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<document ID-CLB-Dataset="7610" ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="3F398A2222BB50968728D1949338C2C4" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina waaf Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="F172B13F-AF66-40E4-B0C3-67267C6811C3" docUuidSource="ZooBank" docVersion="7" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1732748175969" updateUser="ExternalLinkService">
<mods:mods id="77CCEDC570C3F9C8AE2AB6CE1C64B8E5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8EB1A7DEE23E3277375870666A82071D">
<mods:title id="90ED467BB78BBB7F50B87D88654CFDA5">Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="DF875C185FB0C9A2F7BF7F3AF46461A7" type="personal">
<mods:role id="73CAA67B4B1D468CC5902B7C83718770">
<mods:roleTerm id="5AA1AA58266009EE3FAA301F16D1DE52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="654549A06CC1652237DD61E986106A73">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="D10E547D5303697AEA73EE9E5E78BC4A">Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="34B14C4510D57A8699F8B3E41A6C72BF" type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="D54D6EFC1DAECEF9401E702F8C643A26" type="personal">
<mods:role id="A208A91D0C107C028BAA3E0D96B48799">
<mods:roleTerm id="79203ABC4C8A882BCB5BAD908B7A055F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05AAB45247DC4C2B9637DABADA59C261">Surbakti, Suriani</mods:namePart>
<mods:affiliation id="79716EA8EE20192EF9CC130703DFBBE2">Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="A2A0C8CA0EDCB57DB723F79A24C24602" type="personal">
<mods:role id="613445EE8C53C1A19F978C9BCD402CAD">
<mods:roleTerm id="4C5D1CA9C294814972C351C643652408">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C4CE7FE48FAD4A2B7335343C0F74488">Sumoked, Bob</mods:namePart>
<mods:affiliation id="490EF2028B3767F5E7921D45B844DC37">Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="337BFF3395194097F67DA005AD21D3D0" type="personal">
<mods:role id="996098BAD294D79A9B415D0FB9CD43C4">
<mods:roleTerm id="6A50933173CBE3CE1212AA0E698C9A4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EDBF6A89B2BCDE8FA9A70762D22E11C">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="38177C0079C28AF4C7E994BB4F914636" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="D9AD8AE004684266C5181B0AFBBE093D">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F4D702CDFA73B65F0A35329495D0531B">text</mods:typeOfResource>
<mods:relatedItem id="770D21358CF9DE0ADD4B8BF465854BAA" type="host">
<mods:titleInfo id="5348856B13E48326E38DC23F50C9F618">
<mods:title id="2912AAF0E455FD18370329BC3036F9EC">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="40D531818A1F5517ED4C0A64196883F7">
<mods:date id="6118F1BD5C593862DC342790D53BD7A9">2021</mods:date>
<mods:detail id="049F2C6FA3471E797147798A9A63FE74" type="pubDate">
<mods:number id="C5A319C34C18F2D641A728EC0EE84053">2021-03-25</mods:number>
</mods:detail>
<mods:detail id="6686BCE9755162A58FB62EE0858CB592" type="volume">
<mods:number id="BCCC27FEF4C0A861646F78CE66154F66">1026</mods:number>
</mods:detail>
<mods:extent id="E7324B3D5665507302701100B1FE2A06" unit="page">
<mods:start id="A42B01BEB9ADFB23691D9946A11ED83C">45</mods:start>
<mods:end id="52254D5960DE917C565B57A4642E9DB4">67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1F4D9A68BC9EBBFEB5BB308F354C3A75">
<mods:url id="32CC737F3BF0AB2EB42F2D5100E3FF43">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification id="25A3445E2F0B6B71BF8C988A1B995FF0">journal article</mods:classification>
<mods:identifier id="F59AAAA899F20EB391764DF1AC151DBE" type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier id="B9015A26E9A3EFCE058B95BCA9E17162" type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier id="BE20C9A71D4E34B1E2CDC76794A45153" type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier id="6D9BB4BF1AEB7DBEF0B5FAB451730E71" type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment id="3F398A2222BB50968728D1949338C2C4" ID-GBIF-Taxon="180680997" LSID="urn:lsid:zoobank.org:act:F172B13F-AF66-40E4-B0C3-67267C6811C3" httpUri="http://treatment.plazi.org/id/3F398A2222BB50968728D1949338C2C4" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="42D7CA1720ED4F244A98C487F4503B93" pageId="0" pageNumber="45" type="nomenclature">
<paragraph id="E6426772BE72A87B8B9CB2DB9C5413FB" pageId="0" pageNumber="45">
<taxonomicName id="DF8D8F5B223B866421440E41A9C36F8E" ID-CoL="8LZ8S" LSID="http://zoobank.org/F172B13F-AF66-40E4-B0C3-67267C6811C3" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina waaf" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="waaf" status="sp. nov.">Exocelina waaf Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel id="30E8B692537D73AAFB813576F300C41C" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation id="21A1E907CBADEE25A47A95695BF30A51" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 2</figureCitation>
<figureCitation id="C48F28748CF511452DDF8C4C5D6551C6" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">, 6</figureCitation>
<figureCitation id="24EC540032716580608950A0DF6B7613" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9352961E1A25BA2A59E4ADE499A564FE" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="B7A18C30C91E873B30392C79F0695FC6" pageId="0" pageNumber="45">
<taxonomicName id="0FE18B6E3F7372E98B9F8A01F9F21FC3" authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina utowaensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="utowaensis" status="sp. nov.">Exocelina nr. utowaensis</taxonomicName>
#7281:
<bibRefCitation id="E5F9EFB3D97239A916F60A013FFDF211" DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection id="6A72CCE41F5C3E361F958B10C725DE1A" pageId="0" pageNumber="45" type="type locality">
<paragraph id="E1575CC035BE86EFDF2027ACACBADB72" pageId="0" pageNumber="45">
<typeStatus id="F8B8C5DBA68D2B50FFA8B9108BF6AF56">Type</typeStatus>
locality.
</paragraph>
<paragraph id="7C68412C2042FC24A163A6B49B4BD60F" pageId="0" pageNumber="45">
<materialsCitation id="61E689F02A980384BC0507A7CAA5721D" ID-GBIF-Occurrence="3062138301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Foja Mts, N" country="Indonesia" elevation="115" latitude="-2.374889" location="Waaf Village" longLatPrecision="1" longitude="138.73886" municipality="Foja Mts" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry id="D2FA1408212D09D351B37DABB07A696B" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="CC7FAF0F29D78F72F94E63909B8CA633" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
, Sarmi Regency,
<collectorName id="39900A03EF36C47589288BE4861B6D4A">
<collectingMunicipality id="B42C4BCB2EC63D8F18E02E9A4798A576">Foja Mts</collectingMunicipality>
, N
</collectorName>
<location id="0F45CD7ADC66DF13A5A1ACD7FF6ABB8C" LSID="urn:lsid:plazi:treatment:3F398A2222BB50968728D1949338C2C4:0F45CD7ADC66DF13A5A1ACD7FF6ABB8C" country="Indonesia" latitude="-2.374889" longLatPrecision="1" longitude="138.73886" municipality="Foja Mts" name="Waaf Village" stateProvince="Papua Barat">Waaf Village</location>
,
<geoCoordinate id="29D8F2F585870547D1EAA839D63FDF5D" degrees="02" direction="south" minutes="22" orientation="latitude" precision="1" seconds="29.6" value="-2.374889">02°22'29.6&quot;S</geoCoordinate>
,
<geoCoordinate id="5974BEF9A9690DD072B2D02BF9F0774E" degrees="138" direction="east" minutes="44" orientation="longitude" precision="1" seconds="19.9" value="138.73886">138°44'19.9&quot;E</geoCoordinate>
,
<elevation id="AF87F97AAC22274D9D4C244DB76C2301" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<quantity id="DEB337DC4DB026211097F383A6F01023" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A777F4F8DF594A4EFC39B91B4A3169C3" pageId="0" pageNumber="45" type="type material">
<paragraph id="34D36161828B8608B84E094B8EAD1C11" pageId="0" pageNumber="45">
<typeStatus id="AD6FEEBE0A381C7801240A43BCB473E8">Type</typeStatus>
material.
</paragraph>
<paragraph id="D586B0EE7A2A59C1FBB57B42545C4749" pageId="0" pageNumber="45">
<emphasis id="ACD9D0DF55654A9693837FF80BE0643A" bold="true" pageId="0" pageNumber="45">
<emphasis id="F472B4AD656320A2A1966245657C4983" italics="true" pageId="0" pageNumber="45">
<typeStatus id="51A7500E79D571DF705D95CA5BCBDAF6">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation id="A3B8698F5F9D8FD47A5D6D92899AF3FE" ID-GBIF-Occurrence="3062138323" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="115" latitude="-2.374874" location="Foja Mountains" longLatPrecision="1" longitude="138.73886" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot;
<collectingCountry id="EC16E99BAAF149CC51EE32C67EDD7F29" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="C124AB0147151D6EEC49D72C26C47859" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location id="C0881F0394BFE40C6C002DF504C36C7D" LSID="urn:lsid:plazi:treatment:3F398A2222BB50968728D1949338C2C4:C0881F0394BFE40C6C002DF504C36C7D" country="Indonesia" latitude="-2.374874" longLatPrecision="1" longitude="138.73886" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
N foot, N Waaf vill,
<quantity id="60E2A5ABB7F8FCD1E29A8A18B0E96688" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<elevation id="208F2C93A2F6B8D618B1F602F1884206" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115m</elevation>
</quantity>
,
<date id="FF41F71D9371E6B0FDE1F5BF71530D1B" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="5F4D471DBBA63215DBACA7C4C8CFB627" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate id="94F3FBC96F3CD12ADC119571EB424279" degrees="2.374874" direction="south" orientation="latitude" precision="1" value="-2.374874">-2.374874</geoCoordinate>
<geoCoordinate id="F0132449DF76ADF6A78F3D0FBC3B2BE0" degrees="138.738855" direction="east" orientation="longitude" precision="1" value="138.73886">138.738855</geoCoordinate>
, Sumoked (Pap060)&quot; (
<collectionCode id="38FA1DD527AFC0B75D2E3312B2C1DDAF" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3D44018CAF0C909821AF12EAF0801958" pageId="0" pageNumber="45">
<materialsCitation id="3DE9CA449B84E58700EFD9DC79F591F8" ID-GBIF-Occurrence="3062138316" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="9" specimenCount-female="2" specimenCount-male="7" typeStatus="paratype">
<emphasis id="ABA3E82D1D3B482292B661129833238C" bold="true" pageId="0" pageNumber="45">
<emphasis id="F1AA18576A9B7B47F45203582FC8CA3E" italics="true" pageId="0" pageNumber="45">
<typeStatus id="A558B2A8272E31D2FC9C69083254E15C">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount id="2F873021568A85192BCD15F6B59FDBC9" type="male">7 males</specimenCount>
,
<specimenCount id="61398AE893A38EA63C4C7DDB4F42D32E" type="female">2 females</specimenCount>
with the same label as the
<typeStatus id="5439FB99B95CEB5ED27EAE213C4890A0">holotype</typeStatus>
, one male with an additional green text label
<normalizedToken id="B797632F00A6B858D6E3FCB1BD12BF53" originalValue="“7281”">&quot;7281&quot;</normalizedToken>
(
<collectionCode id="D63F5BCF23BA96857607C8D3383EDFD9" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode id="98013826AC750B62F1A0BE397A036C70" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="478A576674D6289DE833D435B326F3E6" pageId="0" pageNumber="45" type="description">
<paragraph id="A8AECEF187E4D1E7A84A4004DA80EF19" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="D559D0E633140F02FDDA1756DDFC06D1" pageId="0" pageNumber="45">
<emphasis id="83BD22F395FC2CE65D5488D82017E471" bold="true" pageId="0" pageNumber="45">
<emphasis id="0039A9888675C04D51ECF7D2CFE96490" italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small to medium-sized: TL-H
<quantity id="A680908C1670AF6A7DEA11F5F60D8348" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.75" metricValueMin="3.55" unit="mm" value="3.65" valueMax="3.75" valueMin="3.55">3.55-3.75 mm</quantity>
, TL
<quantity id="39D08687CEA741A5E5E5F4EB59F383BA" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.9-4.2 mm</quantity>
, MW 1.95-2.0 mm (
<typeStatus id="F4AB9B5419F715BE3787E62694FCDA98">holotype</typeStatus>
: TL-H
<quantity id="74A39640F25D55B384E395676A3D1A0F" metricMagnitude="-3" metricUnit="m" metricValue="3.65" unit="mm" value="3.65">3.65 mm</quantity>
, TL
<quantity id="7FF06EBDABFF5465B6EFD2BC00C42D27" metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
,
<collectionCode id="0B69FA0334259B5F9DBC997F29A399D0" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity id="E2B52E7E2913086F235BF06736A24F59" metricMagnitude="-3" metricUnit="m" metricValue="1.95" unit="mm" value="1.95">1.95 mm</quantity>
), with oblong-oval habitus (Fig.
<figureCitation id="E4EB2E4032F4C8A2101B3B86BAE64CF4" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">2</figureCitation>
).
</paragraph>
<paragraph id="C0E108822EFD1D3FFA7B52FE33D30342" pageId="0" pageNumber="45">
<emphasis id="7F75A289297C643820BAC0112AECFD55" bold="true" pageId="0" pageNumber="45">
<emphasis id="20A93047E729BF17448A8BAA518B2544" italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally piceous, with paler lateral sides of pronotum (Fig.
<figureCitation id="DB9770C37D6BAF66BCC56C9169BED478" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">2</figureCitation>
). Head piceous, with slightly paler, dark brown, anterior margin; pronotum piceous, slightly paler towards lateral sides, lateral sides brown to dark brown, yellowish red to reddish brown at anterior angles; elytra uniformly piceous or with dark brown sutural lines; head appendages and proximal part of legs yellowish brown, legs distally brown.
</paragraph>
<paragraph id="144DFDDA344271DBA338A95AB0A65A6D" pageId="0" pageNumber="45">
<emphasis id="A695819BE82B347F15CB151FA88C3CF6" bold="true" pageId="0" pageNumber="45">
<emphasis id="E78B23943F20193AF7E8F63465FB811B" italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with inconspicuous, almost invisible elytral punctation and weakly impressed microreticulation. Head with uneven, sparse punctation (spaces between punctures 1-4 times size of punctures); diameter of punctures smaller than or almost equal to diameter of cells of microreticulation; punctation sparser and finer anteriorly and posteriorly. Pronotum with distinctly finer, sparser, and more evenly distributed punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Elytra and pronotum with weakly impressed microreticulation; head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and fine punctation.
</paragraph>
<paragraph id="6812A3D08607EB798D6BBCFFE95754F4" pageId="0" pageNumber="45">
<emphasis id="F270E45D9648CD007C74309BFCC46411" bold="true" pageId="0" pageNumber="45">
<emphasis id="75AC158EB55CE3043FE31DF78442F94D" italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, narrow, convex, with distinct lateral bead and few setae. Abdominal ventrite 6 concave apically.
</paragraph>
<paragraph id="917A5461FA753A1ED85EA58B2C65D22C" pageId="0" pageNumber="45">
<emphasis id="3DD14DB468B3ACC29623FAE4EA083111" bold="true" pageId="0" pageNumber="45">
<emphasis id="99706430967995432A09C89ACBD74CF1" italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple (Fig.
<figureCitation id="42E9EA146CBE798BC86C7273349EB659" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">2</figureCitation>
). Pro- and mesotarsomeres 1-3 not dilated, relatively narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, long, relatively slender, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 14 and posterior row of five short setae (Fig.
<figureCitation id="C00B3B534394AC110CF41EB9B3B4AAE3" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">6A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with large, evenly tapering and curved downwards apex; in ventral view, with weak submedian constriction, distal part distinctly narrower than proximal one, apex truncate (Fig.
<figureCitation id="5FB2E7A0632F12C37A68C71F56EE996C" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">6B, C</figureCitation>
). Paramere with strong notch on dorsal side and subdistal part subquadrate, large and broad; subdistal setae long, dense, thick, and flattened; proximal setae hair-like, numerous, more inconspicuous than subdistal ones (Fig.
<figureCitation id="B257A75DBB1DE1DAE5FA90A47FEC4AB7" captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">6D</figureCitation>
). Abdominal ventrite 6 distinctly concave, with 9-11 lateral striae on each side.
</paragraph>
<paragraph id="2DED899573851DCFD9CE5158FCBC3B3C" pageId="0" pageNumber="45">
<emphasis id="D1E1A466D69D24FEFE957A016216B45A" bold="true" pageId="0" pageNumber="45">
<emphasis id="A84B948FF2AC5691E88890E36BB01C31" italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 slightly concave, without lateral striae.
</paragraph>
</subSubSection>
<subSubSection id="D1A543FE3F4329D438250A427570C4C7" pageId="0" pageNumber="45" type="affinities">
<paragraph id="609BC299828AAE01DB5E9268F6803C50" pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph id="ABC077992DC57829603DCC28457AC1B6" pageId="0" pageNumber="45">
The new species evidently belongs to the
<taxonomicName id="A8E583641B9E18576EF369077837F21C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis id="E34195306A7079D2B61D1E3ED34109B1" italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. The species is very similar to
<taxonomicName id="69398641286C05D8590E8F56E1796C22" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. utowaensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="utowaensis">
<emphasis id="F794F8D7837CFAB41D7A21A64FA32E04" italics="true" pageId="0" pageNumber="45">E. utowaensis</emphasis>
</taxonomicName>
<bibRefCitation id="A2F70B5C65280CE69C1C5C836AA65F99" DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">
Shaverdo et. al.,
<quantity id="B8E4FE6AB538805F62446CA6461CD804" metricMagnitude="1" metricUnit="m" metricValue="5.11048" unit="in" value="2012.0">2012 in</quantity>
</bibRefCitation>
general appearance, apically concave abdominal ventrite 6, and structure of the male genitalia, but differs from in more slender male antennae and shape of the median lobe and paramere.
</paragraph>
</subSubSection>
<subSubSection id="D6F2C5D5D8529657C00F382EDED73691" pageId="0" pageNumber="45" type="distribution">
<paragraph id="10A873B3D0EEC49DA35E4D6179F05AEA" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="8F7BC412154D68354290B636A431198B" pageId="0" pageNumber="45">
<materialsCitation id="3E4BD5865335310186E442AF8EF91F41" ID-GBIF-Occurrence="3062138341" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry id="3A1B6D94DCF9D8AF28445DC785BA21FE" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="6A91F31D881B1FA58850AB9F061840A8" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
.
<collectingMunicipality id="90BB724323A17790A2DAA5F0E9E8BB0F">This</collectingMunicipality>
species is known only from the
<typeStatus id="8E6D0BB31204FFC2F0F63BB0F370124E">type</typeStatus>
locality in
<location id="5E6F526B9E66800AC39B1CAC4BA92CFE" LSID="urn:lsid:plazi:treatment:3F398A2222BB50968728D1949338C2C4:5E6F526B9E66800AC39B1CAC4BA92CFE" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
(Fig.
<figureCitation id="461C000DA6C9D797F222CEB5AFAE62D9" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E2072160C44FC952434CDF2625DBC42" pageId="0" pageNumber="45" type="habitat">
<paragraph id="66D04C2BDCE097E39BCA674C641FBA8A" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="29024741A60ED45582A1AAD4FC09FBFA" pageId="0" pageNumber="45">The specimens were collected from shaded waterholes on a riverbank.</paragraph>
</subSubSection>
<subSubSection id="420B5998CA4F386CC2BA7324EE72A2BB" pageId="0" pageNumber="45" type="etymology">
<paragraph id="F8202CDA80616250390D6E363B84410F" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="72B5703DAC96BF57D32E3602F457022C" pageId="0" pageNumber="45">The species is named after Waaf Village. The name is a noun in the nominative singular standing in apposition.</paragraph>
</subSubSection>
</treatment>
</document>
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