121 lines
12 KiB
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121 lines
12 KiB
XML
<document id="7F1BF20B85CE3DA6E768C074EA3F69AB" ID-CLB-Dataset="30184" ID-DOI="http://dx.doi.org/10.3897/zookeys.753.22366" ID-GBIF-Dataset="fe320c78-20cd-45de-babe-5c2fd18c7a1d" ID-PMC="PMC5934354" ID-Pensoft-Pub="1313-2970-753-23" ID-PubMed="29731681" ID-ZooBank="BBC59C6140DD44F3B9F34C011E0D0B75" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-753-23" ModsDocOrigin="ZooKeys 753" ModsDocTitle="Taxonomic revision of the Graphipterusserrator (Forskål) group (Coleoptera, Carabidae): an increase from five to 15 valid species" checkinTime="1524810864104" checkinUser="pensoft" docAuthor="Renan, Ittai, Assmann, Thorsten & Freidberg, Amnon" docDate="2018" docId="3F331E47F3E3F0B23160994812E2318B" docLanguage="en" docName="ZooKeys 753: 23-82" docOrigin="ZooKeys 753" docSource="http://dx.doi.org/10.3897/zookeys.753.22366" docTitle="Graphipterus barthelemyi Dejean 1830" docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="123E1514FFF23830FFA1FFB0FFE0DA05" masterDocTitle="Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species" masterLastPageNumber="82" masterPageNumber="23" pageNumber="32" updateTime="1701380747961" updateUser="ExternalLinkService">
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<mods:titleInfo id="5AF858E6413453F7D133A05ECCA0B0B1">
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<mods:title id="5865C49470B930C969D75B5258FA2FBA">Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species</mods:title>
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<mods:namePart id="A05D0EE3541EECAFFA3E87B69E655A44">Renan, Ittai</mods:namePart>
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<mods:namePart id="6218F3B1F1ECB0D5087F8075F179F7F7">Assmann, Thorsten</mods:namePart>
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<mods:roleTerm id="A2E9B680962805BFEAE0E14289CEF4AB">Author</mods:roleTerm>
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<mods:namePart id="7695B3E6DF873A04418D1B6709FB155F">Freidberg, Amnon</mods:namePart>
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<mods:typeOfResource id="4430651B80D145CEB0F480ECEB7687C4">text</mods:typeOfResource>
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<mods:title id="5574C7C084B3C863043C5194D9ED8FD7">ZooKeys</mods:title>
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<mods:part id="B77596131C7AF18CDA91EFB2BB4A00FE">
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<mods:date id="2EA09B1DF613CDA1BF5995032BD5A824">2018</mods:date>
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<mods:detail id="E549CDBB0660DEB4D11FFEB3238FF280" type="volume">
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<mods:number id="DE67105E2F5DD5160FD0BF5CDB704ABF">753</mods:number>
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<mods:start id="D919136C929E08AB2AEAF619BECAD0F4">23</mods:start>
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<mods:url id="7C64C34DCA4A781BC7E1186D3940CB8E">http://dx.doi.org/10.3897/zookeys.753.22366</mods:url>
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<mods:classification id="42B3A9A80BEF957FA01664606024A190">journal article</mods:classification>
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<mods:identifier id="2CBB031579C0D406B439F9717D02A39C" type="DOI">http://dx.doi.org/10.3897/zookeys.753.22366</mods:identifier>
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<mods:identifier id="155754EF8124FF7A4F6DF007869FEFE5" type="Pensoft-Pub">1313-2970-753-23</mods:identifier>
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<treatment id="3F331E47F3E3F0B23160994812E2318B" ID-GBIF-Taxon="143706395" LSID="urn:lsid:plazi:treatment:3F331E47F3E3F0B23160994812E2318B" httpUri="http://treatment.plazi.org/id/3F331E47F3E3F0B23160994812E2318B" lastPageNumber="32" pageId="9" pageNumber="32">
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<subSubSection id="F6BD3BE9C650C55EF7E3DAB850E6B30D" pageId="9" pageNumber="32" type="nomenclature">
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<paragraph id="1C860C93327AFBD6BB0C78CABB5CBE6E" pageId="9" pageNumber="32">
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<taxonomicName id="59DD0501C2FE1938C401BB8C2E2A7159" ID-CoL="3H95V" authority="Dejean, 1830" authorityName="Dejean" authorityYear="1830" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus barthelemyi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="barthelemyi">Graphipterus barthelemyi Dejean, 1830</taxonomicName>
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Figs 3a, 6c, 9a, 17, 20
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<normalizedToken id="7BBD64600519DC16CB3FAA2FF482D9AF" originalValue="a–b">a-b</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="D13869A2208AC4BC418E34B1BD4745B2" pageId="9" pageNumber="32" type="types">
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<paragraph id="D08852D23117FD3DD71F8DA061C21174" pageId="9" pageNumber="32">Types.</paragraph>
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<paragraph id="2D4337049E526EA79F6CFEB6A65EA7CA" pageId="9" pageNumber="32">Holotype: ♂ (Blue label, black handwritten): <Barthelemyi. Solier/. in Barbaria. Tunis. D. Barthelemyi>. (White label with brown margin, brown letters, handwritten): <EX Musaeo/Chaudoir>. (Red label, black letters, type written): <TYPE>. Deposited in NHMB, Chaudoir collection [examined].</paragraph>
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</subSubSection>
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<subSubSection id="981C68CE836273D6DE56968BDBCE30E4" pageId="9" pageNumber="32" type="diagnosis">
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<paragraph id="A15A5B95F2AE72E90930CB6B3A26A961" pageId="9" pageNumber="32">Diagnosis.</paragraph>
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<paragraph id="2AC283C53A1954588DA1FA138FB2168B" pageId="9" pageNumber="32">Medium-sized species with grayish or yellowish scales usually cover the elytra and sometimes also on the pronotum. Elytra pattern rarely visible with six lateral margin extensions and 18-24 isolated white circular to elongated spots occur on elytra.</paragraph>
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</subSubSection>
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<subSubSection id="68B0FB62F5B52FA25A3F5F5298C28EB2" pageId="9" pageNumber="32" type="description">
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<paragraph id="A76FA7EA75EEBA8CFDD8E970256E4291" pageId="9" pageNumber="32">Description.</paragraph>
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<paragraph id="49BFDE8E3E2347F672747E73724BF4D0" pageId="9" pageNumber="32">
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BL male: 13.0-17.0 mm, average 15.8
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<normalizedToken id="E6F1FAB24D9898B32BB479C6F3267D7F" originalValue="±">+/-</normalizedToken>
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1.5 mm; BL female: 15.5-17.0 mm, average 16.3
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<normalizedToken id="8816F535BE5C3641777BFD8367655880" originalValue="±">+/-</normalizedToken>
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0.6 mm. Grayish with elytral white blurred spots and extensions.
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</paragraph>
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<paragraph id="B58BC3B8A0B46341E90C943059148C95" pageId="9" pageNumber="32">Head slender: HW/PW: 0.72; EYL: 1.1-1.4 mm; EYL/EL: 0.15. Mentum without teeth (Fig. 3a). Frontal ridge reduced. In male, apical white frons stripes slenderer than exposed frons (cf. Fig. 4a).</paragraph>
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<paragraph id="23DCE2F0BD87554A2AED37373E44B028" pageId="9" pageNumber="32">Pronotum wide; PL/PW: 0.63; BPW/BPW/PW: 0.6; posteromedially concave and with white lateral margin, as wide as antennomere I long: white slushy scales cover disc sometimes.</paragraph>
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<paragraph id="88F9BC0868001FFE65D9DD8EE0AD94AD" pageId="9" pageNumber="32">Elytra wide, elytron margin almost continuously rounded from humeri to posterolateral angles; EL: 7.1-8.8 mm, average 8.2 mm; EW: 6.5-7.8 mm, average 7.3 mm; EL/EW: 1.1. Elytra longitudinally flat, usually with grayish scales, disc visible between scales (Fig. 6c); extensions and lateral margin blurred. Lateral margin nearly as wide as antennomere I long and with six extensions; extension I usually elongated; white posterior margin almost touches suture at apex. Disc with 18-24 rounded, usually elongated spots, anterior pair of spots elongate, as wide as extension I; posterior pair of spots rounded, located toward suture; round spots located posterior to third extensions laterally in imaginary lateral line as posterior spots. Apical sinuation slightly developed to straight, apex not protuberant, broadly rounded, especially medially (Fig. 7c). Suture inconspicuous.</paragraph>
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<paragraph id="09BD42AAB96CBD7B77D2A2426FF96FF4" pageId="9" pageNumber="32">Legs medium; MTIL: 4.5-6.5 mm, average 5.7 mm; El/MTIL: 1.7. Metatibial secondary spur brown. MTAL: 3.5-4.4 mm, average 4 mm; MTAL/MTIL: 0.8. Claws of hind legs brown at base.</paragraph>
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<paragraph id="B3218D2C54DEAFC50FA7C6CC8F273958" pageId="9" pageNumber="32">Median lobe of aedeagus with short unbent tip (Fig. 9a).</paragraph>
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</subSubSection>
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<subSubSection id="68FF2FC4A1842E193722974B27F4A6B8" pageId="9" pageNumber="32" type="comparisons">
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<paragraph id="40FD010980707CC61BFB0E8E343E927D" pageId="9" pageNumber="32">Comparisons.</paragraph>
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<paragraph id="BEB844C742D03CA32C78D6F9E25D37BC" pageId="9" pageNumber="32">
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Distinguished from all other species of the
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<taxonomicName id="A61F7A30E5A1BF19DCBE8534B966F165" lsidName="G. serrator" pageId="9" pageNumber="32" rank="species" species="serrator">G. serrator</taxonomicName>
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group by white lateral margins merged at the posterior margin of the pronotum. Median lobe of aedeagus with short, straight tip.
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</paragraph>
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</subSubSection>
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<subSubSection id="A567D1C5759AD56F658AF3A11AD19D59" pageId="9" pageNumber="32" type="habitat">
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<paragraph id="E2BE57A7E65A1D1559E8866EE2C8475D" pageId="9" pageNumber="32">Habitat.</paragraph>
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<paragraph id="8CA824BC5626BA5BFCB8BD3C53174B9E" pageId="9" pageNumber="32">Unknown. The species was found exclusively in coastal dune habitats.</paragraph>
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</subSubSection>
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<subSubSection id="1BBDC666558A6D0FD2B0E79634B945AA" pageId="9" pageNumber="32" type="co-occurring species">
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<paragraph id="5B0D09C57F5735F7A365316F4A19C575" pageId="9" pageNumber="32">Co-occurring species.</paragraph>
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<paragraph id="6019788393FAA376783FC07BA6FD7B2B" pageId="9" pageNumber="32">
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<taxonomicName id="B607DDF14736D7CB8123105015633B1D" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Graphipterus barthelemyi" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="barthelemyi">Graphipterus barthelemyi</taxonomicName>
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lives in sympatry with
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<taxonomicName id="C969490ABA44012009A823927FB1A94C" lsidName="G. luctuosus" pageId="9" pageNumber="32" rank="species" species="luctuosus">G. luctuosus</taxonomicName>
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in Tunisia.
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</paragraph>
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</subSubSection>
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<subSubSection id="5298CF1468B1A87DBE7BA398DCCE8CCC" pageId="9" pageNumber="32" type="distribution">
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<paragraph id="CF462210D90F815BFA0D956751CE3A06" pageId="9" pageNumber="32">Distribution.</paragraph>
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<paragraph id="B756DC92B4095A91931B219F3A643DEE" pageId="9" pageNumber="32">Restricted to north-east Tunisia (Fig. 17).</paragraph>
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</subSubSection>
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<subSubSection id="CB3F506296B0E0F22D7C4DC567B0A93D" pageId="9" pageNumber="32" type="conservation">
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<paragraph id="3D123A2A7DF95E51A7C33A67A61CB010" pageId="9" pageNumber="32">Conservation.</paragraph>
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<paragraph id="B8E75C3C89841315C8A02C663AB490AD" pageId="9" pageNumber="32">The restricted distribution range of the endemic species and the decline of the coastal sandy habitat as a result of increasing anthropogenic pressures (e.g., tourism activities, urbanization, etc.) threaten at least the long-term survival of the species.</paragraph>
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</subSubSection>
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<subSubSection id="9C7B7D982CCA65634DFEB109A974EE58" pageId="9" pageNumber="32" type="comments">
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<paragraph id="93018A5553C472E332C739EDE987CD3B" pageId="9" pageNumber="32">Comments.</paragraph>
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<paragraph id="0A2BD252400BD210406ECDC66C819C67" pageId="9" pageNumber="32">
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Both
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<bibRefCitation id="3563A34723629020411BAAF19D84FAE4" pageId="9" pageNumber="32">Basilewsky (1977)</bibRefCitation>
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and
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<bibRefCitation id="9438BB1FBCBFB8816798F8E1BD5F24D7" author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="51" pageNumber="74" title="Systematic list of extant ground beetles of the world (Coleoptera " Geadephaga ": Trachypachidae and Carabidae, incl. Paussinae, Cicindelinae, Rhysodinae). Second edition." year="2005">Lorenz (2005)</bibRefCitation>
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note in error that
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<taxonomicName id="31014FAD7CBBF2E9C1ECE55740A035BA" lsidName="G. barthelemyi" pageId="9" pageNumber="32" rank="species" species="barthelemyi">G. barthelemyi</taxonomicName>
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was described by Dejean (1831).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |