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<document ID-CLB-Dataset="56262" ID-DOI="http://dx.doi.org/10.3897/zookeys.1170.103834" ID-GBIF-Dataset="bc6405f2-e1d8-4e3d-8652-b0cd68b396f1" ID-PMC="PMC10372772" ID-Pensoft-Pub="1313-2970-1170-1" ID-Pensoft-UUID="AB30D9571F635294A0C3C7126F1CAF36" ID-PubMed="37521854" ID-ZooBank="17F0C88A2F0B414AAA7C8B0AB89B6E6E" ModsDocID="1313-2970-1170-1" checkinTime="1689838476568" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Hajek, Jiri, Hendrich, Lars, Surbakti, Suriani, Panjaitan, Rawati &amp; Balke, Michael" docDate="2023" docId="403F20D6F179587AB69A53737961415B" docLanguage="en" docName="ZooKeys 1170: 1-164" docOrigin="ZooKeys 1170" docPubDate="2023-07-19" docSource="http://dx.doi.org/10.3897/zookeys.1170.103834" docTitle="Austrelatus adelbert Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke 2023, comb. nov." docType="treatment" docVersion="4" id="AB30D9571F635294A0C3C7126F1CAF36" lastPageNumber="1" masterDocId="AB30D9571F635294A0C3C7126F1CAF36" masterDocTitle="Austrelatus gen. nov., a new genus of Australasian diving beetles (Coleoptera, Dytiscidae, Copelatinae), with the discovery of 31 new species from New Guinea" masterLastPageNumber="164" masterPageNumber="1" pageNumber="1" updateTime="1732590595498" updateUser="ExternalLinkService">
<mods:mods id="2AD7B796AAE1528B048CFF995529D2AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5D4E8579088EAC269BC326828645C117">
<mods:title id="F1EC055E2604D4464C05E7ADE4C003C6">Austrelatus gen. nov., a new genus of Australasian diving beetles (Coleoptera, Dytiscidae, Copelatinae), with the discovery of 31 new species from New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="0CDD600F3258C0832E538FED154A1389" type="personal">
<mods:role id="45D26E4556806D6251233901F9D356A9">
<mods:roleTerm id="B57EE0C2999DFA183A3B3BFBC758A12C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F6BC1E648A03C08C011A0AD89D4ED4A">Shaverdo, Helena</mods:namePart>
<mods:nameIdentifier id="BCE79FEC858D3970811C5A76751CF1A0" type="ORCID">https://orcid.org/0000-0001-5034-7342</mods:nameIdentifier>
<mods:affiliation id="0AF3A34521F2A904F09B1AFC94FD8D1F">Naturhistorisches Museum Wien, Burgring 7, 1010, Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="028310CC0F52CA8F89C043ABC9D95ABE" type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="6F753BAD7F8DAAB93B4E06FF828C2166" type="personal">
<mods:role id="F00BB75046972A9B87031AFA4F7B3EAD">
<mods:roleTerm id="A3D6AD9C2E03F1BE217376533DEC9D4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36479A5DB230C7A7181C7225E621425F">Hajek, Jiri</mods:namePart>
<mods:nameIdentifier id="8E7950AD1015B01AA4DF524139D8FEFA" type="ORCID">https://orcid.org/0000-0001-5779-1542</mods:nameIdentifier>
<mods:affiliation id="62CE1B9CA20B8BA4DF2DA6696C165D75">Department of Entomology, National Museum, Cirkusova 1740, CZ- 193 00 Praha 9 - Horni Pocernice, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="4C71FF86490DEC276C8B2F0CBE737493" type="personal">
<mods:role id="BBEA9DCA074243722B35F0559B42E47F">
<mods:roleTerm id="5D4600CFFFD690199313E696274EE402">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F86A291C76E331E32485F70610AEEAB">Hendrich, Lars</mods:namePart>
<mods:nameIdentifier id="905454254FC3A7263BF9533CF66556B4" type="ORCID">https://orcid.org/0000-0001-8366-0749</mods:nameIdentifier>
<mods:affiliation id="623648DFC028683999283A9335ABB7CF">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:name id="40BA6C1A3C0C79554CC1C3FBE1150A3E" type="personal">
<mods:role id="AB5254C5EFBE9B5745267E153804DFD8">
<mods:roleTerm id="ACDF9E32BAFD76F1E6E602D93BFB26D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B2129D77820B1BBB56370B312D5A8A3">Surbakti, Suriani</mods:namePart>
<mods:nameIdentifier id="7686D1E82DDC249D7F9C06936AD25CD0" type="ORCID">https://orcid.org/0000-0003-3984-2187</mods:nameIdentifier>
<mods:affiliation id="281A611370FA301560588841F0729335">Department of Biology, Universitas Cendrawasih, Jayapura, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="987F37B4DA7F7911B269A88B393DE9C4" type="personal">
<mods:role id="A03FE34BBE0E2F75D2C0171A01EBFD56">
<mods:roleTerm id="5EE111183B9C5D752B92F800418DC6E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB81B1115EB1A5598CC77B585EA9A095">Panjaitan, Rawati</mods:namePart>
<mods:affiliation id="2EE4D2AD3CC54164475E0382C67000EC">Department of Biology, Faculty of Sciences and Mathematics, State University of Papua (UNIPA), Jalan Gunung Salju Amban, Manokwari 98314, West Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="CC42C128EB8B0FE0DA9CC8CF417813A9" type="personal">
<mods:role id="2B44293BB721F394432631B2A4A07B3F">
<mods:roleTerm id="EAE49E7C3785D724254D26C8E811AF3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="023FAACDC3CE301E1D428242417D9EA2">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="F972AE9B48EAF4B478F65A8A9A0B7724" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="72243CCEB903956B80C650389B9EB507">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1C5549B01FBDB2363D50379B9C4DE8D8">text</mods:typeOfResource>
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<mods:titleInfo id="15260D08FE4590295D84EECC15E7D07E">
<mods:title id="27AF946568EF13A2F129B793393D713B">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="2C8E111628C8A5D05C5E75092B68853E">
<mods:date id="A371A2DFA15DA24FA70E69D08C789301">2023</mods:date>
<mods:detail id="983FEBFB51840B3B11CF7AEAD5A82DF9" type="pubDate">
<mods:number id="7B8122E232E98D9FB2A9E8889B41A170">2023-07-19</mods:number>
</mods:detail>
<mods:detail id="89F6A99725CC1E780EC4AE0B38FCEF70" type="volume">
<mods:number id="873037DE8E8127F17ABCBBBD9D2995B8">1170</mods:number>
</mods:detail>
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<mods:start id="1B610AA1699907026998E6F427665767">1</mods:start>
<mods:end id="548E580E8D2D89BACB28D7ED8C5D036B">164</mods:end>
</mods:extent>
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<mods:location id="F9291E85B2332351579DF056370EEE5E">
<mods:url id="87B0C70C0DA27A7D6881A3B9DA0F4B68">http://dx.doi.org/10.3897/zookeys.1170.103834</mods:url>
</mods:location>
<mods:classification id="5AFAA3B1D599E62EE9A03119854A69B5">journal article</mods:classification>
<mods:identifier id="02F9CBBCE71862EAAD26DDE09DC4E50F" type="DOI">http://dx.doi.org/10.3897/zookeys.1170.103834</mods:identifier>
<mods:identifier id="ED6E0F5B6ECF1770801648218A7C7C7C" type="Pensoft-Pub">1313-2970-1170-1</mods:identifier>
<mods:identifier id="B1C68E87FBBEA8C50C0324EEDFB28D13" type="ZooBank">17F0C88A2F0B414AAA7C8B0AB89B6E6E</mods:identifier>
<mods:identifier id="C0968303B1B8DBC6941BC7F4D5561162" type="Pensoft-UUID">AB30D9571F635294A0C3C7126F1CAF36</mods:identifier>
</mods:mods>
<treatment id="403F20D6F179587AB69A53737961415B" ID-GBIF-Taxon="212415879" LSID="urn:lsid:plazi:treatment:403F20D6F179587AB69A53737961415B" httpUri="http://treatment.plazi.org/id/403F20D6F179587AB69A53737961415B" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="DF46AFC3F1727EA94EAAA85FBEDB7212" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="3D4EFFA85D55878AF4BFDC6DF23D2B38" pageId="0" pageNumber="1">
1.
<taxonomicName id="B495257D00F7BBCE948FDB023DE531E7" LSID="403F20D6-F179-587A-B69A-53737961415B" authority="(Megna, Atthakor, Manaono, Hendrich &amp; Balke, 2017)" authorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" authorityYear="2023" baseAuthorityName="Megna, Atthakor, Manaono, Hendrich &amp; Balke" baseAuthorityYear="2017" class="Insecta" family="Dytiscidae" genus="Austrelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrelatus adelbert" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="adelbert" status="comb. nov.">Austrelatus adelbert (Megna, Atthakor, Manaono, Hendrich &amp; Balke, 2017)</taxonomicName>
<taxonomicNameLabel id="4D9250055D154C0F82FE3F47D6231DAE" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F788536C73809680036902EDCB2622C6" pageId="0" pageNumber="1" type="description">
<paragraph id="399EB46654B8BCBC8F803847832AD7D5" pageId="0" pageNumber="1">
<figureCitation id="2125E4811945F6A82E13FDB3348F5000" captionStart="Figures 14–17" captionStartId="F12" captionText="Figures 14 - 17. Habitus and colouration of 14 Austrelatus adelbert (Megna et al., 2017) 15 A. lembenensis sp. nov. 16 A. nadjae sp. nov. 17 A. pseudoneoguineensis sp. nov." figureDoi="10.3897/zookeys.1170.103834.figures14-17" httpUri="https://binary.pensoft.net/fig/881119" pageId="0" pageNumber="1">Figs 14</figureCitation>
<figureCitation id="3E46602AC3964BEDF0C847FA8226CC2F" captionStart="Figure 18" captionStartId="F13" captionText="Figure 18. Austrelatus adelbert (Megna et al., 2017), median lobe A left lateral view B right lateral view C ventral view D left paramere in external view E right paramere in internal view. Scale bar: one unit - 0.1 mm (A-C)." figureDoi="10.3897/zookeys.1170.103834.figure18" httpUri="https://binary.pensoft.net/fig/881120" pageId="0" pageNumber="1">, 18</figureCitation>
<figureCitation id="FB505DC19DD1FC58B01EDF362346E6E2" captionStart="Figure 82" captionStartId="F53" captionText="Figure 82. Map of New Guinea showing distribution of 11 Austrelatus species." figureDoi="10.3897/zookeys.1170.103834.figure82" httpUri="https://binary.pensoft.net/fig/881160" pageId="0" pageNumber="1">, 82</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C01E25603ED4BCF2846B42BA5185B4B6" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="E7AF914FF657E0C3377DE3DE6FF001FD" pageId="0" pageNumber="1">
<taxonomicName id="FA22595A4383ECCC849459DC313CB4C0" authorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" authorityYear="2023" baseAuthorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" baseAuthorityYear="2023" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus adelbert" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="adelbert" status="comb. nov.">Copelatus adelbert</taxonomicName>
Megna, Atthakor, Manaono, Hendrich &amp; Balke, 2017: 50.
</paragraph>
</subSubSection>
<subSubSection id="26F40BA913923FEB642EC79F966C35BC" pageId="0" pageNumber="1" type="type locality">
<paragraph id="BF66C9246A2DF3EF98CB11F0DBFA770F" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="56C352A02A2D766900F0EBC632588100" pageId="0" pageNumber="1">
Papua New Guinea: Madang Province, Adelbert Range, Keki birdwatching area,
<geoCoordinate id="22D8D8262E3D560C00AB4F3F0DE117EA" degrees="04" direction="south" minutes="42" orientation="latitude" precision="1" seconds="21.5" value="-4.705972">04°42'21.5&quot;S</geoCoordinate>
,
<geoCoordinate id="7766B0FC9E1B2B0A25713A69E717E01E" degrees="145" direction="east" minutes="25" orientation="longitude" precision="1" seconds="15.4" value="145.42094">145°25'15.4&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="764DA9897B80F36E06DCF7955B6DA1AA" pageId="0" pageNumber="1" type="type material">
<paragraph id="B6573E255B899BC597B24B33CCB4E26D" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="440194E8E8A9A9E33FECD93E792EF87B" pageId="0" pageNumber="1">
<emphasis id="E882E20921CE73EA177173E0DECD9947" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: male &quot;Papua New Guinea: Madang, Adelbert Mts., Sewan - Keki, 700 m, 04°42'215&quot;S, 145°25'154&quot;E, 4.v.2006, leg Balke &amp; Manaono (PNG 51)&quot;, &quot;HOLOTYPE
<taxonomicName id="C25E589B636E5AF8F430FA89DF48AF33" authorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" authorityYear="2023" baseAuthorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" baseAuthorityYear="2023" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus adelbert" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="adelbert">Copelatus adelbert</taxonomicName>
sp. n. Det.: Y. S. Megna, W. Atthakor, M. Manaono, L. Hendrich &amp; M. Balke 2017&quot; [red],
<normalizedToken id="68CBEF07FD8D820AEFB1F796E1A10016" originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
(ZSM).
