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<document id="C5F34E67D887F97F9816719419EEA150" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FFCF9854FF63D881769CF97C" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) echyra Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="29" updateTime="1700188109802" updateUser="ExternalLinkService">
<mods:mods id="539AB0ABBDAC10A3B2B2CD62FCD0CA09" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9AA359A774AB1799E137B2CCE4BDD7D">
<mods:title id="0DB0E6E3FF02D1A246111D0C5374AC3E">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
</mods:titleInfo>
<mods:name id="E1386A0632BED1DA77085CA36C349258" type="personal">
<mods:role id="B562F413720B4A665B33CCE444C0B627">
<mods:roleTerm id="8C4C6D690065D39DA674731844C5937D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA0EE124491A20B8DC6AF23FD01121DF">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="8D506CF761899E5BF6F5C154F09233DD">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
</mods:name>
<mods:name id="A5B8E7A62ED8C977958CAA2E4DF1A4DC" type="personal">
<mods:role id="64B15D6B646D9E79C8050BF36B3089D1">
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</mods:role>
<mods:namePart id="2E4F78678E3D0ED37C1398265FBD29EF">Silva, Valéria Juliete Da</mods:namePart>
<mods:nameIdentifier id="A78018D036AFE2EB320ABCD56858ED56" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
<mods:affiliation id="D205182C66C73AD7E8055A05EF736D7F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
<mods:nameIdentifier id="BD40C7CA891CB94AF6297143E694B0E3" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B8652EA59EF3BE11E51F9F797B3C99BA" type="personal">
<mods:role id="B6F1A62D6618FDC9EFC194C42030FB7C">
<mods:roleTerm id="548ED4C80B040634664984B639477CAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="652720BE16EE9C4EA9E0792D408F9AB5">Fernandes, José Antônio Marin</mods:namePart>
<mods:nameIdentifier id="1424AF45E9979C8BB1BEBF4B9FA354D4" type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
<mods:affiliation id="341CE4E776DEE8F14D2F5B3C2473DA74">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
<mods:nameIdentifier id="8676258C3070FBE9A01F8CC7ECE3186D" type="email">joseamf@ufpa.br</mods:nameIdentifier>
</mods:name>
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<mods:title id="9D8683865EA5FB568C35B78DAEB10387">Zootaxa</mods:title>
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<mods:part id="7640C0D4ECE1D2993F5C38797E45F1F5">
<mods:date id="8141FBBEFCE7A048EE2D51CA70BECFF7">2023</mods:date>
<mods:detail id="C9222EFC00D2A15BD104403B49766DF2" type="pubDate">
<mods:number id="F5ECAF2743206B0AEF731F29A802CE43">2023-11-14</mods:number>
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<mods:detail id="FD5CEBB4051567BF1478B78159EBFC54" type="volume">
<mods:number id="ADEB7BE01A2EA8B5B9476C090F7D0CC2">5372</mods:number>
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<mods:number id="25FD65D95FCEB8EEAAE5E7C44CBCEF31">1</mods:number>
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<mods:url id="CBAD3B200D81E7863F56BB96A777B2B1">https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272</mods:url>
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<mods:identifier id="10EDEED3F994B6620FE09AE69DA75D82" type="DOI">10.11646/zootaxa.5372.1.1</mods:identifier>
<mods:identifier id="BEB5EF96DE23D7AEBD74211D4621DD0F" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="80C928EC77BE44CD9CD9F1689321E3D5" type="Zenodo-Dep">10146342</mods:identifier>
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<treatment id="414B87B5FFCF9854FF63D881769CF97C" LSID="urn:lsid:plazi:treatment:414B87B5FFCF9854FF63D881769CF97C" httpUri="http://treatment.plazi.org/id/414B87B5FFCF9854FF63D881769CF97C" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="81F86528FFCF9856FF63D88174EDF920" box="[151,510,1734,1761]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="C95D36A3FFCF9856FF63D88174EDF920" blockId="28.[151,510,1734,1797]" box="[151,510,1734,1761]" pageId="28" pageNumber="29">
<heading id="921581CFFFCF9856FF63D88174EDF920" bold="true" box="[151,510,1734,1761]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="FB96EAB1FFCF9856FF63D88174EDF920" bold="true" box="[151,510,1734,1761]" pageId="28" pageNumber="29">
<taxonomicName id="0EE24D20FFCF9856FF63D88174A7F920" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,436,1734,1761]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="echyra" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFCF9856FF63D88174A7F920" bold="true" box="[151,436,1734,1761]" italics="true" pageId="28" pageNumber="29">Edessa (Edessa) echyra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A557CAFFCF9856FE48D88074EDF920" box="[444,510,1735,1761]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCF9856FF63D8AC7480F8C4" box="[151,403,1770,1797]" pageId="28" pageNumber="29" type="description">
<paragraph id="C95D36A3FFCF9856FF63D8AC7480F8C4" blockId="28.[151,510,1734,1797]" box="[151,403,1770,1797]" pageId="28" pageNumber="29">
(
<figureCitation id="51D92A26FFCF9856FF6BD8AC75EFF8C4" box="[159,252,1771,1797]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="28" pageNumber="29">Figs. 11</figureCitation>
,
<figureCitation id="51D92A26FFCF9856FEFCD8AD7471F8C4" box="[264,354,1770,1797]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="28" pageNumber="29">57 C–D</figureCitation>
,
<figureCitation id="51D92A26FFCF9856FE9AD8AC7498F8C4" box="[366,395,1771,1797]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="28" pageNumber="29">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCF9856FF63D97476A2F88C" box="[151,945,1843,1869]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="C95D36A3FFCF9856FF63D97476A2F88C" blockId="28.[151,1437,1843,2013]" box="[151,945,1843,1869]" pageId="28" pageNumber="29">
<emphasis id="FB96EAB1FFCF9856FF63D974740BF88C" bold="true" box="[151,280,1843,1869]" pageId="28" pageNumber="29">Etymology</emphasis>
. The name refers to robust body size (Gr.
<emphasis id="FB96EAB1FFCF9856FD05D974765FF88C" box="[753,844,1843,1869]" italics="true" pageId="28" pageNumber="29">Echyros</emphasis>
, strong).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCF9856FF33D911768DF81C" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="C95D36A3FFCF9856FF33D91175E5F854" blockId="28.[151,1437,1843,2013]" pageId="28" pageNumber="29">
<emphasis id="FB96EAB1FFCF9856FF33D91174ABF8B1" bold="true" box="[199,440,1878,1904]" pageId="28" pageNumber="29">Material examined.</emphasis>
<materialsCitation id="798A3CFEFFCF9856FE33D91075E5F854" collectingDate="1991-01-29" collectionCode="UFES" collectorName="J. S. Santos" country="Brazil" location="Linhares" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<typeStatus id="16598801FFCF9856FE33D910773CF8B0" box="[455,559,1879,1905]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male.
<emphasis id="FB96EAB1FFCF9856FD7ED91176DBF8B1" bold="true" box="[650,968,1878,1905]" pageId="28" pageNumber="29">
<collectingCountry id="B1F57633FFCF9856FD7ED91177E6F8B1" box="[650,757,1878,1904]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
,
<collectingRegion id="0B26F841FFCF9856FCFFD91176D0F8B1" box="[779,963,1878,1905]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="29">Espírito Santo</collectingRegion>
:
</emphasis>
<location id="CC3D6078FFCF9856FC22D9107128F8B0" LSID="urn:lsid:plazi:treatment:414B87B5FFCF9854FF63D881769CF97C:CC3D6078FFCF9856FC22D9107128F8B0" box="[982,1083,1879,1905]" country="Brazil" name="Linhares" pageId="28" pageNumber="29" stateProvince="Espirito Santo">Linhares</location>
,
<date id="BD5C1063FFCF9856FBBAD91071D3F8B0" box="[1102,1216,1879,1905]" pageId="28" pageNumber="29" value="1991-01-29">
<collectingDate id="AD18E98BFFCF9856FBBAD91071D3F8B0" box="[1102,1216,1879,1905]" pageId="28" pageNumber="29" value="1991-01-29">29-I-1991</collectingDate>
</date>
,
<collectorName id="64175375FFCF9856FB20D9107076F8B0" box="[1236,1381,1879,1905]" pageId="28" pageNumber="29">J. S. Santos</collectorName>
col. (
<collectionCode id="AFF3AE66FFCF9856FF6BD93C75FAF854" box="[159,233,1915,1941]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="28" pageNumber="29">UFES</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="C95D36A3FFCF9856FF33D9D8768DF81C" blockId="28.[151,1437,1843,2013]" pageId="28" pageNumber="29">
<materialsCitation id="798A3CFEFFCF9856FF33D9D8708BF878" box="[199,1432,1950,1977]" collectingDate="1939-11-12" collectingDateMax="1939-11-15" collectingDateMin="1939-11-12" collectionCode="CEIOC" collectorName="Martins &amp; Lopes" country="Brazil" location="Rio Doce" municipality="Calado" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="16598801FFCF9856FF33D9D8742BF878" box="[199,312,1951,1977]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
.
<emphasis id="FB96EAB1FFCF9856FEB3D9D97767F879" bold="true" box="[327,628,1950,1977]" pageId="28" pageNumber="29">
<collectingCountry id="B1F57633FFCF9856FEB3D9D974A1F879" box="[327,434,1950,1976]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
,
<collectingRegion id="0B26F841FFCF9856FE31D9D9777CF879" box="[453,623,1950,1976]" country="Brazil" name="Minas Gerais" pageId="28" pageNumber="29">Minas Gerais</collectingRegion>
:
</emphasis>
<specimenCount id="DFE4FD2AFFCF9856FD8BD9D877B1F878" box="[639,674,1951,1977]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
,
<collectingMunicipality id="2939ACD9FFCF9856FD41D9D87614F878" box="[693,775,1951,1977]" pageId="28" pageNumber="29">Calado</collectingMunicipality>
,
<location id="CC3D6078FFCF9856FCE3D9D8769AF878" LSID="urn:lsid:plazi:treatment:414B87B5FFCF9854FF63D881769CF97C:CC3D6078FFCF9856FCE3D9D8769AF878" box="[791,905,1951,1977]" country="Brazil" municipality="Calado" name="Rio Doce" pageId="28" pageNumber="29" stateProvince="Minas Gerais">Rio Doce</location>
,
<date id="BD5C1063FFCF9856FC6DD9D9715AF878" box="[921,1097,1950,1977]" pageId="28" pageNumber="29" value="1939-11-12" valueMax="1939-11-15" valueMin="1939-11-12">
<collectingDate id="AD18E98BFFCF9856FC6DD9D9715AF878" box="[921,1097,1950,1977]" pageId="28" pageNumber="29" value="1939-11-12" valueMax="1939-11-15" valueMin="1939-11-12">12–15-XI-1939</collectingDate>
</date>
,
<collectorName id="64175375FFCF9856FBAED9D871A2F878" box="[1114,1201,1951,1977]" pageId="28" pageNumber="29">Martins</collectorName>
&amp;
<collectorName id="64175375FFCF9856FB2AD9D87031F878" box="[1246,1314,1951,1977]" pageId="28" pageNumber="29">Lopes</collectorName>
(
<collectionCode id="AFF3AE66FFCF9856FAC2D9D87083F878" box="[1334,1424,1951,1977]" pageId="28" pageNumber="29">CEIOC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="798A3CFEFFCF9856FF63D985768AF81C" box="[151,921,1986,2013]" collectingDate="1990-09-03" collectionCode="UFES" collectorName="J. S. Santos" country="Brazil" location="Linhares" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Espirito Santo" typeStatus="paratype">
<emphasis id="FB96EAB1FFCF9856FF63D985745FF81D" bold="true" box="[151,332,1986,2013]" pageId="28" pageNumber="29">
<collectingRegion id="0B26F841FFCF9856FF63D985745BF81D" box="[151,328,1986,2013]" country="Brazil" name="Espirito Santo" pageId="28" pageNumber="29">Espírito Santo</collectingRegion>
:
</emphasis>
<specimenCount id="DFE4FD2AFFCF9856FEA7D9847465F81C" box="[339,374,1987,2013]" count="1" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
,
<location id="CC3D6078FFCF9856FE70D98474FAF81C" LSID="urn:lsid:plazi:treatment:414B87B5FFCF9854FF63D881769CF97C:CC3D6078FFCF9856FE70D98474FAF81C" box="[388,489,1987,2013]" country="Brazil" name="Linhares" pageId="28" pageNumber="29" stateProvince="Espirito Santo">Linhares</location>
,
<date id="BD5C1063FFCF9856FE00D9847768F81C" box="[500,635,1987,2013]" pageId="28" pageNumber="29" value="1990-09-03">
<collectingDate id="AD18E98BFFCF9856FE00D9847768F81C" box="[500,635,1987,2013]" pageId="28" pageNumber="29" value="1990-09-03">03-IX-1990</collectingDate>
</date>
,
<collectorName id="64175375FFCF9856FD73D984761BF81C" box="[647,776,1987,2013]" pageId="28" pageNumber="29">J. S. Santos</collectorName>
col. (
<collectionCode id="AFF3AE66FFCF9856FCB3D9847682F81C" box="[839,913,1987,2013]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="28" pageNumber="29">UFES</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="9D9D662BFFCE9857FF6CD9D07680F812" ID-DOI="http://doi.org/10.5281/zenodo.10146369" ID-Zenodo-Dep="10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="29" pageNumber="30" startId="29.[152,255,1943,1967]" targetBox="[189,1398,192,1911]" targetPageId="29" targetType="figure">
<paragraph id="C95D36A3FFCE9857FF6CD9D07680F812" blockId="29.[151,1436,1943,2004]" pageId="29" pageNumber="30">
<emphasis id="FB96EAB1FFCE9857FF6CD9D07435F86E" bold="true" box="[152,294,1943,1967]" pageId="29" pageNumber="30">FIGURE 11.</emphasis>
A–G.
<taxonomicName id="0EE24D20FFCE9857FE9BD9DF7727F871" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[367,564,1944,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="echyra" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFCE9857FE9BD9DF7727F871" box="[367,564,1944,1968]" italics="true" pageId="29" pageNumber="30">Edessa (E.) echyra</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCE9857FDC9D9DF776AF871" bold="true" box="[573,633,1944,1968]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="E0A557CAFFCE9857FDC9D9DF776AF871" box="[573,633,1944,1968]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm.
</paragraph>
</caption>
<subSubSection id="81F86528FFCD9854FF33DED076E0FF38" pageId="30" pageNumber="31" type="description">
<paragraph id="C95D36A3FFCD9854FF33DED076E0FF38" blockId="30.[151,1437,150,1725]" pageId="30" pageNumber="31">
<emphasis id="FB96EAB1FFCD9854FF33DED07465FF70" bold="true" box="[199,374,151,177]" pageId="30" pageNumber="31">Measurements</emphasis>
(n= 3). Total length: 21.1–23.4; head length: 1.7; head width: 3.5–3.9; pronotum length: 4.6– 5.3; pronotum width: 16.5–17.2; scutellum length: 9.6–10.4; scutellum width: 7.5–8.0; abdominal width: 12.5–13.3; length antennomeres: I: 1.0; II: 2.0–2.2; III: 2.0–2.0; IV: 4.9–5.0; V: 5.3–5.3.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCD9854FF33DF457776FAF0" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="C95D36A3FFCD9854FF33DF457776FAF0" blockId="30.[151,1437,150,1725]" pageId="30" pageNumber="31">
<emphasis id="FB96EAB1FFCD9854FF33DF457451FEDD" bold="true" box="[199,322,258,284]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FFCD9854FE51DF447747FEDC" box="[421,596,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.225" metricValueMax="2.34" metricValueMin="2.11" pageId="30" pageNumber="31" unit="mm" value="22.25" valueMax="23.4" valueMin="21.1">21.1–23.4 mm</quantity>
). Dorsal body surface brownish olive, densely punctured (
<figureCitation id="51D92A26FFCD9854FAE1DF44709CFEDC" box="[1301,1423,258,285]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
). Ventral surface dark yellow to dark reddish brown with transversal black bands on thorax and abdomen (
<figureCitation id="51D92A26FFCD9854FA9BDF6075C0FEA4" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
), densely punctured. Antennae reddish brown (
<figureCitation id="51D92A26FFCD9854FD05DF0C7695FEA4" box="[753,902,330,357]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C–D</figureCitation>
). Pronotum with brown punctures (
<figureCitation id="51D92A26FFCD9854FAE9DF0C709DFEA4" box="[1309,1422,330,357]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
); anterolateral margin and cicatrices with black punctures; anterolateral margin coarse. Humeral angles short (1.25 times wider than long), laterally directed; apex with black spot restricted to the angles in dorsal view, reddish brown in ventral view; abruptly bent backward (
<figureCitation id="51D92A26FFCD9854FD9EDFF077EFFE10" box="[618,764,438,465]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C–D</figureCitation>
). Scutellum with brown punctures; posterior part excavated medially; apex not reaching the end of coria (
<figureCitation id="51D92A26FFCD9854FD7FDF9C77E5FE34" box="[651,758,474,501]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
). Coria with all veins slightly lighter than surface (
<figureCitation id="51D92A26FFCD9854FAD0DF9C709CFE34" box="[1316,1423,474,501]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
). Connexival segments with concavities almost entirely covered by subrectangular black spots punctured separated by a large reddish median spot (
<figureCitation id="51D92A26FFCD9854FDFEDC64776FFDFC" box="[522,636,546,573]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
); black spots extending ventrally, roundish (
<figureCitation id="51D92A26FFCD9854FB8DDC6471FFFDFC" box="[1145,1260,546,573]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="51D92A26FFCD9854FD82DC0077FBFDA0" box="[630,744,582,609]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 C</figureCitation>
). Ventral surface, thorax with black bands; dark band of the propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FFCD9854FD28DC2C765DFD44" box="[732,846,618,645]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Proepisternum coarse, with dark band (
<figureCitation id="51D92A26FFCD9854FAEADC2C7083FD44" box="[1310,1424,618,645]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Evaporatorium concolorous with the surrounding surface (
<figureCitation id="51D92A26FFCD9854FCDDDCC87689FD68" box="[809,922,654,681]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FFCD9854FB69DCC87000FD68" box="[1181,1299,655,681]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Figs. 11 G</figureCitation>
,
<figureCitation id="51D92A26FFCD9854FAD6DCC97049FD68" box="[1314,1370,654,681]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">57 D</figureCitation>
) with arms of anterior bifurcation straight and laterally well expanded, margin acuminated at apex; anterior bifurcation broadly excavated receiving fourth rostral segment. Legs with femora darker than the other articles (
<figureCitation id="51D92A26FFCD9854FAE8DC907083FD30" box="[1308,1424,726,753]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Abdomen with spine of segment III rounded (
<figureCitation id="51D92A26FFCD9854FD5ADCBC763AFCD4" box="[686,809,763,789]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Figs. 11 G</figureCitation>
,
<figureCitation id="51D92A26FFCD9854FCCFDCBD766AFCD4" box="[827,889,762,789]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">57 D</figureCitation>
). Intersegmental areas covered by narrow solid black bands with well-defined margins, not reaching the lateral spots (
<figureCitation id="51D92A26FFCD9854FC43DD58713AFCF8" box="[951,1065,798,825]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Pseudosutures solid black with well-defined margins (
<figureCitation id="51D92A26FFCD9854FE62DD047715FC9C" box="[406,518,834,861]" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Median longitudinal brown band incomplete, restricted to the segment VII (
<figureCitation id="51D92A26FFCD9854FA9BDD0475C1FC40" captionStart="FIGURE 57" captionStartId="110.[152,255,1943,1967]" captionTargetBox="[254,1334,187,1913]" captionTargetId="figure-22@110.[248,1340,181,1919]" captionTargetPageId="110" captionText="FIGURE 57. A–F. Dorsal and ventral surface:A–B, Edessa (E.) decolorata sp. n.; C–D, Edessa (E.) echyra sp. n.; E–F, Edessa (E.) fabricii sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146463" httpUri="https://zenodo.org/record/10146463/files/figure.png" pageId="30" pageNumber="31">Fig. 57 D</figureCitation>
). Trichobothria one in line with spiracle and the other laterad. Posterolateral angles of segment VII at the sam level as the apices of laterotergites IX (
<figureCitation id="51D92A26FFCD9854FD94DDCC77DDFC64" box="[608,718,907,933]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 F</figureCitation>
). Male genitalia, dorsal side of the pygophore with a suffused brown area occupying 1/3 of the surface (
<figureCitation id="51D92A26FFCD9854FD90DDE877E3FC08" box="[612,752,943,969]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 A–B</figureCitation>
). Posterolateral angles of the pygophore developed, projected laterally (
<figureCitation id="51D92A26FFCD9854FEF6DD94749BFC2C" box="[258,392,979,1005]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 A,C</figureCitation>
). Superior processes of the genital cup subrectangular, thick, in flattened and coarse posterior view (
<figureCitation id="51D92A26FFCD9854FF15DDB07447FBD0" box="[225,340,1015,1041]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 B</figureCitation>
), continuing ventrally in a crenulate, long and high carina (
<figureCitation id="51D92A26FFCD9854FBE0DDB071BBFBD0" box="[1044,1192,1015,1041]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 D–E</figureCitation>
, crenulate carina not visible). Parameres with brown margins; anterior lobe inconspicuous and rounded; dorsal lobe acuminated and curved laterally; posterior lobe subrectangular, with curved apex. Proctiger, posterior face triangular (
<figureCitation id="51D92A26FFCD9854FAF7DA787082FB98" box="[1283,1425,1087,1113]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 D–E</figureCitation>
). Ventral surface with a suffused brown area occupying 1/2 of the surface; densely punctuated; presence of dark spots laterally and medially (
<figureCitation id="51D92A26FFCD9854FE6FDAC07714FB60" box="[411,519,1159,1185]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 C</figureCitation>
). Ventral rim setose, with medial tuft (
<figureCitation id="51D92A26FFCD9854FC41DAC0712EFB60" box="[949,1085,1159,1185]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11A–C</figureCitation>
); with expansions undeveloped and brown with surface (
<figureCitation id="51D92A26FFCD9854FE5EDAEC7707FB04" box="[426,532,1195,1221]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 C</figureCitation>
). Female genitalia, densely punctured. Valvifers VIII with dark punctures; sutural margins brown, contiguous, and not divergent; posterior margin brown and subrectangular; with brown spot in tumescence. Laterotergites VIII with dark band on outer lateral margins. Laterotergites IX with apices acuminate passing the mediotergite VIII (
<figureCitation id="51D92A26FFCD9854FE1ADB50774BFAF0" box="[494,600,1303,1329]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="30" pageNumber="31">Fig. 11 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCD9854FF33DB7D74E6F958" pageId="30" pageNumber="31" type="discussion">
<paragraph id="C95D36A3FFCD9854FF33DB7D74E6F958" blockId="30.[151,1437,150,1725]" pageId="30" pageNumber="31">
<emphasis id="FB96EAB1FFCD9854FF33DB7D745CFA95" bold="true" box="[199,335,1338,1364]" pageId="30" pageNumber="31">Comments.</emphasis>
<taxonomicName id="0EE24D20FFCD9854FEADDB7C7721FA94" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[345,562,1339,1365]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="echyra" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FEADDB7C7721FA94" box="[345,562,1339,1365]" italics="true" pageId="30" pageNumber="31">Edessa (E.) echyra</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FDC8DB7C776DFA94" bold="true" box="[572,638,1339,1365]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FDC8DB7C776DFA94" box="[572,638,1339,1365]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="0EE24D20FFCD9854FCF4DB7C76D8FA94" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[768,971,1339,1365]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FCF4DB7C76D8FA94" box="[768,971,1339,1365]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FC21DB7C7104FA94" bold="true" box="[981,1047,1339,1365]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FC21DB7C7104FA94" box="[981,1047,1339,1365]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the robust and brownish olive body; humeral angles black, curved, projected and globose; coria with veins lighter than surface; body ventrally with intersegmental areas black; dorsal side of the pygophore with a suffused brown area occupying 1/3 of the surface and ventral rim setose, with medial tuft. On the other hand, both species can be easily separated by the black spot on humeral angles restricted to the dorsal surface of the pronotum (in
<taxonomicName id="0EE24D20FFCD9854FC40DB8C716FFA24" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[948,1148,1483,1509]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FC40DB8C716FFA24" box="[948,1148,1483,1509]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FB70DB8C71D6FA24" bold="true" box="[1156,1221,1483,1509]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FB70DB8C71D6FA24" box="[1156,1221,1483,1509]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
black spot slightly expanded); ventral surface of the humeral angles reddish brown (in
<taxonomicName id="0EE24D20FFCD9854FC65DBA87144F9C8" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[913,1111,1519,1545]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FC65DBA87144F9C8" box="[913,1111,1519,1545]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FBABDBA8718CF9C8" bold="true" box="[1119,1183,1519,1545]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FBABDBA8718CF9C8" box="[1119,1183,1519,1545]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
black); proepisternum with dark band (
<taxonomicName id="0EE24D20FFCD9854FEB9D8547702F9EC" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[333,529,1555,1581]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FEB9D8547702F9EC" box="[333,529,1555,1581]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FDE3D8547745F9EC" bold="true" box="[535,598,1555,1581]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FDE3D8547745F9EC" box="[535,598,1555,1581]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
without this band); ventrally intersegmental areas black bands not reaching the lateral spots (
<taxonomicName id="0EE24D20FFCD9854FEA3D870770FF990" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[343,540,1591,1617]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FEA3D870770FF990" box="[343,540,1591,1617]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FDD7D8707770F990" bold="true" box="[547,611,1591,1617]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FDD7D8707770F990" box="[547,611,1591,1617]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
black bands reaching the lateral spots); ventral surface densely punctured (
<taxonomicName id="0EE24D20FFCD9854FF55D81C7463F9B4" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[161,368,1627,1653]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FF55D81C7463F9B4" box="[161,368,1627,1653]" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
lightly punctuated); and the female valvifers VIII with brown spot in tumescence (
<taxonomicName id="0EE24D20FFCD9854FABAD81C741FF958" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="glomerosa" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCD9854FABAD81C741FF958" italics="true" pageId="30" pageNumber="31">E. (E.) glomerosa</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCD9854FEE7D8387440F958" bold="true" box="[275,339,1663,1689]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="E0A557CAFFCD9854FEE7D8387440F958" box="[275,339,1663,1689]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
without spot).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCD9854FF33D8E5769CF97C" box="[199,911,1698,1725]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="C95D36A3FFCD9854FF33D8E5769CF97C" blockId="30.[151,1437,150,1725]" box="[199,911,1698,1725]" pageId="30" pageNumber="31">
<emphasis id="FB96EAB1FFCD9854FF33D8E5744AF97D" bold="true" box="[199,345,1698,1724]" pageId="30" pageNumber="31">Distribution</emphasis>
(
<figureCitation id="51D92A26FFCD9854FE9DD8E474A9F97C" box="[361,442,1699,1725]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="30" pageNumber="31">Fig. 72</figureCitation>
).
<collectingCountry id="B1F57633FFCD9854FE24D8E47729F97C" box="[464,570,1699,1725]" name="Brazil" pageId="30" pageNumber="31">BRAZIL</collectingCountry>
:
<collectingRegion id="0B26F841FFCD9854FDB0D8E477CEF97C" box="[580,733,1699,1725]" country="Brazil" name="Minas Gerais" pageId="30" pageNumber="31">Minas Gerais</collectingRegion>
,
<collectingRegion id="0B26F841FFCD9854FD1CD8E47698F97C" box="[744,907,1699,1725]" country="Brazil" name="Espirito Santo" pageId="30" pageNumber="31">Espirito Santo</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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