159 lines
13 KiB
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159 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.724.21415" ID-GBIF-Dataset="99799f60-7061-439a-855d-53f781229ca9" ID-PMC="PMC5769737" ID-Pensoft-Pub="1313-2970-724-53" ID-PubMed="29362533" ID-ZBK="BCDF3F207E274CCFA47467DA61308A78" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-724-53" ModsDocOrigin="ZooKeys 724" ModsDocTitle="Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species" checkinTime="1513881635527" checkinUser="pensoft" docAuthor="Ivkovic, Marija, Cevid, Josipa, Horvat, Bogdan & Sinclair, Bradley J." docDate="2017" docId="41215C92AF02C646F784120C7462903E" docLanguage="en" docName="ZooKeys 724: 53-100" docOrigin="ZooKeys 724" docSource="http://dx.doi.org/10.3897/zookeys.724.21415" docTitle="Wiedemannia pseudoberthelemyi Ivkovic & Sinclair, sp. n." docType="treatment" docUuid="BD1AFAB9-06BD-4BEC-A08F-F3D80E8FAFE0" docUuidSource="ZooBank" docVersion="5" lastPageNumber="61" masterDocId="FFA4FFD8AC29FFE5CB1EFF8C2D70FFAC" masterDocTitle="Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species" masterLastPageNumber="100" masterPageNumber="53" pageNumber="59" updateTime="1668165242534" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Aquatic dance flies (Diptera, Empididae, Clinocerinae and Hemerodromiinae) of Greece: species richness, distribution and description of five new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ivkovic, Marija</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cevid, Josipa</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Horvat, Bogdan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sinclair, Bradley J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>724</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>53</mods:start>
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<mods:end>100</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.724.21415</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.724.21415</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-724-53</mods:identifier>
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<mods:identifier type="ZBK">BCDF3F207E274CCFA47467DA61308A78</mods:identifier>
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<mods:identifier type="ZooBank">BCDF3F207E274CCFA47467DA61308A78</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="139438682" LSID="urn:lsid:zoobank.org:act:BD1AFAB9-06BD-4BEC-A08F-F3D80E8FAFE0" httpUri="http://treatment.plazi.org/id/41215C92AF02C646F784120C7462903E" lastPageId="8" lastPageNumber="61" pageId="6" pageNumber="59">
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<subSubSection pageId="6" pageNumber="59" type="nomenclature">
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<paragraph pageId="6" pageNumber="59">
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<taxonomicName LSID="http://zoobank.org/BD1AFAB9-06BD-4BEC-A08F-F3D80E8FAFE0" authority="Ivkovic & Sinclair" class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia pseudoberthelemyi" order="Diptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="pseudoberthelemyi">
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Wiedemannia pseudoberthelemyi
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<normalizedToken originalValue="Ivković">Ivkovic</normalizedToken>
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& Sinclair
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</taxonomicName>
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<taxonomicNameLabel pageId="6" pageNumber="59">sp. n.</taxonomicNameLabel>
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Figs 1, 3, 10
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="59" type="type locality">
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<paragraph pageId="6" pageNumber="59">Type locality.</paragraph>
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<paragraph pageId="6" pageNumber="59">Greece: Etolia, River Mornos, Nafpaktos, 38°23'N, 21°51'E.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="59" type="type material">
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<paragraph pageId="6" pageNumber="59">Type material.</paragraph>
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<paragraph pageId="6" pageNumber="59">
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Holotype ♂ (in 80% ethanol), labelled: "HOLOTYPE/
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<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="genus">Wiedemannia</taxonomicName>
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//
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<taxonomicName lsidName="pseudoberthelemyi" pageId="6" pageNumber="59" rank="species" species="pseudoberthelemyi">pseudoberthelemyi</taxonomicName>
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<normalizedToken originalValue="IVKOVIĆ">IVKOVIC</normalizedToken>
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// et SINCLAIR/, GREECE, Etolia,// River Mornos,/ Nafpaktos,/ 38°23'N, 21°51'E, 23.iv.1990,// leg. B. Horvat, I. Sivec" (UZC). Paratypes: same data as holotype (1 ♂, 3 ♀, UZC; 3 ♂, 6 ♀, CNC, dried from alcohol).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="60" pageId="6" pageNumber="59" type="materials_examined">
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<paragraph pageId="6" pageNumber="59">Additional material.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="60" pageId="6" pageNumber="59">
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GREECE: Central Greece, Panaitoliko Mts., R. Tavropos, Kalesmeno, 300 m, 38°56'N, 21°40'E, 29.iv.1989 (SMNH); Central Greece, Etolia, Agrinio, Agia Soufia, 100 m, 38°36'N, 21°26'E, 24.iv.1990 (SMNH);
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<pageBreakToken pageId="7" pageNumber="60" start="start">Etolia</pageBreakToken>
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, Vardousia Mts., R. Evinos, Grammeni Oxia, 800 m, 38°43'N, 22°00'E, 28.iv.1990 (SMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="60" type="diagnosis">
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<paragraph pageId="7" pageNumber="60">Diagnosis.</paragraph>
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<paragraph pageId="7" pageNumber="60">
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This species of
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<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="genus">Wiedemannia</taxonomicName>
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is distinguished by the long gena and the mitten-shaped clasping cercus, which is extremely similar to that of
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<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia (Chamaedipsia) berthelemyi" order="Diptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="berthelemyi" subGenus="Chamaedipsia">Wiedemannia (Chamaedipsia) berthelemyi</taxonomicName>
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Vaillant &
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<normalizedToken originalValue="Vinçon">Vincon</normalizedToken>
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, 1987.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="60" type="description">
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<paragraph pageId="7" pageNumber="60">Description.</paragraph>
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<paragraph pageId="7" pageNumber="60">Male. Body length 3.5-4.1 mm, wing length 3.5-3.6 mm (colouration bleached by prolonged storage in alcohol). Head (Fig. 3) in lateral view higher than long; gena broad, three-quarters height of eye. Frons short, broader than face. Face wide, with distinct carina on lower margin, bare, lacking setae. One pair of short ocellar setae and one pair of vertical setae; about 6 distinct upper postocular setae; lower postocular setae finer and merging with longer setae on middle and lower occiput; few small setulae present on vertex and between ocellar area. Antenna brown; postpedicel and stylus minutely pubescent; scape longer than pedicel, with setulae dorsally; pedicel with complete circlet of apical setae; postpedicel apically pointed; stylus nearly twice length of postpedicel.</paragraph>
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<paragraph pageId="7" pageNumber="60">Mesonotum with 5 pairs of dorsocentral setae with short setulae interspersed. Acrostichal setae short and fine, biserial, extending to prescutellar depression; 1 strong postpronotal seta; 2-3 notopleural setae and several short setae of variable size; 1 presutural supra-alar seta and numerous small setulae; 1 postalar seta. Antepronotum with 1 pair of strong setae and a few shorter setae. Proepisternum with some fine setulae. Katepisternum with a few (1-3) setulae. Laterotergite with fine, pale setae. One pair of strong marginal scutellar setae with many scattered setae on disc.</paragraph>
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<paragraph pageId="7" pageNumber="60">Wing membrane infuscate, veins darker; 1 short basal costal seta ending before humeral crossvein. Cell dm produced anteroapically. Veins M1 and M2 originating together with short stem vein proximal to M1+2 fork. Vein CuA+CuP extremely faint. Pterostigma elongate, indistinct. Squama with setulae. Halter yellowish.</paragraph>
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<paragraph pageId="7" pageNumber="60">Legs mostly brown; fore femur with 1 strong preapical anterior seta; uniformly covered with rows of small dark setulae. All coxae with longer setae anteriorly. Fore and mid femora ventrally with some longer setulae on proximal half.</paragraph>
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<paragraph pageId="7" pageNumber="60">Abdomen concolourous with thorax, covered in short setae. Pubescence darker on tergites than sternites. Terminalia (Fig. 10): hypandrium subequal in length with epandrium, with 6 pairs of setae. Epandrium irregularly subquadrate, with 2-3 stouter and longer setae (shown by enlarged sockets) in addition to regular setae ventrally and laterally; surstylus very slender, hook-shaped. Clasping cercus yellowish-brown, broad, mitten-shaped, with thumb-like anterior lobe; posterior lobe truncate apically; thumb-like lobe with long outer setae; stout setae with multi-branched apex covering most of inner face of cercus. Phallus more or less linear, slender; distiphallus with swelling at mid-length.</paragraph>
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<paragraph pageId="7" pageNumber="60">Female. Similar to male. Terminalia: cercus short ovate and minutely pilose.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="60" type="etymology">
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<paragraph pageId="7" pageNumber="60">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="60">
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The species name is derived from the name
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<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia (Chamaedipsia) berthelemyi" order="Diptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="berthelemyi" subGenus="Chamaedipsia">Wiedemannia (Chamaedipsia) berthelemyi</taxonomicName>
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because of the similarity of the clasping cercus with that of this species.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60" type="remarks">
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<paragraph pageId="7" pageNumber="60">Remarks.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60">
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<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia pseudoberthelemyi" order="Diptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="pseudoberthelemyi">Wiedemannia pseudoberthelemyi</taxonomicName>
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sp. n. is known only from parts of Greece. This new species differs from
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<taxonomicName lsidName="W. berthelemyi" pageId="7" pageNumber="60" rank="species" species="berthelemyi">W. berthelemyi</taxonomicName>
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on the basis of the truncate pos
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<pageBreakToken pageId="8" pageNumber="61" start="start">terior</pageBreakToken>
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lobe of the clasping cercus (pointed in
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<taxonomicName lsidName="W. berthelemyi" pageId="8" pageNumber="61" rank="species" species="berthelemyi">W. berthelemyi</taxonomicName>
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) and in having only a single preapical seta on the fore femur (2 in
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<taxonomicName lsidName="W. berthelemyi" pageId="8" pageNumber="61" rank="species" species="berthelemyi">W. berthelemyi</taxonomicName>
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). The odd stout setae with multi-branched tips on the inner face of the clasping cercus were not noted by
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<bibRefCitation pageId="8" pageNumber="61">
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Vaillant and
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<normalizedToken originalValue="Vinçon">Vincon</normalizedToken>
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(1987)
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</bibRefCitation>
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and the absence of the swelling on the distiphallus (
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<bibRefCitation pageId="8" pageNumber="61">
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Vaillant and
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<normalizedToken originalValue="Vinçon">Vincon</normalizedToken>
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1987
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</bibRefCitation>
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, fig. 32) is likely an artefact caused by the acid clearing process. The holotype of
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<taxonomicName lsidName="W. berthelemyi" pageId="8" pageNumber="61" rank="species" species="berthelemyi">W. berthelemyi</taxonomicName>
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was not examined.
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</paragraph>
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<paragraph pageId="8" pageNumber="61">
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Additional similar species that could be included in this group based on the shape of the clasping cercus include:
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<taxonomicName lsidName="W. angelieri" pageId="8" pageNumber="61" rank="species" species="angelieri">W. angelieri</taxonomicName>
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Vaillant, 1967 (Pyrenees),
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<taxonomicName lsidName="W. vedranae" pageId="8" pageNumber="61" rank="species" species="vedranae">W. vedranae</taxonomicName>
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<normalizedToken originalValue="Ivković">Ivkovic</normalizedToken>
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& Sinclair, 2014 (Sierra Nevada, Spain), and
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<taxonomicName lsidName="W. queyrasiana" pageId="8" pageNumber="61" rank="species" species="queyrasiana">W. queyrasiana</taxonomicName>
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Vaillant, 1956 (European Alps).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |