id="C95D36A3FFE2987BFF33DB73745BFA56" blockId="49.[151,1437,1188,2043]" pageId="49" pageNumber="50">
<emphasis id="FB96EAB1FFE2987BFF33DB737465FA8F" bold="true" box="[199,374,1332,1358]" pageId="49" pageNumber="50">Measurements</emphasis>
(n= 1). Total length: 19.3; head length: 1.8; head width: 3.8; pronotum length: 4.3; pronotum width: 14.0; scutellum length: 8.8; scutellum width: 7.0; abdominal width: 10.5; length antennomeres: I: 1.0; II: 2.0; III: 2.0; IV: 4.5.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE29879FF33DBE7741EFE6C" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="C95D36A3FFE2987BFF33DBE7701AF83B" blockId="49.[151,1437,1188,2043]" pageId="49" pageNumber="50">
<emphasis id="FB96EAB1FFE2987BFF33DBE77451FA7B" bold="true" box="[199,322,1440,1466]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FFE2987BFE6EDBE77717FA7B" box="[410,516,1440,1466]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="49" pageNumber="50" unit="mm" value="19.3">19.3 mm</quantity>
). Dorsal body surface olive green, densely punctured (
<figureCitation id="51D92A26FFE2987BFB9BDBE771CDFA7B" box="[1135,1246,1440,1466]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
). Ventral surface dark yellow to brown with transversal black bands on thorax and abdomen (
<figureCitation id="51D92A26FFE2987BFC16DB83715EFA1F" box="[994,1101,1476,1502]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Head with concolorous with the surrounding surface to brown punctures. Antennae reddish brown (
<figureCitation id="51D92A26FFE2987BFC44DBAF710DF9C3" box="[944,1054,1512,1538]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
). Pronotum with brown punctures (
<figureCitation id="51D92A26FFE2987BFF6BD84B7403F9E7" box="[159,272,1548,1574]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
); anterolateral margin and cicatrices with black punctures. Humeral angles short (1.2 times wider than long); apex with dark brown restricted to the angles in dorsal view; slightly bent backward (
<figureCitation id="51D92A26FFE2987BFB78D877700AF98B" box="[1164,1305,1584,1610]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 AB</figureCitation>
). Scutellum with brown to black punctures; posterior part excavated medially; apex not reaching the end of coria (
<figureCitation id="51D92A26FFE2987BFAEBD8137083F9AF" box="[1311,1424,1620,1646]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
). Coria with all veins concolorous with the surrounding surface (
<figureCitation id="51D92A26FFE2987BFC91D83F76C5F953" box="[869,982,1656,1682]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
). Connexival segments with concavities almost entirely covered by subrectangular black spots separated by a large yellow median spot (
<figureCitation id="51D92A26FFE2987BFB29D8DB705EF977" box="[1245,1357,1692,1718]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
), spots extending ventrally, subrectangular (
<figureCitation id="51D92A26FFE2987BFDCDD88777B8F91B" box="[569,683,1728,1754]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="51D92A26FFE2987BFABDD88775BCF93F" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 A</figureCitation>
). Ventral surface, thorax with black bands; dark band of the propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FFE2987BFF6BD94F7403F8E3" box="[159,272,1800,1826]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Proepisternum, densely punctured, with dark band (
<figureCitation id="51D92A26FFE2987BFC81D94F76F5F8E3" box="[885,998,1800,1826]" captionStart="FIGURE 60" captionStartId="113.[152,255,1943,1967]" captionTargetBox="[252,1335,187,1913]" captionTargetId="figure-20@113.[246,1341,181,1919]" captionTargetPageId="113" captionText="FIGURE 60. AF. Dorsal and ventral surface:AB, Edessa (E.) glomerosa sp. n.; CD, Edessa (E.) guatemalensis sp. n.; EF, Edessa (E.) humeroglobosa sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146471" httpUri="https://zenodo.org/record/10146471/files/figure.png" pageId="49" pageNumber="50">Fig. 60 B</figureCitation>
). Evaporatorium concolorous with the surrounding surface (
<figureCitation id="51D92A26FFE2987BFE79D96B7713F887" box="[397,512,1836,1862]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FFE2987BFCFAD96B766EF887" box="[782,893,1836,1862]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="49" pageNumber="50">Fig. 22 F</figureCitation>
) with arms of anterior bifurcation straight and laterally well expanded at apex; anterior bifurcation slightly excavated receiving fourth and a small part of the third rostral segment. Legs with tibiae and tarsi reddish, remaining yellow to orange (
<figureCitation id="51D92A26FFE2987BFBE2D9337191F84F" box="[1046,1154,1908,1934]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Abdomen with spine of segment III rounded (
<figureCitation id="51D92A26FFE2987BFE72D9DF74EAF872" box="[390,505,1944,1971]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="49" pageNumber="50">Figs. 22 F</figureCitation>
;
<figureCitation id="51D92A26FFE2987BFDF7D9DF7729F873" box="[515,570,1944,1970]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">61 B</figureCitation>
). Intersegmental areas covered by narrow solid black bands with well-defined margins, reaching the lateral spots (
<figureCitation id="51D92A26FFE2987BFDDAD9FB778EF817" box="[558,669,1980,2006]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Pseudosutures solid black with well-defined margins (
<figureCitation id="51D92A26FFE2987BFAEBD9FB709DF817" box="[1311,1422,1980,2006]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Median longitudinal black band incomplete (
<figureCitation id="51D92A26FFE2987BFD6AD9A7761DF83B" box="[670,782,2016,2042]" captionStart="FIGURE 61" captionStartId="114.[152,255,1943,1967]" captionTargetBox="[266,1321,184,1913]" captionTargetId="figure-22@114.[260,1327,181,1919]" captionTargetPageId="114" captionText="FIGURE 61. AF. Dorsal and ventral surface:AB, Edessa (E.) humerorotunda sp. n.; CD, Edessa (E.) illuminocornis sp. n.; EF, Edessa (E.) imitans sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146473" httpUri="https://zenodo.org/record/10146473/files/figure.png" pageId="49" pageNumber="50">Fig. 61 B</figureCitation>
). Trichobothria both parallel to the spiracle.
</paragraph>
<caption id="9D9D662BFFE19878FF6CD9D077F3F812" ID-DOI="http://doi.org/10.5281/zenodo.10146391" ID-Zenodo-Dep="10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="50" pageNumber="51" startId="50.[152,255,1943,1967]" targetBox="[201,1393,192,1914]" targetPageId="50" targetType="figure">
<paragraph id="C95D36A3FFE19878FF6CD9D077F3F812" blockId="50.[151,1436,1943,2004]" pageId="50" pageNumber="51">
<emphasis id="FB96EAB1FFE19878FF6CD9D07430F86E" bold="true" box="[152,291,1943,1967]" pageId="50" pageNumber="51">FIGURE 22.</emphasis>
AG.
<taxonomicName id="0EE24D20FFE19878FE94D9D07762F86E" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[352,625,1943,1967]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="humerorotunda" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFE19878FE94D9D07762F86E" box="[352,625,1943,1967]" italics="true" pageId="50" pageNumber="51">Edessa (E.) humerorotunda</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFE19878FD80D9DF77B8F871" bold="true" box="[628,683,1944,1968]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="E0A557CAFFE19878FD80D9DF77B8F871" box="[628,683,1944,1968]" pageId="50" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
:AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. Scale= 1 mm.
</paragraph>
</caption>
<paragraph id="C95D36A3FFE09879FF63DED0741EFE6C" blockId="51.[151,1437,151,537]" pageId="51" pageNumber="52">
Male genitalia, posterolateral angles of the pygophore developed, projected laterally, apices internally with spots blacks (
<figureCitation id="51D92A26FFE09879FF1BDEFC7493FF14" box="[239,384,187,213]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 AC</figureCitation>
). Superior processes of the genital cup subrectangular, thick, in flattened, coarse and slightly concave posterior view, continuing ventrally in a crenulate, long and high carina (
<figureCitation id="51D92A26FFE09879FBDBDE9871CDFF38" box="[1071,1246,223,249]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 B, DE</figureCitation>
, crenulate carina barely visible). Parameres (
<figureCitation id="51D92A26FFE09879FE21DF44777AFEDC" box="[469,617,259,285]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 DE</figureCitation>
) with black margins; anterior lobe rounded; dorsal lobe subtriangular; posterior lobe subrectangular, with curved apex. Proctiger, posterior face subtriangular, reddish posterior half (
<figureCitation id="51D92A26FFE09879FA9BDF6075E1FEA4" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 DE</figureCitation>
). Ventral surface densely punctuated; presence of dark spots laterally and medially (
<figureCitation id="51D92A26FFE09879FB69DF0C7019FEA4" box="[1181,1290,331,357]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 C</figureCitation>
). Ventral rim setose, with medial tuft (
<figureCitation id="51D92A26FFE09879FE44DF287771FE48" box="[432,610,367,393]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 A, CE</figureCitation>
); with expansions undeveloped and brown (
<figureCitation id="51D92A26FFE09879FBA1DF2871D6FE48" box="[1109,1221,367,393]" captionStart="FIGURE 22" captionStartId="50.[152,255,1943,1967]" captionTargetBox="[201,1393,192,1914]" captionTargetId="figure-22@50.[195,1393,181,1920]" captionTargetPageId="50" captionText="FIGURE 22.AG. Edessa(E.) humerorotundasp.n.:AE,pygophore indorsal,dorsolateral,ventral, posteriorandposterolateral views, respectively; F, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146391" httpUri="https://zenodo.org/record/10146391/files/figure.png" pageId="51" pageNumber="52">Fig. 22 C</figureCitation>
). Female genitalia. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE09879FF33DFF175A6FE34" pageId="51" pageNumber="52" type="discussion">
<paragraph id="C95D36A3FFE09879FF33DFF175A6FE34" blockId="51.[151,1437,151,537]" pageId="51" pageNumber="52">
<emphasis id="FB96EAB1FFE09879FF33DFF1745CFE11" bold="true" box="[199,335,438,464]" pageId="51" pageNumber="52">Comments.</emphasis>
See comments on
<taxonomicName id="0EE24D20FFE09879FDD5DFF077E9FE11" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[545,762,439,465]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="crassicornis" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFE09879FDD5DFF077E9FE11" box="[545,762,439,465]" italics="true" pageId="51" pageNumber="52">E. (E.) crassicornis</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFE09879FCF4DFF07653FE10" bold="true" box="[768,832,439,465]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="E0A557CAFFE09879FCF4DFF07653FE10" box="[768,832,439,465]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="0EE24D20FFE09879FCB9DFF07159FE10" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[845,1098,439,465]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="humeroglobosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFE09879FCB9DFF07159FE10" box="[845,1098,439,465]" italics="true" pageId="51" pageNumber="52">E. (E.) humeroglobosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFE09879FBA4DFF0719CFE10" bold="true" box="[1104,1167,439,465]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="E0A557CAFFE09879FBA4DFF0719CFE10" box="[1104,1167,439,465]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="0EE24D20FFE09879FB38DFF07066FE10" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1228,1397,439,465]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="limeirai" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFE09879FB38DFF07066FE10" box="[1228,1397,439,465]" italics="true" pageId="51" pageNumber="52">E. (E.) limeirai</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFE09879FA8FDFF075BDFE34" bold="true" pageId="51" pageNumber="52">
<taxonomicNameLabel id="E0A557CAFFE09879FA8FDFF075BDFE34" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFE09879FF33DFB977D2FDD8" box="[199,705,510,537]" pageId="51" pageNumber="52" type="distribution">
<paragraph id="C95D36A3FFE09879FF33DFB977D2FDD8" blockId="51.[151,1437,151,537]" box="[199,705,510,537]" pageId="51" pageNumber="52">
<emphasis id="FB96EAB1FFE09879FF33DFB9744AFDD9" bold="true" box="[199,345,510,536]" pageId="51" pageNumber="52">Distribution</emphasis>
(
<figureCitation id="51D92A26FFE09879FE9DDFB874A9FDD8" box="[361,442,511,537]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="51" pageNumber="52">Fig. 72</figureCitation>
).
<collectingCountry id="B1F57633FFE09879FE24DFB87729FDD8" box="[464,570,511,537]" name="Brazil" pageId="51" pageNumber="52">BRAZIL</collectingCountry>
: Amazonas.
</paragraph>
</subSubSection>
</treatment>
</document>