</paragraph>
<paragraph id="6D6C288BCB7E3468323F740AB9F92101" pageId="0" pageNumber="1">
<emphasis id="4112BA45ABC03FF738708C0ED8026F6F" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 1 male, 1 female with the same label as the holotype and with &quot;PARATYPE
<taxonomicName id="DE9EF618C12CCB10C35279AED79A3FED" authorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" authorityYear="2023" baseAuthorityName="Shaverdo &amp; Hájek &amp; Hendrich &amp; Surbakti &amp; Panjaitan &amp; Balke" baseAuthorityYear="2023" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus adelbert" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="adelbert">Copelatus adelbert</taxonomicName>
sp. n. Det.: Y. S. Megna, W. Atthakor, M. Manaono, L. Hendrich &amp; M. Balke 2017&quot; [red] (ZSM). 2 males &quot;Papua New Guinea: Madang, Adalbert [sic!] Mts., Keki, 850 m, 4.v.2006, nr 04.42.300S 145.25.089E, Balke &amp; Manaono (PNG 52)&quot; (ZMS). 3 males, 3 females &quot;Papua New Guinea: Madang, Adalbert [sic!] Mts., below Keki, 790 m, 5.v.2006, 04.42.300S 145.25.089E, Balke &amp; Manaono (PNG 53)&quot; (ZMS). 1 male, 2 females &quot;Papua New Guinea: Madang, Adalbert [sic!] Mts., creek nr Keki, 790 m, 26.xi.2006, 04.42.300S 145.25.089E, Binatang Boys leg. (PNG 53a)&quot; (ZMS).
</paragraph>
</subSubSection>
<subSubSection id="3A1ABFE606A91DC32DE6C1E478D019B2" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="3758F2A6C16320EB77F7F7108FD5A4B5" pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph id="9E55634EF04884F07BBD70379438C09D" pageId="0" pageNumber="1">
<materialsCitation id="CDC31D9C4778A3A9C7707AD6F0FACCBF" collectingDate="2006-05-04" collectingDateMax="2006-11-29" collectingDateMin="2006-05-04" collectorName="Balke, Manaono, Keki, Adalbert, Mts., Kinibel" country="Papua New Guinea" elevation="850" location="Keki" municipality="Adalbert" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Madang">
<specimenCount id="93E83FBB3C0FA99888D425C0992D20A7" type="male">1 male</specimenCount>
<normalizedToken id="4B955BDDF9EE7632C408CB439F54A314" originalValue="“3204”">&quot;3204&quot;</normalizedToken>
[green label], &quot;
<collectingCountry id="0A7CC1B3BFF478B31E541BC728251CA1" name="Papua New Guinea">Papua New Guinea</collectingCountry>
:
<collectingRegion id="A1331D776CC964A419C95B8413F5D9A5" country="Papua New Guinea" name="Madang">Madang</collectingRegion>
,
<collectingMunicipality id="09FCE226DE91A5FAFF2E8EF0A41C3948">Adalbert</collectingMunicipality>
[sic!] Mts.,
<location id="2393B810483D8A35253D414A6C950A7E" LSID="urn:lsid:plazi:treatment:403F20D6F179587AB69A53737961415B:2393B810483D8A35253D414A6C950A7E" country="Papua New Guinea" municipality="Adalbert" name="Keki" stateProvince="Madang">Keki</location>
,
<quantity id="6596A777EC645182C8BD49F519537981" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<elevation id="A627339D4DEC95107C4EE49651CE977E" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<collectingDate id="5976CC88A4F26CB516575633BAF119A5" value="2006-05-04">4.v.2006</collectingDate>
, nr 04.42.300S 145.25.089E,
<collectorName id="FB5CE43B4024FA1AE9FAFED53407FFF1">Balke</collectorName>
&amp;
<collectorName id="047ABE492EC084E3B4196A215BB626EF">Manaono</collectorName>
(PNG 52)&quot; (ZMS).
<specimenCount id="8A6DD6DBCE41BD042E7F33F603DEAFC5" type="male">1 male</specimenCount>
&quot;
<collectingCountry id="3DC1F07088FE5E4BB327AA468731BF5A" name="Papua New Guinea">Papua New Guinea</collectingCountry>
:
<collectingRegion id="9DDCCFD83FC18590EB9FE3AE401026D0" country="Papua New Guinea" name="Madang">Madang</collectingRegion>
, Adalbert [sic!] Mts.,
<quantity id="EA6B6DE3D1245E8CDD556A3FC235E465" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="DDA0B026784F2061BB7E75CBC47D8616" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<collectingDate id="AEACFD50E9617092278813E782433FC3" value="2006-11-29">29.xi.2006</collectingDate>
, 04.43.058S 145.24.437E, Binatang Boys (PNG 119)&quot; (ZMS).
<specimenCount id="0F429CAE629E054B29C087AEAC86A0F8" type="male">3 males</specimenCount>
,
<specimenCount id="C7D550388911A00243FAECBB0F481668" type="female">1 female</specimenCount>
&quot;
<collectingCountry id="CD21263644D73F8EB028C0C2AA8D2902" name="Papua New Guinea">Papua New Guinea</collectingCountry>
:
<collectingRegion id="40B0C75328816FD988BB59CA62BB64B5" country="Papua New Guinea" name="Madang">Madang</collectingRegion>
,
<collectorName id="2B87A6399B7ED6D173C47EC219AA8798">Keki</collectorName>
,
<collectorName id="F328A5B62D1C537E8DE99491A021974B">Adalbert</collectorName>
[sic!]
<collectorName id="3B23BB01765CC2B69C4B87499F3652F5">Mts.</collectorName>
,
<quantity id="A6A6BD116C9AB72238866BB630ABC78D" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="1A168168C65226C19BD1DBDAE4AAE592" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<collectingDate id="95E414B6757B14DB6C35358BD17167AC" value="2006-11-29">29.xi.2006</collectingDate>
, 04.43.058S 145.24.437E,
<collectorName id="61BEAB7038BC045B83766521184E5BDE">Balke</collectorName>
&amp;
<collectorName id="2BC3EA0E00C648ABA0CF313C09458C19">Kinibel</collectorName>
(PNG 118)&quot; (NHMW, ZMS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F54D9828E92464AAD76C3E57C93CCC14" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="C2E225F718FF852ED58B2412A9AEF56F" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="BA44249E4593E68142B70F63EB2F1222" pageId="0" pageNumber="1">
For complete description, see
<bibRefCitation id="97205D340298C511CFD690DB3B11E2CD" author="Megna, YS" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="1" pagination="49 - 55" publicationUrl="https://search.informit.org/doi/10.3316/informit.718392650619836" refId="B23" refString="Megna, YS, Atthakor, W, Manaono, M, Hendrich, L, Balke, M, 2017. A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea. Australian Entomologist 44 (1): 49 - 55, https://search.informit.org/doi/10.3316/informit.718392650619836" title="A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea." url="https://search.informit.org/doi/10.3316/informit.718392650619836" volume="44" year="2017">Megna et al. (2017</bibRefCitation>
: 51-53).
</paragraph>
<paragraph id="0AE19C432BBC6854AA0507E5D770E8B7" pageId="0" pageNumber="1">
Beetle medium-sized: TL 5.9-6.4 mm, TL-H 5.5-5.9 mm, with oblong-oval habitus of continuous outline. Dorsal colouration very distinct: yellow to orange head, black pronotum with yellow to orange lateral bands, elytron with broad basal and apical yellow to orange parts. Elytron without striae and strioles in both sexes. Proclaws relatively long, slender, equal in in length. Median lobe of aedeagus relatively broad; its dorsal and ventral sclerites almost equal in length, relatively thin apically, pointed; ventral sclerite weakly sclerotised: its right lobe membranous, long, and usually sticking out ventrally, with small lateral sclerotised area; left ventral lobe shorter, almost completely but weakly sclerotised, without a lateral sclerotised area. Paramere of narrow triangular form, more or less evenly tapering to apex, with setae not divided into distal and proximal, slightly sparser and shorter proximally (Figs
<figureCitation id="B635654B55FB328F2A20AEA6AC19DC12" captionStart="Figures 14–17" captionStartId="F12" captionText="Figures 14 - 17. Habitus and colouration of 14 Austrelatus adelbert (Megna et al., 2017) 15 A. lembenensis sp. nov. 16 A. nadjae sp. nov. 17 A. pseudoneoguineensis sp. nov." figureDoi="10.3897/zookeys.1170.103834.figures14-17" httpUri="https://binary.pensoft.net/fig/881119" pageId="0" pageNumber="1">14</figureCitation>
,
<figureCitation id="81060AC613C054719F4466CFF060F6F9" captionStart="Figure 18" captionStartId="F13" captionText="Figure 18. Austrelatus adelbert (Megna et al., 2017), median lobe A left lateral view B right lateral view C ventral view D left paramere in external view E right paramere in internal view. Scale bar: one unit - 0.1 mm (A-C)." figureDoi="10.3897/zookeys.1170.103834.figure18" httpUri="https://binary.pensoft.net/fig/881120" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="69A233BF7A339718F528A5DAE97BE64A" pageId="0" pageNumber="1" type="affinities">
<paragraph id="95E13D17F278B6438D4158A3953B6CD1" pageId="0" pageNumber="1">Affinities.</paragraph>
<paragraph id="2B78B20D45C973BDBFF6F769E92C2869" pageId="0" pageNumber="1">
Due to its dorsal colouration and smooth elytra, the species cannot be mixed up with any other NG species. In addition, it differs from all other representatives of the
<taxonomicName id="B20C8C6BB50D75A1D38DA1FC1B0D1476" lsidName="A. neoguineensis" pageId="0" pageNumber="1" rank="species" species="neoguineensis">
<emphasis id="36D04CB9FF8FE44A21674C99027873F7" italics="true" pageId="0" pageNumber="1">A. neoguineensis</emphasis>
</taxonomicName>
group in less modified sclerites of the median lobe and less sclerotised ventral sclerite (left ventral lobe without lateral sclerotised area).
</paragraph>
</subSubSection>
<subSubSection id="4EB160A530F60E7F5B75A49404ECDB85" pageId="0" pageNumber="1" type="distribution">
<paragraph id="9FEA45806449729766BFEE6E523FD14A" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="D98E582E62FF758B29062791FD98AC26" pageId="0" pageNumber="1">
New Guinean endemic. Papua New Guinea: Madang Province, Adelbert Range (Fig.
<figureCitation id="59D8354A0A26CC12028A40922289A516" captionStart="Figure 82" captionStartId="F53" captionText="Figure 82. Map of New Guinea showing distribution of 11 Austrelatus species." figureDoi="10.3897/zookeys.1170.103834.figure82" httpUri="https://binary.pensoft.net/fig/881160" pageId="0" pageNumber="1">82</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="444E05C8ADB256E0298F1D3C96BBAFC6" pageId="0" pageNumber="1" type="habitat">
<paragraph id="22EB81DE0616166CC45F5C741C1F93CC" pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph id="FB6CB3ECE5A179E671C10721DCCC3AAE" pageId="0" pageNumber="1">
The species is collected from small puddles along forest creeks; the ground was red clay with sand, gravel and sometimes leaves (
<bibRefCitation id="88CF05D147819951900A9E38AFA17950" author="Megna, YS" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="1" pagination="49 - 55" publicationUrl="https://search.informit.org/doi/10.3316/informit.718392650619836" refId="B23" refString="Megna, YS, Atthakor, W, Manaono, M, Hendrich, L, Balke, M, 2017. A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea. Australian Entomologist 44 (1): 49 - 55, https://search.informit.org/doi/10.3316/informit.718392650619836" title="A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea." url="https://search.informit.org/doi/10.3316/informit.718392650619836" volume="44" year="2017">Megna et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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