treatments-xml/data/41/49/87/414987D12A17FFF8FF21FE2CFC1C7E38.xml

<document id="AC47634DBEDAAF83BDE071995E88B921" ID-CLB-Dataset="55701" ID-DOI="10.11646/zootaxa.5133.2.1" ID-GBIF-Dataset="49bbe6cd-9052-42a3-8da7-1039e8c8de60" ID-ISSN="1175-5326" ID-Zenodo-Dep="6521560" ID-ZooBank="6B6B0F37-8D98-4930-9038-EEE9F0E7B674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651754288764" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Matsuzaki, Koji" docDate="2022" docId="414987D12A17FFF8FF21FE2CFC1C7E38" docLanguage="en" docName="zootaxa.5133.2.1.pdf" docOrigin="Zootaxa 5133 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebbeus subtilis Komai &amp; Matsuzaki 2022, n. sp." docType="treatment" docUuid="F9365A97-E59B-4C1A-B924-14AE5EB1AB26" docUuidSource="ZooBank" docVersion="9" lastPageNumber="167" masterDocId="BD70FFA92A1CFFE8FFB6FFBEFD5C7975" masterDocTitle="Three new species of the thorid shrimp genus Lebbeus White, 1847 (Decapoda: Caridea) from Nemuro Strait, Hokkaido, Japan, with supplemental note on L. fujimotoi Matsuzaki, Hibino &amp; Komai, 2015" masterLastPageNumber="181" masterPageNumber="151" pageNumber="162" updateTime="1701998325065" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BA11E93132F0F3B35DF3ED6AD12F87A9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DF4EAEA95AD2839F04424DC8F452AE94">
<mods:title id="9372A2C60217A9D40EFF4E2FA30C6D09">Three new species of the thorid shrimp genus Lebbeus White, 1847 (Decapoda: Caridea) from Nemuro Strait, Hokkaido, Japan, with supplemental note on L. fujimotoi Matsuzaki, Hibino &amp; Komai, 2015</mods:title>
</mods:titleInfo>
<mods:name id="F5AFB1FEAE2597C582B80900AA86E72B" type="personal">
<mods:role id="C32F16615FDAD1D519351B13F91F023E">
<mods:roleTerm id="FDC66FB87E80272B2C2660660EB983A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA20880E9C9E5BEE4836C38C9D4F039E">Komai, Tomoyuki</mods:namePart>
<mods:nameIdentifier id="B19605568E5C03146E6879A04455BEB5" type="ORCID">0000-0003-0892-2555</mods:nameIdentifier>
<mods:affiliation id="4134ED12B896E2E52ACBACD47471D39E">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, Chiba, 260 - 8682 Japan. komai @ chiba-muse. or. jp; https: // orcid. org / 0000 - 0003 - 0892 - 2555</mods:affiliation>
<mods:nameIdentifier id="7496CB3D96E6A5C2B0E807EFD48F04CA" type="email">komai@chiba-muse.or.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="58AD81E1362086F61F85CD45F463A7C5" type="personal">
<mods:role id="978E4309C81D2187E68A2408B3DA0672">
<mods:roleTerm id="0BE1FB0E84D885550B360EF48A101888">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F887D60C41C3CFFAFEA0FB5C1C525F3">Matsuzaki, Koji</mods:namePart>
<mods:nameIdentifier id="269B1A459A64FD58D636D292D46F94FC" type="ORCID">0000-0002-2583-4293</mods:nameIdentifier>
<mods:affiliation id="0CD6E04B4A9E4D6ACAEF5E334858E159">Marine Science Museum, Fukushima Prefecture, Aquamarine Fukushima, 50 Tatsumi-cho, Onahama, Iwaki, Fukushima, 971 - 8101 Japan. matsu @ aquamarine. or. jp; https: // orcid. org / 0000 - 0002 - 2583 - 4293</mods:affiliation>
<mods:nameIdentifier id="F909B0438DEE94470D802B91D9767B7F" type="email">matsu@aquamarine.or.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1F2F771809B3A030CCB33A5F3FDD19EC">text</mods:typeOfResource>
<mods:relatedItem id="173F54D94B55E8CE44A6AE8C99FED834" type="host">
<mods:titleInfo id="D3D734B848AE848262BAC39B5C1D5A02">
<mods:title id="41505DEDB31207101592EAF5A99C0A6C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FAE6D1A7624046219886BA7E28187A36">
<mods:date id="C0F9DE86BBCD4E65DB20E902AF6FD1F2">2022</mods:date>
<mods:detail id="009A2E5142C9AA4FDE77F648B528470B" type="pubDate">
<mods:number id="C9E2B6CF6AAE79B6838D44E5F4E08539">2022-05-04</mods:number>
</mods:detail>
<mods:detail id="07C563045FE3BFD8CF025C18A2A007E5" type="volume">
<mods:number id="A96EFE32B75BBB7E17B5C04467597171">5133</mods:number>
</mods:detail>
<mods:detail id="B246E662DE19134DEF3A50E482D70878" type="issue">
<mods:number id="D32B5060B0AB1E75D42862F5B1EDA57B">2</mods:number>
</mods:detail>
<mods:extent id="449D7353D9D69858E7B3DF3EE2B08587" unit="page">
<mods:start id="16C3C67277109BD2F11583FD53F8A8C6">151</mods:start>
<mods:end id="119497800886D938F2B309CE788E9E18">181</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8C96A4FFE1B59841760653D8D9ACDF0F">journal article</mods:classification>
<mods:identifier id="5AF3135226F03D657CB6039F3FC02A02" type="CLB-Dataset">55701</mods:identifier>
<mods:identifier id="05D6083CE34BB870B66DC0FE9B999A01" type="DOI">10.11646/zootaxa.5133.2.1</mods:identifier>
<mods:identifier id="14517471E9BE784E344D4727856296E3" type="GBIF-Dataset">49bbe6cd-9052-42a3-8da7-1039e8c8de60</mods:identifier>
<mods:identifier id="0D431724038D3E485411CFB3AA2619BF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34F693CE65387146064AC5172BAE2C7A" type="Zenodo-Dep">6521560</mods:identifier>
<mods:identifier id="C5778699827C5FC4E43DBE14C9FFFD4D" type="ZooBank">6B6B0F37-8D98-4930-9038-EEE9F0E7B674</mods:identifier>
</mods:mods>
<treatment id="414987D12A17FFF8FF21FE2CFC1C7E38" ID-DOI="http://doi.org/10.5281/zenodo.6521131" ID-GBIF-Taxon="195192414" ID-Zenodo-Dep="6521131" LSID="urn:lsid:zoobank.org:act:F9365A97-E59B-4C1A-B924-14AE5EB1AB26" httpUri="http://treatment.plazi.org/id/414987D12A17FFF8FF21FE2CFC1C7E38" lastPageId="16" lastPageNumber="167" pageId="11" pageNumber="162">
<subSubSection id="81FA654C2A17FFE3FF21FE2CFFDF78A4" pageId="11" pageNumber="162" type="nomenclature">
<paragraph id="C95F36C72A17FFE3FF21FE2CFCF878D8" blockId="11.[151,643,402,501]" box="[151,420,402,429]" pageId="11" pageNumber="162">
<heading id="921781AB2A17FFE3FF21FE2CFCF878D8" bold="true" box="[151,420,402,429]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
<emphasis id="FB94EAD52A17FFE3FF21FE2CFCF878D8" bold="true" box="[151,420,402,429]" pageId="11" pageNumber="162">
<taxonomicName id="0EE04D442A17FFE3FF21FE2CFC0678D8" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[151,346,402,429]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A17FFE3FF21FE2CFC0678D8" bold="true" box="[151,346,402,429]" italics="true" pageId="11" pageNumber="162">Lebbeus subtilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A757AE2A17FFE3FED7FE2DFCF878D8" box="[353,420,403,429]" pageId="11" pageNumber="162" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C95F36C72A17FFE3FF21FE09FFDF78A4" blockId="11.[151,643,402,501]" box="[151,643,438,465]" pageId="11" pageNumber="162">[New Japanese name: Sakura-ibara-mo-ebi]</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A17FFE3FF21FE65FC4E7880" box="[151,274,474,501]" pageId="11" pageNumber="162" type="description">
<paragraph id="C95F36C72A17FFE3FF21FE65FC4E7880" blockId="11.[151,643,402,501]" box="[151,274,474,501]" pageId="11" pageNumber="162">
(
<figureCitation id="51DB2A422A17FFE3FF29FE65FC567880" box="[159,266,474,501]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="11.[151,250,1605,1629]" captionStartId-1="12.[151,250,1903,1927]" captionStartId-2="14.[151,250,1759,1783]" captionStartId-3="15.[151,250,1943,1967]" captionTargetBox-0="[151,1435,633,1581]" captionTargetBox-1="[151,1428,605,1875]" captionTargetBox-2="[247,1340,190,1727]" captionTargetBox-3="[206,1379,200,1919]" captionTargetId-0="figure-125@11.[151,1435,633,1581]" captionTargetId-1="figure-200@12.[151,1436,597,1879]" captionTargetId-2="figure-19@14.[238,1349,181,1735]" captionTargetId-3="figure-17@15.[186,1401,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURE 5. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698, habitus in lateral view, showing living colouration." captionText-1="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." captionText-2="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." captionText-3="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.6521572" figureDoi-1="http://doi.org/10.5281/zenodo.6521574" figureDoi-2="http://doi.org/10.5281/zenodo.6521580" figureDoi-3="http://doi.org/10.5281/zenodo.6521582" httpUri-0="https://zenodo.org/record/6521572/files/figure.png" httpUri-1="https://zenodo.org/record/6521574/files/figure.png" httpUri-2="https://zenodo.org/record/6521580/files/figure.png" httpUri-3="https://zenodo.org/record/6521582/files/figure.png" pageId="11" pageNumber="162">Figs. 5–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A17FFE3FF21FD9CF9037B14" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="C95F36C72A17FFE3FF21FD9CF9037B14" blockId="11.[151,1436,546,609]" pageId="11" pageNumber="162">
<emphasis id="FB94EAD52A17FFE3FF21FD9CFCDE7B49" bold="true" box="[151,386,546,572]" pageId="11" pageNumber="162">Material examined.</emphasis>
<materialsCitation id="79883C9A2A17FFE3FE3CFD9DF9037B14" ID-GBIF-Occurrence="3763099305" collectingDate="2015-09-09" collectionCode="CBM-ZC" collectorName="K. Matsuzaki" elevation="650" pageId="11" pageNumber="162" specimenCode="CBM-ZC 16698" specimenCount="1" typeStatus="holotype">
<typeStatus id="165B88652A17FFE3FE3CFD9DFCAA7B48" box="[394,502,547,573]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
. 
<specimenCode id="99469EBC2A17FFE3FDB7FD9DFFE17B48" box="[513,701,547,573]" collectionCode="CBM-ZC" pageId="11" pageNumber="162">CBM-ZC 16698</specimenCode>
, ovigerous female (cl 
<quantity id="0E189B222A17FFE3FC0CFD9CF97F7B48" box="[954,1059,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="11" pageNumber="162" unit="mm" value="15.9">15.9 mm</quantity>
), off Rausu, Nemuro Strait, 
<quantity id="0E189B222A17FFE3FAD2FD9CFDBC7B14" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="162" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<elevation id="42CDD1F42A17FFE3FAD2FD9CFDBC7B14" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="162" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500– 800 m</elevation>
</quantity>
, 
<date id="BD5E10072A17FFE3FF5AFDF9FCE77B14" box="[236,443,582,609]" pageId="11" pageNumber="162" value="2015-09-09">
<collectingDate id="AD1AE9EF2A17FFE3FF5AFDF9FCE77B14" box="[236,443,582,609]" pageId="11" pageNumber="162" value="2015-09-09">9 September 2015</collectingDate>
</date>
, 
<collectingMethod id="10A14ED02A17FFE3FE70FDF9FF887B14" box="[454,724,583,609]" pageId="11" pageNumber="162">commercial shrimp trap</collectingMethod>
, coll. 
<collectorName id="641553112A17FFE3FCA1FDF9FEED7B14" box="[791,945,583,609]" pageId="11" pageNumber="162">K. Matsuzaki</collectorName>
, DNA voucher.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="9D9F664F2A17FFE3FF21F9FBFC0F7FF4" ID-DOI="http://doi.org/10.5281/zenodo.6521572" ID-Zenodo-Dep="6521572" httpUri="https://zenodo.org/record/6521572/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1605,1629]" targetBox="[151,1435,633,1581]" targetPageId="11">
<paragraph id="C95F36C72A17FFE3FF21F9FBFC0F7FF4" blockId="11.[151,1436,1605,1665]" pageId="11" pageNumber="162">
<emphasis id="FB94EAD52A17FFE3FF21F9FBFC4E7F28" bold="true" box="[151,274,1605,1629]" pageId="11" pageNumber="162">FIGURE 5.</emphasis>
<taxonomicName id="0EE04D442A17FFE3FEAEF9FBFCE77F28" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[280,443,1605,1629]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A17FFE3FEAEF9FBFCE77F28" box="[280,443,1605,1629]" italics="true" pageId="11" pageNumber="162">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A17FFE3FE77F9FBFCA57F28" bold="true" box="[449,505,1605,1629]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="E0A757AE2A17FFE3FE77F9FBFCA57F28" box="[449,505,1605,1629]" pageId="11" pageNumber="162" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698, habitus in lateral view, showing living colouration.
</paragraph>
</caption>
<subSubSection id="81FA654C2A17FFE4FF71F911F9147880" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162" type="diagnosis">
<paragraph id="C95F36C72A17FFE4FF71F911F9147880" blockId="11.[151,1436,1711,2026]" lastBlockId="12.[151,1437,151,573]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
<emphasis id="FB94EAD52A17FFE3FF71F911FC1E7FBC" bold="true" box="[199,322,1711,1737]" pageId="11" pageNumber="162">Diagnosis.</emphasis>
Rostrum (
<figureCitation id="51DB2A422A17FFE3FE08F911FF457FBF" box="[446,537,1711,1738]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="11" pageNumber="162">Fig. 6A</figureCitation>
) somewhat curving dorsally, distinctly overreaching distal margin of antennal scaphocerite, 1.6 times as long as carapace; dorsal margin armed with 4 small teeth, including 2 on proximal 0.3 of rostrum and 2 postrostral (posteriormost tooth located at 0.3 of carapace length); ventral blade poorly developed, armed with 4 widely spaced teeth; tip simple, acuminate. Carapace (
<figureCitation id="51DB2A422A17FFE3FC31F8A5FE827E43" box="[903,990,1819,1846]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="11" pageNumber="162">Fig. 6A</figureCitation>
) with low postrostral ridge; supraorbital spine small, arising at level of posterior margin of orbit; no notch below supraorbital spine; pterygostomial spine small; anterolateral margin between antennal and pterygostomial spines gently sinuous. Pleon (
<figureCitation id="51DB2A422A17FFE3FB0CF8DDF8537E0B" box="[1210,1295,1891,1918]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="11" pageNumber="162">Fig. 6B</figureCitation>
) with pleura of anterior 3 pleomeres rounded, unarmed, that of pleomere 4 with angular posteroventral margin, that of pleomere 5 with strongly produced posteroventral spine.Telson (
<figureCitation id="51DB2A422A17FFE3FD6AF815FE137EB3" box="[732,847,1963,1990]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="11" pageNumber="162">Fig. 7C, D</figureCitation>
) armed with 6 (right) or 9 (left) dorsolateral spiniform setae on either side; posterior margin rounded, with 2 pairs of unequal spiniform setae and some short setae on very narrow space. Antennular peduncle (
<figureCitation id="51DB2A422A10FFE4FD85FF29FFC479C4" box="[563,664,151,177]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="12" pageNumber="163">Figs. 6A</figureCitation>
; 
<figureCitation id="51DB2A422A10FFE4FD12FF29FF9B79C4" box="[676,711,151,177]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="12" pageNumber="163">7A</figureCitation>
) slender, reaching distal 0.2 of scaphocerite; article 1 with small spine on dorsolateral distal angle; stylocerite falling short of distal margin of peduncular article 1, proximolateral lobe gently convex. Antenna (
<figureCitation id="51DB2A422A10FFE4FE5BFF61FF0F798C" box="[493,595,223,249]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="12" pageNumber="163">Figs. 6A</figureCitation>
; 
<figureCitation id="51DB2A422A10FFE4FDD6FF61FFF8798C" box="[608,676,223,249]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="12" pageNumber="163">7A, E</figureCitation>
) with scaphocerite 3.5 times as long as wide, distolateral spine far exceeded by distomesially produced, rounded lamella; carpocerite reaching midlength of scaphocerite. Maxilliped 3 (
<figureCitation id="51DB2A422A10FFE4FF00FE99FC537834" box="[182,271,295,321]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="12" pageNumber="163">Fig. 8A</figureCitation>
) overreaching distal margin of antennal scaphocerite by half-length of ultimate article; ultimate article bearing row of 4 cuspidate setae distomesially (
<figureCitation id="51DB2A422A10FFE4FD0DFEF5FE4E7810" box="[699,786,331,357]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="12" pageNumber="163">Fig. 7F</figureCitation>
). Pereopod 1 (
<figureCitation id="51DB2A422A10FFE4FC0CFEF5F94E7810" box="[954,1042,331,357]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="12" pageNumber="163">Fig. 8B</figureCitation>
) relatively slender, reaching distal 0.2 of scaphocerite. Pereopod 3 (
<figureCitation id="51DB2A422A10FFE4FDB1FED1FF0678FC" box="[519,602,367,393]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="12" pageNumber="163">Fig. 8F</figureCitation>
) relatively slender, overreaching antennal scale by 0.6 length of propodus; dactylus (
<figureCitation id="51DB2A422A10FFE4FEB0FE2DFC0A78D8" box="[262,342,403,429]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="12" pageNumber="163">Fig. 7J</figureCitation>
) 0.13 times as long as propodus, 2.6 times longer than wide, with 6 narrowly spaced accessory spiniform setae on entire flexor margin; merus armed with 10 robust spiniform setae on lateral surface. Strap-like epipods on maxilliped 3 and pereopods 1 and 2, no epipod on pereopod 3 (
<figureCitation id="51DB2A422A10FFE4FC6AFE65F9677880" box="[988,1083,475,501]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="12" pageNumber="163">Fig. 7M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A10FFF8FF71FE40F91F7A98" lastPageId="16" lastPageNumber="167" pageId="12" pageNumber="163" type="description">
<paragraph id="C95F36C72A10FFE4FF71FE40FDA37B48" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="163">
<emphasis id="FB94EAD52A10FFE4FF71FE40FC057B6D" bold="true" box="[199,345,510,536]" pageId="12" pageNumber="163">Description.</emphasis>
<emphasis id="FB94EAD52A10FFE4FEDEFE41FC937B6C" box="[360,463,511,537]" italics="true" pageId="12" pageNumber="163">
<typeStatus id="165B88652A10FFE4FEDEFE41FC937B6C" box="[360,463,511,537]" pageId="12" pageNumber="163" type="holotype">Holotype</typeStatus>
</emphasis>
. Body (
<figureCitation id="51DB2A422A10FFE4FD8DFE41FFCF7B6C" box="[571,659,510,537]" captionStart="FIGURE 5" captionStartId="11.[151,250,1605,1629]" captionTargetBox="[151,1435,633,1581]" captionTargetId="figure-125@11.[151,1435,633,1581]" captionTargetPageId="11" captionText="FIGURE 5. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698, habitus in lateral view, showing living colouration." figureDoi="http://doi.org/10.5281/zenodo.6521572" httpUri="https://zenodo.org/record/6521572/files/figure.png" pageId="12" pageNumber="163">Figs. 5</figureCitation>
, 
<figureCitation id="51DB2A422A10FFE4FD1EFE41FFA87B6C" box="[680,756,511,537]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="12" pageNumber="163">6A, B</figureCitation>
) moderately stout; integument moderately firm, surface glabrous.
</paragraph>
<caption id="9D9F664F2A10FFE4FF21F8D1FEBB7EDE" ID-DOI="http://doi.org/10.5281/zenodo.6521574" ID-Zenodo-Dep="6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1903,1927]" targetBox="[151,1428,605,1875]" targetPageId="12">
<paragraph id="C95F36C72A10FFE4FF21F8D1FEBB7EDE" blockId="12.[151,1436,1903,1963]" pageId="12" pageNumber="163">
<emphasis id="FB94EAD52A10FFE4FF21F8D1FC487EF2" bold="true" box="[151,276,1903,1927]" pageId="12" pageNumber="163">FIGURE 6.</emphasis>
<taxonomicName id="0EE04D442A10FFE4FEADF8D1FC9C7EF2" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[283,448,1903,1927]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A10FFE4FEADF8D1FC9C7EF2" box="[283,448,1903,1927]" italics="true" pageId="12" pageNumber="163">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A10FFE4FE71F8D1FF5D7EF2" bold="true" box="[455,513,1903,1927]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="E0A757AE2A10FFE4FE71F8D1FF5D7EF2" box="[455,513,1903,1927]" pageId="12" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view.
</paragraph>
</caption>
<paragraph id="C95F36C72A11FFE5FF71FF29F9517B6C" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Rostrum (
<figureCitation id="51DB2A422A11FFE5FE8EFF29FCCD79C7" box="[312,401,151,178]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Fig. 6A</figureCitation>
) somewhat curving dorsally, distinctly overreaching distal margin of antennal scaphocerite, 1.6 times as long as carapace; dorsal margin armed with 4 small teeth including 2 on rostrum proper (anteriormost tooth located at proximal 0.3 of rostral length, and then anterior 0.7 leaving unarmed) and 2 postrostral (posteriormost tooth located at 0.3 of carapace length); ventral blade very narrow, deepest at 0.3 length, margin armed with 4 subequal teeth more widely spaced anteriorly; lateral carina obsolete. Carapace (
<figureCitation id="51DB2A422A11FFE5FBF3FE99F9FD7837" box="[1093,1185,295,322]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Fig. 6A</figureCitation>
) with low postrostral ridge extending beyond midlength of carapace length, becoming broader and blunt posteriorly; dorsal profile in lateral view nearly straight; orbital margin evenly concave, ventral part limbed marginally (
<figureCitation id="51DB2A422A11FFE5FB05FED1F85178FC" box="[1203,1293,367,393]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7B</figureCitation>
), postorbital region shallowly depressed; supraorbital spine small, arising at level of posterior margin of orbit; no notch below supraorbital spine; suborbital lobe distinct, rounded (
<figureCitation id="51DB2A422A11FFE5FCB8FE09FE3078A4" box="[782,876,439,465]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7B</figureCitation>
); antennal spine small, well exceeding beyond suborbital lobe; pterygostomial spine small, directed anteroventrally; anterolateral margin between antennal and pterygostomial spines gently sinuous; no conspicuous ridges on lateral surface.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FD9DF9EC7A60" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Pleon (
<figureCitation id="51DB2A422A11FFE5FEACFD9DFC287B4B" box="[282,372,547,574]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Fig. 6B</figureCitation>
) with pleura of anterior four pleomeres rounded, unarmed, that of pleomere 5 with strong posteroventral spine. Pleomere 2 tergum with deep transverse groove. Pleomere 3 rounded dorsally, posterodorsal margin somewhat produced posteriorly. Pleomere 6 1.7 times as long as pleomere 5, 2.4 times as long as high, with small posteroventral spine and posterolateral process terminating in small spine. Telson (
<figureCitation id="51DB2A422A11FFE5FB02FD31F8707BDC" box="[1204,1324,655,681]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7C, D</figureCitation>
) reaching slightly beyond posterior margin of uropods, 1.5 times as long as pleomere 6, relatively narrow, armed with 6 (right) or 9 (left) dorsolateral spiniform setae; posterior margin triangular, with 2 pairs of unequal spiniform setae (mesial pair longer and stouter than lateral pair) and several short setae on very narrow median space.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FCA1FF247A28" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Eye (
<figureCitation id="51DB2A422A11FFE5FEB4FCA1FC347A4C" box="[258,360,799,825]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Figs. 6A</figureCitation>
; 
<figureCitation id="51DB2A422A11FFE5FEC2FCA1FCCB7A4C" box="[372,407,799,825]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">7A</figureCitation>
) subpyriform; cornea darkly pigmented, its maximum width about 0.24 of carapace length; ocellus absent. Eyestalk short, cup-shaped.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FCD9FE2C7D2C" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Antennular peduncle (
<figureCitation id="51DB2A422A11FFE5FE7FFCD9FF6D7AF4" box="[457,561,871,897]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Figs. 6A</figureCitation>
; 
<figureCitation id="51DB2A422A11FFE5FDF7FCD9FF387AF4" box="[577,612,871,897]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">7A</figureCitation>
) slender, reaching distal 0.2 of antennal scaphocerite. Article 1 longer than distal two articles combined, with small spine on dorsolateral distal angle; stylocerite moderately slender, partially in touch with first article mesially, slightly curving dorsally, terminating in acute spine falling far short of distal margin of peduncular article 1, proximolateral lobe with small tubercle, making proximolateral lobe as hooklike projection. Article 2 subcylindrical, armed with small spine at dorsolateral distal angle. Article 3 short, with small spine on dorsodistal margin. Upper flagellum with thickened aesthetasc-bearing portion 0.4 times as long as carapace; lower flagellum distinctly longer than upper flagellum.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FBDDFF757D9C" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Antenna (
<figureCitation id="51DB2A422A11FFE5FE81FBDDFCC07D08" box="[311,412,1123,1149]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="13" pageNumber="164">Figs. 6A</figureCitation>
; 
<figureCitation id="51DB2A422A11FFE5FE1FFBDDFCB17D08" box="[425,493,1123,1149]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">7A, E</figureCitation>
) with moderately stout basicerite, bearing blunt dorsolateral projection and slender ventrolateral spine. Scaphocerite 0.9 times as long as carapace, 3.5 times as long as wide; lateral margin nearly straight; distolateral spine relatively slender, far exceeded by distomesially produced, rounded lamella. Carpocerite reaching midlength of scaphocerite.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FB4DF9B97CE8" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Mouthparts not dissected. Maxilliped 3 (
<figureCitation id="51DB2A422A11FFE5FD28FB4DFFA57C78" box="[670,761,1267,1293]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="13" pageNumber="164">Fig. 8A</figureCitation>
) relatively slender, overreaching distal margin of antennal scaphocerite by 0.5 length of ultimate article; ultimate article 3.7 times as long as penultimate article, bearing 8, including 4 on mesial margin, darkly pigmented cuspidate setae partially circumscribing terminal margin (
<figureCitation id="51DB2A422A11FFE5FA8CFA85F8D27C20" box="[1338,1422,1339,1365]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7F</figureCitation>
); antepenultimate article subequal in length to ultimate article, with 1 minute spiniform seta at ventrolateral distal angle, dorsodistal margin unarmed, distolateral margin with 1 tiny spine (
<figureCitation id="51DB2A422A11FFE5FC7BFA3DF97B7CE8" box="[973,1063,1411,1437]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7G</figureCitation>
); exopod absent.
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71FA19F91C7FC8" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Pereopod 1 (
<figureCitation id="51DB2A422A11FFE5FEEAFA19FC867CB4" box="[348,474,1447,1473]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="13" pageNumber="164">Fig. 8B; C</figureCitation>
) relatively slender, reaching midlength of scaphocerite. Chela 1.3 times as long as carpus, 5.7 times as long as wide; dactylus 0.6 times as long as palm, terminating in 2 black corneous claws and slender, accessory claw at base of shorter terminal claw (
<figureCitation id="51DB2A422A11FFE5FC9FFA51FED97F7C" box="[809,901,1519,1545]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7H</figureCitation>
); fixed finger terminating in single black claw (
<figureCitation id="51DB2A422A11FFE5FF16F9ADFDA07F58" box="[160,252,1555,1581]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7H</figureCitation>
); palm 3.6 times as long as wide, with short row of grooming setae on mesial face proximally. Carpus subcylindrical; grooming apparatus consisting of rows of stiff setae circumscribing deep depression on subdistal portion on dorsal face and subdistal cluster of short stiff setae on mesial face. Merus 7.7 times as long as wide, with minute spine on near dorsal margin proximally and few minute spiniform setae on ventral margin proximally (
<figureCitation id="51DB2A422A11FFE5FAD9F9C1FDEE7FC8" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7I</figureCitation>
). Ischium with cluster of minute spiniform setae on ventrodistal margin (
<figureCitation id="51DB2A422A11FFE5FC52F91DF96F7FC8" box="[996,1075,1699,1725]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7I</figureCitation>
).
</paragraph>
<paragraph id="C95F36C72A11FFE5FF71F979FE6A7E38" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="164">
Pereopod 2 (
<figureCitation id="51DB2A422A11FFE5FEE3F979FCF17F94" box="[341,429,1735,1761]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="13" pageNumber="164">Fig. 8D</figureCitation>
) moderately slender, overreaching distal margin of scaphocerite by length of chela. Chela slightly less than 0.2 times as long as carpus, with tufts of short setae on fingers; dactylus 0.4 times as long as palm. Carpus subdivided into 7 segments, length ratio of segments: 1: 0.6: 2.7: 0.8: 0.6: 0.4: 0.8; distalmost segment with prominent tuft of setae. Merus 1.2 times as long as ischium.
</paragraph>
<paragraph id="C95F36C72A11FFF8FF71F8E9FF1A78FC" blockId="13.[151,1437,151,2013]" lastBlockId="16.[151,1437,151,1869]" lastPageId="16" lastPageNumber="167" pageId="13" pageNumber="164">
Pereopod 3–5 (
<figureCitation id="51DB2A422A11FFE5FEC1F8E9FCAC7E04" box="[375,496,1879,1905]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="13" pageNumber="164">Fig. 8E–G</figureCitation>
) relatively slender. Pereopod 3 (
<figureCitation id="51DB2A422A11FFE5FCDFF8E9FE9C7E04" box="[873,960,1879,1905]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="13" pageNumber="164">Fig. 8E</figureCitation>
) overreaching scaphocerite by 0.6 length of propodus; dactylus (
<figureCitation id="51DB2A422A11FFE5FE29F8C5FCB37EE0" box="[415,495,1915,1941]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="13" pageNumber="164">Fig. 7J</figureCitation>
) short, 0.13 times as long as propodus, 2.6 times longer than wide, terminating in moderately slender, clearly demarcated unguis, with 6 accessory spiniform setae on entire flexor margin, increasing in length distally, distalmost accessory seta slightly curved; propodus 20 times longer than wide, with evenly spaced minute spiniform setae arranged in 2 rows on flexor margin; carpus 0.6 times as long as propodus, unarmed: merus 13 times longer than wide, armed with 10 robust spiniform setae on lateral surface; ischium unarmed. Pereopod 4 (
<figureCitation id="51DB2A422A0CFFF8FF16FF61FDA8798C" box="[160,244,223,249]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="16" pageNumber="167">Fig. 8F</figureCitation>
) similar to pereopod 3, overreaching antennal scale by 0.2 length of propodus; dactylus (
<figureCitation id="51DB2A422A0CFFF8FB5DFF61F818798C" box="[1259,1348,223,249]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7K</figureCitation>
) with 6 accessory spiniform setae; merus with 8 robust spiniform setae on lateral surface. Pereopod 5 (
<figureCitation id="51DB2A422A0CFFF8FB7DFEBDF87F7868" box="[1227,1315,259,285]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[206,1379,200,1919]" captionTargetId="figure-17@15.[186,1401,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="16" pageNumber="167">Fig. 8G</figureCitation>
) generally similar to pereopods 3 and 4, overreaching antennal scale by length of dactylus; dactylus (
<figureCitation id="51DB2A422A0CFFF8FB36FE99F9897834" box="[1152,1237,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7L</figureCitation>
) with 6 accessory spiniform setae; propodus with cluster of grooming setae on flexor surface in distal 0.1 of flexor margin; merus with 4 robust spiniform seta on lateral face.
</paragraph>
<caption id="9D9F664F2A12FFE6FF21F961FF5E7E86" ID-DOI="http://doi.org/10.5281/zenodo.6521580" ID-Zenodo-Dep="6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1759,1783]" targetBox="[247,1340,190,1727]" targetPageId="14">
<paragraph id="C95F36C72A12FFE6FF21F961FF5E7E86" blockId="14.[151,1437,1759,2035]" pageId="14" pageNumber="165">
<emphasis id="FB94EAD52A12FFE6FF21F961FC4E7F82" bold="true" box="[151,274,1759,1783]" pageId="14" pageNumber="165">FIGURE 7.</emphasis>
<taxonomicName id="0EE04D442A12FFE6FEAEF961FCE77F82" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[280,443,1759,1783]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A12FFE6FEAEF961FCE77F82" box="[280,443,1759,1783]" italics="true" pageId="14" pageNumber="165">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A12FFE6FE77F961FCA67F82" bold="true" box="[449,506,1759,1783]" pageId="14" pageNumber="165">
<taxonomicNameLabel id="E0A757AE2A12FFE6FE77F961FCA67F82" box="[449,506,1759,1783]" pageId="14" pageNumber="165" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod.
</paragraph>
</caption>
<caption id="9D9F664F2A13FFE7FF21F829F9087E8D" ID-DOI="http://doi.org/10.5281/zenodo.6521582" ID-Zenodo-Dep="6521582" httpUri="https://zenodo.org/record/6521582/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1943,1967]" targetBox="[206,1379,200,1919]" targetPageId="15">
<paragraph id="C95F36C72A13FFE7FF21F829F9087E8D" blockId="15.[151,1436,1943,2040]" pageId="15" pageNumber="166">
<emphasis id="FB94EAD52A13FFE7FF21F829FC497EDA" bold="true" box="[151,277,1943,1967]" pageId="15" pageNumber="166">FIGURE 8.</emphasis>
<taxonomicName id="0EE04D442A13FFE7FEABF826FC9F7EC5" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[285,451,1944,1968]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A13FFE7FEABF826FC9F7EC5" box="[285,451,1944,1968]" italics="true" pageId="15" pageNumber="166">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A13FFE7FE7AF826FF5B7EC5" bold="true" box="[460,519,1944,1968]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="E0A757AE2A13FFE7FE7AF826FF5B7EC5" box="[460,519,1944,1968]" pageId="15" pageNumber="166" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, left maxilliped 3, lateral view; B, right pereopod 1, lateral view; C, same, chela and carpus, dorsal (extensor) view; D, right pereopod 2, lateral view; E, right pereopod 3, lateral view; F, left pereopod 4, lateral view; G, left pereopod 5, lateral view.
</paragraph>
</caption>
<paragraph id="C95F36C72A0CFFF8FF71FE2DFF6F7880" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
Strap-like epipods present on maxilliped 3 and pereopods 1 and 2; anterior 2 epipods terminally hooked, interacting with setobranchs on pereopods 1 and 2, respectively; epipod on pereopod 2 simple, no corresponding setobranch on pereopod 3 (
<figureCitation id="51DB2A422A0CFFF8FE71FE65FF7B7880" box="[455,551,475,501]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7M</figureCitation>
).
</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71FE41FF317B6C" blockId="16.[151,1437,151,1869]" box="[199,621,511,537]" pageId="16" pageNumber="167">Pleopods without distinctive features.</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71FD9DFCBF7BF0" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
Uropod (
<figureCitation id="51DB2A422A0CFFF8FE85FD9DFCC07B48" box="[307,412,547,573]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="16" pageNumber="167">Figs. 6B</figureCitation>
; 
<figureCitation id="51DB2A422A0CFFF8FE19FD9DFC8D7B48" box="[431,465,547,573]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">7C</figureCitation>
) reaching posterior margin of telson. Protopod terminating in strong acute spine posterolaterally. Exopod subequal in length of endopod, with small posterolateral spine exceeded by tiny spiniform seta arising at its mesial base.
</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71FD31FF467BDC" blockId="16.[151,1437,151,1869]" box="[199,538,654,681]" pageId="16" pageNumber="167">
Eggs oval, large, 2.4× 
<quantity id="0E189B222A0CFFF8FE0BFD30FF4A7BDC" box="[445,534,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="167" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71FD0CF91F7A98" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
<emphasis id="FB94EAD52A0CFFF8FF71FD0CFCF27BB9" bold="true" box="[199,430,690,716]" pageId="16" pageNumber="167">Colouration in life.</emphasis>
Carapace and rostrum pale pink, tip of rostrum reddish; pleon also pale pink generally, pleomeres 1 to 3 each with irregular red transverse band medially, pleomeres 4 and 5 each with red transverse band posteriorly; pleomere 6 with red tint posteriorly; tail fan (telson and uropods) reddish; cornea black; antennular and antennal peduncles generally pinkish, antennal carpocerite red; flagella uniformly pink or reddish; maxilliped 3 antepenultimate and penultimate articles reddish proximally and distally, otherwise white, penultimate article red, with white blotch; pereopod 1 pink to red, fingers red; pereopod 2 ischium and merus mottled with pink and red; carpus and chela pale pink; pereopods 3–5 ischia and meri mottled pink and red; carpi and propodi generally white, but latter articles with tint of red distally; dactyli red; pleopods 2–5 generally pale pink, ventral expansions of protopods each with proximal red and distal white blotches; eggs yellow (
<figureCitation id="51DB2A422A0CFFF8FC47FC6DF9697A98" box="[1009,1077,978,1005]" captionStart="FIGURE 5" captionStartId="11.[151,250,1605,1629]" captionTargetBox="[151,1435,633,1581]" captionTargetId="figure-125@11.[151,1435,633,1581]" captionTargetPageId="11" captionText="FIGURE 5. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698, habitus in lateral view, showing living colouration." figureDoi="http://doi.org/10.5281/zenodo.6521572" httpUri="https://zenodo.org/record/6521572/files/figure.png" pageId="16" pageNumber="167">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A0CFFF8FF71FC48F9917D64" box="[199,1229,1014,1041]" pageId="16" pageNumber="167" type="distribution">
<paragraph id="C95F36C72A0CFFF8FF71FC48F9917D64" blockId="16.[151,1437,151,1869]" box="[199,1229,1014,1041]" pageId="16" pageNumber="167">
<emphasis id="FB94EAD52A0CFFF8FF71FC48FC3D7D65" bold="true" box="[199,353,1014,1040]" pageId="16" pageNumber="167">Distribution.</emphasis>
Presently known only from off Rausu, Nemuro Strait, at depth of 
<quantity id="0E189B222A0CFFF8FBF1FC48F9957D64" box="[1095,1225,1014,1041]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="16" pageNumber="167" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
500– 
<specimenCount id="DFE6FD4E2A0CFFF8FB36FC48F9957D64" box="[1152,1225,1014,1041]" pageId="16" pageNumber="167" type="generic">800 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A0CFFF8FF71FBA5FC587E70" pageId="16" pageNumber="167" type="discussion">
<paragraph id="C95F36C72A0CFFF8FF71FBA5FE9D7C78" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
<emphasis id="FB94EAD52A0CFFF8FF71FBA5FC677D40" bold="true" box="[199,315,1051,1077]" pageId="16" pageNumber="167">Remarks.</emphasis>
<taxonomicName id="0EE04D442A0CFFF8FEF0FBA5FF5C7D40" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[326,512,1051,1077]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FEF0FBA5FF5C7D40" box="[326,512,1051,1077]" italics="true" pageId="16" pageNumber="167">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FDBDFBA5FF127D40" bold="true" box="[523,590,1051,1077]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FDBDFBA5FF127D40" box="[523,590,1051,1077]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is referred to the species group characterised by having epipods on the maxilliped 3 and pereopods 1 and 2, represented by 23 previously described species (cf. 
<tableCitation id="8462037C2A0CFFF8FBCFFB81F9977D2C" box="[1145,1227,1087,1113]" captionStart="TABLE 1" captionStartId="1.[149,237,469,495]" captionTargetPageId="1" captionText="TABLE 1. List of species of Lebbeus White, 1847, grouped according to the development of epipods on the pereopods 1–3. *: species recorded from the Sea of Okhotsk." httpUri="http://table.plazi.org/id/9D9F664F2A1DFFE9FF23FE6BFAFB789A" pageId="16" pageNumber="167" tableUuid="9D9F664F2A1DFFE9FF23FE6BFAFB789A">Table 1</tableCitation>
). The new species is characteristic in having an elongate rostrum being distinctly longer than the carapace, with a poorly developed ventral lamina and a greatly unarmed dorsal margin (the distal 0.6 is unarmed). None of the other species in this group has such an elongate rostrum with few dorsal teeth and a poorly developed ventral lamina (cf. 
<bibRefCitation id="AD714B362A0CFFF8FA97FB15FD8E7D9C" author="Kobjakova, Z. I." pageId="16" pageNumber="167" pagination="93 - 154" refId="ref15778" refString="Kobjakova, Z. I. (1937) Systematisch Ubersicht der Dekapoden aus dem Ochotskischen und Japanischen Meere. Uchenie Zapiski Leningrad Universitet, 15, 93 - 154, pls. 1 - 3. [in Russian with German summary]" type="journal article" year="1937">Kobjakova 1937</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FF68FB71FCD77D9C" author="Zarenkov, N. A." box="[222,395,1231,1257]" pageId="16" pageNumber="167" pagination="343 - 350" refId="ref18253" refString="Zarenkov, N. A. (1960) Observations about some decapod Crustacea from the Okhotsk and Bering Seas. Trudy Instituta Okeanologii, 34, 343 - 350. [in Russian]" type="journal article" year="1960">Zarenkov 1960</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FE20FB71FFE27D9C" author="Wicksten, M. K. &amp; Mendez, M. G." box="[406,702,1231,1257]" pageId="16" pageNumber="167" pagination="106 - 120" refId="ref18202" refString="Wicksten, M. K. &amp; Mendez, M. G. (1982) New records and new species of the genus Lebbeus (Caridea: Hippolytidae) in the eastern Pacific Ocean. Bulletin of the Southern California Academy of Sciences, 81, 106 - 120." type="journal article" year="1982">Wicksten &amp; Méndez 1982</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FD7CFB71FEFA7D9C" author="Kensley, B. &amp; Tranter, H. A. &amp; Griffin, D. J. G." box="[714,934,1231,1257]" pageId="16" pageNumber="167" pagination="263 - 331" refId="ref15620" refString="Kensley, B., Tranter, H. A. &amp; Griffin, D. J. G. (1987) Deepwater decapod Crustacea from eastern Australia (Penaeidea and Caridea). Records of the Australian Museum, 39 (5), 263 - 331. https: // doi. org / 10.3853 / j. 0067 - 1975.39.1987.171" type="journal article" year="1987">
Kensley 
<emphasis id="FB94EAD52A0CFFF8FC9BFB71FE027D9C" box="[813,862,1231,1257]" italics="true" pageId="16" pageNumber="167">et al</emphasis>
. 1987
</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FC04FB71F90C7D9C" author="Hayashi, K." box="[946,1104,1231,1257]" pageId="16" pageNumber="167" pagination="107 - 138" refId="ref15377" refString="Hayashi, K. (1992 a) Studies on the hippolytid shrimps from Japan - VIII. The genus Lebbeus White. Journal of the Shimonoseki University of Fisheries, 40, 107 - 138." type="journal article" year="1992">Hayashi 1992</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FBEDFB71F9A27D9C" author="Crosnier, A." box="[1115,1278,1231,1257]" pageId="16" pageNumber="167" pagination="453 - 460" refId="ref15056" refString="Crosnier, A. (1999) Une nouvelle espece de Lebbeus d'Indonesie (Crustacea, Decapoda, Caridea, Hippolytidae). Zoosystema, 21, 453 - 460." type="journal article" year="1999">Crosnier 1999</bibRefCitation>
; Komai 2002; 
<bibRefCitation id="AD714B362A0CFFF8FF21FB4DFC797C78" author="Jensen, G. C." box="[151,293,1267,1293]" pageId="16" pageNumber="167" pagination="23 - 43" refId="ref15571" refString="Jensen, G. C. (2006) Three new species of Lebbeus (Crustacea: Decapoda: Hippolytidae) from the Northeastern Pacific. Zootaxa, 1383 (1), 23 - 43. https: // doi. org / 10.11646 / zootaxa. 1383.1.2" type="journal article" year="2006">Jensen 2006</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FE86FB4DFF0C7C78" author="McCallum, A. W. &amp; Poore, G. C. B." box="[304,592,1267,1293]" pageId="16" pageNumber="167" pagination="126 - 137" refId="ref16975" refString="McCallum, A. W. &amp; Poore, G. C. B. (2010) Two crested and colourful new species of Lebbeus (Crustacea: Caridea: Hippolytidae) from the continental margin of Western Australia. Zootaxa, 2372 (1), 126 - 137. https: // doi. org / 10.11646 / zootaxa. 2372.1.13" type="journal article" year="2010">McCallum &amp; Poore 2010</bibRefCitation>
; 
<bibRefCitation id="AD714B362A0CFFF8FDECFB4DFE7B7C78" author="Komai, T. &amp; Tsuchida, S. &amp; Segonzac, M." box="[602,807,1267,1293]" pageId="16" pageNumber="167" pagination="35 - 63" refId="ref16588" refString="Komai, T., Tsuchida, S. &amp; Segonzac, M. (2012) Records of species of the hippolytid genus Lebbeus White, 1847 (Crustacea: Decapoda: Caridea) from hydrothermal vents in the Pacific Ocean, with descriptions of three new species. Zootaxa, 3241 (1), 35 - 63. https: // doi. org / 10.11646 / zootaxa. 3241.1.2" type="journal article" year="2012">
Komai 
<emphasis id="FB94EAD52A0CFFF8FD1BFB4DFF837C78" box="[685,735,1267,1293]" italics="true" pageId="16" pageNumber="167">et al</emphasis>
. 2012
</bibRefCitation>
, 
<bibRefCitation id="AD714B362A0CFFF8FC84FB4DFE317C78" author="Komai, T. &amp; Matsuzaki, K. &amp; Hibino, M." box="[818,877,1267,1293]" pageId="16" pageNumber="167" pagination="390 - 400" refId="ref16448" refString="Komai, T., Matsuzaki, K. &amp; Hibino, M. (2016) Rediscovery and redescription of a deep-sea shrimp Lebbeus longidactylus (Kobjakova, 1936) (Crustacea: Decapoda: Caridea: Thoridae) based on material from the Nemuro Strait, southwestern Sea of Okhotsk. Zootaxa, 4175 (4), 390 - 400. https: // doi. org / 10.11646 / zootaxa. 4175.4.8" type="journal article" year="2016">2016</bibRefCitation>
, 
<bibRefCitation id="AD714B362A0CFFF8FCCEFB4DFEE87C78" author="Komai, T. &amp; Chan, T. - Y. &amp; Chang, S. - C." box="[888,948,1267,1293]" pageId="16" pageNumber="167" pagination="1 - 13" refId="ref16038" refString="Komai, T., Chan, T. - Y. &amp; Chang, S. - C. (2021) Redescription of the colorful deep-sea shrimp Lebbeus compressus Holthuis, 1947 and description of a closely allied new species from Taiwan (Decapoda: Caridea: Thoridae). Journal of Crustacean Biology, 41 (3): 1 - 13. https: // doi. org / 10.1093 / jcbiol / ruab 048" type="journal article" year="2021">2021</bibRefCitation>
).
</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71FAA9FC5E7FEC" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
Morphologically, 
<taxonomicName id="0EE04D442A0CFFF8FE24FAA9FF167C44" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[402,586,1303,1329]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FE24FAA9FF167C44" box="[402,586,1303,1329]" italics="true" pageId="16" pageNumber="167">Lebbeus subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FDE2FAA9FFCA7C44" bold="true" box="[596,662,1303,1329]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FDE2FAA9FFCA7C44" box="[596,662,1303,1329]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is rather similar to 
<taxonomicName id="0EE04D442A0CFFF8FCC8FAA9F9587C44" box="[894,1028,1303,1329]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="fujimotoi">
<emphasis id="FB94EAD52A0CFFF8FCC8FAA9F9587C44" box="[894,1028,1303,1329]" italics="true" pageId="16" pageNumber="167">L. fujimotoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="0EE04D442A0CFFF8FBF6FAA9F8597C44" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[1088,1285,1303,1329]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="rufomaculosus" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FBF6FAA9F8597C44" box="[1088,1285,1303,1329]" italics="true" pageId="16" pageNumber="167">L. rufomaculosus</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FAB9FAA9F80C7C44" bold="true" box="[1295,1360,1303,1329]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FAB9FAA9F80C7C44" box="[1295,1360,1303,1329]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the general body form and the shape and armature of the rostrum, although the development of pereopodal epipods is different among those species. The present new species differs from 
<taxonomicName id="0EE04D442A0CFFF8FC12FAE1F9747C0C" box="[932,1064,1375,1401]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="fujimotoi">
<emphasis id="FB94EAD52A0CFFF8FC12FAE1F9747C0C" box="[932,1064,1375,1401]" italics="true" pageId="16" pageNumber="167">L. fujimotoi</emphasis>
</taxonomicName>
in having fewer ventral teeth on the rostrum (four versus seven to nine), the antennular stylocerite falling short of the distal margin of the peduncular article 1 (versus reaching it) (cf. 
<figureCitation id="51DB2A422A0CFFF8FDBCFA19FF037CB4" box="[522,607,1447,1473]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7A</figureCitation>
versus 
<bibRefCitation id="AD714B362A0CFFF8FD00FA19FEEC7CB4" author="Matsuzaki, K. &amp; Hibino, M. &amp; Komai, T." box="[694,944,1446,1473]" pageId="16" pageNumber="167" pagination="309 - 318" refId="ref16905" refString="Matsuzaki, K., Hibino, M. &amp; Komai, T. (2015) A new species of the caridean shrimp genus Lebbeus White, 1847 (Crustacea: Decapoda: Thoridae) from the southwestern Sea of Okhotsk, Hokkaido, Japan. Zootaxa, 4032 (3), 309 - 318. https: // doi. org / 10.11646 / zootaxa. 4032.3.6" type="journal article" year="2015">
Matsuzaki 
<emphasis id="FB94EAD52A0CFFF8FC85FA19FE397CB4" box="[819,869,1447,1473]" italics="true" pageId="16" pageNumber="167">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 3A, B) and the presence of epipods on the pereopods 1 and 2 (versus epipods are usually absent on those pereopods) (cf. 
<figureCitation id="51DB2A422A0CFFF8FB80FA75F9CD7C90" box="[1078,1169,1483,1509]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7M</figureCitation>
versus 
<bibRefCitation id="AD714B362A0CFFF8FB51FA75FD8F7F7C" author="Matsuzaki, K. &amp; Hibino, M. &amp; Komai, T." pageId="16" pageNumber="167" pagination="309 - 318" refId="ref16905" refString="Matsuzaki, K., Hibino, M. &amp; Komai, T. (2015) A new species of the caridean shrimp genus Lebbeus White, 1847 (Crustacea: Decapoda: Thoridae) from the southwestern Sea of Okhotsk, Hokkaido, Japan. Zootaxa, 4032 (3), 309 - 318. https: // doi. org / 10.11646 / zootaxa. 4032.3.6" type="journal article" year="2015">
Matsuzaki 
<emphasis id="FB94EAD52A0CFFF8FAD2FA75F8C97C90" box="[1380,1429,1483,1509]" italics="true" pageId="16" pageNumber="167">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 3F). The absence of a prominent proximolateral projection on the antennular stylocerite immediately distinguishes 
<taxonomicName id="0EE04D442A0CFFF8FE85F9ADFCF87F58" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[307,420,1555,1581]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FE85F9ADFCF87F58" box="[307,420,1555,1581]" italics="true" pageId="16" pageNumber="167">L. subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FE18F9ADFCB37F58" bold="true" box="[430,495,1555,1581]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FE18F9ADFCB37F58" box="[430,495,1555,1581]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from 
<taxonomicName id="0EE04D442A0CFFF8FD8FF9ADFE197F58" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[569,837,1555,1581]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="rufomaculosus" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FD8FF9ADFE197F58" box="[569,837,1555,1581]" italics="true" pageId="16" pageNumber="167">Lebbeus rufomaculosus</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FCF8F9ADFED37F58" bold="true" box="[846,911,1555,1581]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FCF8F9ADFED37F58" box="[846,911,1555,1581]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(cf. 
<figureCitation id="51DB2A422A0CFFF8FC7FF9ADF9437F58" box="[969,1055,1555,1581]" captionStart="FIGURE 6" captionStartId="12.[151,250,1903,1927]" captionTargetBox="[151,1428,605,1875]" captionTargetId="figure-200@12.[151,1436,597,1879]" captionTargetPageId="12" captionText="FIGURE 6. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698. A, carapace and left cephalic appendages, lateral view; B, pleon, telson and left pleonal appendages, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6521574" httpUri="https://zenodo.org/record/6521574/files/figure.png" pageId="16" pageNumber="167">Fig. 6A</figureCitation>
versus 
<figureCitation id="51DB2A422A0CFFF8FBCCF9ADF98F7F58" box="[1146,1235,1555,1581]" captionStart="FIGURE 3" captionStartId="8.[151,250,1749,1773]" captionTargetBox="[268,1319,189,1714]" captionTargetId="figure-19@8.[259,1328,181,1725]" captionTargetPageId="8" captionText="FIGURE 3. Lebbeus rufomaculosus n. sp., holotype, ovigerous female (cl 16.5 mm), CBM-ZC 16695. A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, proximolateral portion of left antennular stylocerite, lateral view; F, left antenna, ventral view (setae and flagellum omitted); G, chela and carpus of right pereopod 1, extensor view; H, tips of fingers of right pereopod 1 (setae omitted); I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of right pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521566" httpUri="https://zenodo.org/record/6521566/files/figure.png" pageId="16" pageNumber="167">Fig. 3E</figureCitation>
). Furthermore, 
<taxonomicName id="0EE04D442A0CFFF8FA30F9ADFDB47F24" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FA30F9ADFDB47F24" italics="true" pageId="16" pageNumber="167">L. subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FF59F989FC727F24" bold="true" box="[239,302,1591,1617]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FF59F989FC727F24" box="[239,302,1591,1617]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="0EE04D442A0CFFF8FE74F989FFD87F24" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[450,644,1591,1617]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="rufomaculosus" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FE74F989FFD87F24" box="[450,644,1591,1617]" italics="true" pageId="16" pageNumber="167">L. rufomaculosus</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FD3DF989FF967F24" bold="true" box="[651,714,1591,1617]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FD3DF989FF967F24" box="[651,714,1591,1617]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in the more stout dactyli of the pereopods 3–5 (
<figureCitation id="51DB2A422A0CFFF8FB54F989F8127F24" box="[1250,1358,1591,1617]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7J–L</figureCitation>
versus 
<figureCitation id="51DB2A422A0CFFF8FF21F9E5FC587F00" box="[151,260,1627,1653]" captionStart="FIGURE 3" captionStartId="8.[151,250,1749,1773]" captionTargetBox="[268,1319,189,1714]" captionTargetId="figure-19@8.[259,1328,181,1725]" captionTargetPageId="8" captionText="FIGURE 3. Lebbeus rufomaculosus n. sp., holotype, ovigerous female (cl 16.5 mm), CBM-ZC 16695. A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, proximolateral portion of left antennular stylocerite, lateral view; F, left antenna, ventral view (setae and flagellum omitted); G, chela and carpus of right pereopod 1, extensor view; H, tips of fingers of right pereopod 1 (setae omitted); I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of right pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521566" httpUri="https://zenodo.org/record/6521566/files/figure.png" pageId="16" pageNumber="167">Fig. 3J–L</figureCitation>
) and the presence of an epipod on the pereopod 2 (versus epipod absent on pereopod 2) (
<figureCitation id="51DB2A422A0CFFF8FB44F9E5F8117F00" box="[1266,1357,1627,1653]" captionStart="FIGURE 7" captionStartId="14.[151,250,1759,1783]" captionTargetBox="[247,1340,190,1727]" captionTargetId="figure-19@14.[238,1349,181,1735]" captionTargetPageId="14" captionText="FIGURE 7. Lebbeus subtilis n. sp., holotype, ovigerous female (cl 15.9 mm), CBM-ZC 16698.A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, left antenna, ventral view (setae and flagellum omitted); F, distal part of ultimate article of left maxilliped 3, dorsal (extensor) view; G, distal part of antepenultimate article of left maxilliped 3, lateral view; H, terminal claws of fingers of right pereopod 1 chela; I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521580" httpUri="https://zenodo.org/record/6521580/files/figure.png" pageId="16" pageNumber="167">Fig. 7M</figureCitation>
versus 
<figureCitation id="51DB2A422A0CFFF8FF21F9C1FDA97FEC" box="[151,245,1663,1689]" captionStart="FIGURE 3" captionStartId="8.[151,250,1749,1773]" captionTargetBox="[268,1319,189,1714]" captionTargetId="figure-19@8.[259,1328,181,1725]" captionTargetPageId="8" captionText="FIGURE 3. Lebbeus rufomaculosus n. sp., holotype, ovigerous female (cl 16.5 mm), CBM-ZC 16695. A, anterior part of carapace and left cephalic appendages, dorsal view (setae and antennular flagella omitted); B, suborbital lobe and antennal spine of carapace, left side, dorsolateral view; C, telson and left uropod, dorsal view (setae omitted); D, posterior end of telson, dorsal view; E, proximolateral portion of left antennular stylocerite, lateral view; F, left antenna, ventral view (setae and flagellum omitted); G, chela and carpus of right pereopod 1, extensor view; H, tips of fingers of right pereopod 1 (setae omitted); I, junction between merus and ischium of right pereopod 1, showing armature of ventral margin; J, K, L, dactyli of right pereopods 3–5, lateral view; M, coxae of maxilliped 3 and pereopods 1–3, showing development of epipods. Abbreviations: MX3 = maxilliped 3; P = pereopod." figureDoi="http://doi.org/10.5281/zenodo.6521566" httpUri="https://zenodo.org/record/6521566/files/figure.png" pageId="16" pageNumber="167">Fig. 3M</figureCitation>
).
</paragraph>
<paragraph id="C95F36C72A0CFFF8FF71F91DFC587E70" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
The status of 
<taxonomicName id="0EE04D442A0CFFF8FEE9F91DFC927FC8" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[351,462,1699,1725]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FEE9F91DFC927FC8" box="[351,462,1699,1725]" italics="true" pageId="16" pageNumber="167">L. subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FE63F91DFF4F7FC8" bold="true" box="[469,531,1699,1725]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FE63F91DFF4F7FC8" box="[469,531,1699,1725]" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is well supported by the genetic analyses: COI divergence between 
<taxonomicName id="0EE04D442A0CFFF8FAA6F91DF8237FC8" authority="Komai &amp; Matsuzaki, 2022" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[1296,1407,1699,1725]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis" status="sp. nov.">
<emphasis id="FB94EAD52A0CFFF8FAA6F91DF8237FC8" box="[1296,1407,1699,1725]" italics="true" pageId="16" pageNumber="167">L. subtilis</emphasis>
</taxonomicName>
<emphasis id="FB94EAD52A0CFFF8FA30F91DFDE47F94" bold="true" pageId="16" pageNumber="167">
<taxonomicNameLabel id="E0A757AE2A0CFFF8FA30F91DFDE47F94" pageId="16" pageNumber="167" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and the known congeneric species ranges from 4.2 to 19.6% (
<tableCitation id="8462037C2A0CFFF8FCD8F979FE987F94" box="[878,964,1735,1761]" captionStart="TABLE 3" captionStartId="24.[149,237,541,567]" captionTargetPageId="24" captionText="TABLE 3. K2P divergences of the mitochondrial COI gene sequences among species of Lebbeus. Standard error estimates are shown above the diagonal. Codon positions included were 1st+2nd+3rd. All ambiguous positions were removed for each sequence pair (pairwise deletion option). There were a total of 604 bp in the final dataset." httpUri="http://table.plazi.org/id/9D9F664F2A04FFF0FF23FDA3FA2E7B2E" pageId="16" pageNumber="167" tableUuid="9D9F664F2A04FFF0FF23FDA3FA2E7B2E">Table 3</tableCitation>
); 16S divergence ranges from 2.1 to 7.6% (
<tableCitation id="8462037C2A0CFFF8FF29F955FDAA7E70" box="[159,246,1771,1797]" captionStart="TABLE 4" captionStartId="26.[149,237,747,773]" captionTargetPageId="26" captionText="TABLE 4. K2P divergences of the mitochondrial 16 rRNA gene sequences among species of Lebbeus. Standard error estimates are shown above the diagonal. All ambiguous positions were removed for each sequence pair (pairwise deletion option). There were a total of 570 bp in the final dataset." httpUri="http://table.plazi.org/id/9D9F664F2A06FFF2FF23FD55F8E47A5C" pageId="16" pageNumber="167" tableUuid="9D9F664F2A06FFF2FF23FD55F8E47A5C">Table 4</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81FA654C2A0CFFF8FF71F8B1FC1C7E38" pageId="16" pageNumber="167" type="etymology">
<paragraph id="C95F36C72A0CFFF8FF71F8B1FC1C7E38" blockId="16.[151,1437,151,1869]" pageId="16" pageNumber="167">
<emphasis id="FB94EAD52A0CFFF8FF71F8B1FC127E5C" bold="true" box="[199,334,1807,1833]" pageId="16" pageNumber="167">Etymology.</emphasis>
From the Latin “ 
<taxonomicName id="0EE04D442A0CFFF8FDA4F8B1FF3F7E5C" authorityName="Komai &amp; Matsuzaki" authorityYear="2022" box="[530,611,1807,1833]" class="Malacostraca" family="Thoridae" genus="Lebbeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="subtilis">
<emphasis id="FB94EAD52A0CFFF8FDA4F8B1FF3F7E5C" box="[530,611,1807,1833]" italics="true" pageId="16" pageNumber="167">subtilis</emphasis>
</taxonomicName>
” (= slender), in reference to the slender rostrum with a poorly developed ventral lamina.
</paragraph>
</subSubSection>
</treatment>
</document>