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<document id="B397D1CAFB7733BB36AF14A294D41F62" ENCODING="UTF-8" ID-GBIF-Dataset="57481cd6-bd80-45ed-81e5-efa7df94defb" ModsDocID="z01126p001" checkinTime="1247230541569" checkinUser="thomas" docAuthor="H. H. Tan &amp; C. U. M. Leh" docDate="2006" docId="3D58559EE0339DF6DBC0D185440ED220" docLanguage="en" docName="2006_Tan_Leh_gg1.xml" docOrigin="Zootaxa 1126" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:62372BE2-2974-42CF-B88F-0D3CD2B66F6C" docTitle="Gastromyzon crenastus Tan &amp; Leh, 2006, new species" docType="treatment" docVersion="12" lastPageNumber="13" masterDocId="006D0B716E91CA4AD4674B54E7B68FA7" masterDocTitle="Three new species of Gastromyzon (Teleostei: Balitoridae) from southern Sarawak." masterLastPageNumber="19" masterPageNumber="1" pageNumber="9" updateTime="1701314298820" updateUser="plazi">
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<mods:title id="6878BE41637856797E4B3DCAC4A392F3">Three new species of Gastromyzon (Teleostei: Balitoridae) from southern Sarawak.</mods:title>
</mods:titleInfo>
<mods:name id="3E7CD42D41CDADECDD7690C201DDF1DB" type="personal">
<mods:role id="F43D190C84EE434BF9E365D0F705BC4E">
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</mods:role>
<mods:namePart id="C3F422CC02A173935BBD5B64F6B022B7">H. H. Tan</mods:namePart>
</mods:name>
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<mods:role id="B6BA156D2C479B7A41D642272B15D057">
<mods:roleTerm id="17FBD9F82A036D87699980E4E32705AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD38CBC8D5FB5C0BEF327C7DF07632A6">C. U. M. Leh</mods:namePart>
</mods:name>
<mods:typeOfResource id="DE0F4A1F04E987519C92245A141CB54A">text</mods:typeOfResource>
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<mods:date id="3A24A9FF9F6F77E752A9DEDF3A1572B1">2006</mods:date>
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<mods:classification id="B32A63BD1437EBC2C88FB30C8CC753FF">journal article</mods:classification>
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<treatment id="3D58559EE0339DF6DBC0D185440ED220" ID-DOI="http://doi.org/10.5281/zenodo.6256591" ID-GBIF-Taxon="100122025" ID-Zenodo-Dep="6256591" LSID="urn:lsid:plazi:treatment:3D58559EE0339DF6DBC0D185440ED220" httpUri="http://treatment.plazi.org/id/3D58559EE0339DF6DBC0D185440ED220" lastPageNumber="13" pageNumber="9">
<subSubSection id="97EDF32CE244811A1BE31C06C6AB9F0D" type="nomenclature">
<paragraph id="350F1C5F95D24EEFB773B4EA2B5EF4F2" pageNumber="9">
<taxonomicName id="732DD32FBB919D363644D46FD78DF0E9" ID-CoL="3FDR6" LSID-ZBK="urn:lsid:zoobank.org:act:896D0D5B-0278-4732-B2AE-55DA94B684AA" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon crenastus Tan &amp; Leh 2006" order="Cypriniformes" pageNumber="9" phylum="Chordata" rank="species" species="crenastus">Gastromyzon crenastus</taxonomicName>
,
<taxonomicNameLabel id="59FE41AF4D3D4F31483347185CE6229D" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BA609FCC27CF36FA316BCEE38C95A6B6" type="description">
<paragraph id="4A51453CDEFAF249214883370A58F739" pageNumber="9">Figs. 3-4</paragraph>
</subSubSection>
<subSubSection id="97E6E223D03B3DA00787E43499F928EA" type="nomenclature">
<paragraph id="7EB9ED0A01FA894F85A59895B4E3FC54" pageNumber="9">
Material examined:
<collectingRegion id="02F83FDE191C11F0250D9EA375A92679">BORNEO: SARAWAK</collectingRegion>
(Sadong River basin):
</paragraph>
<paragraph id="117D2E2F5D19C1E796C6849C72F4628D" pageNumber="9">
<materialsCitation id="2046937394ADBB4A7756A97A6F191E7A" ID-GBIF-Occurrence="923893633" country="Malaysia" stateProvince="Sarawak">
<typeStatus id="A28A1477AA4D6AA2835ACA69004248A0">HOLOTYPE</typeStatus>
:
<collectionCode id="2D412DC1BF683739121ACC6702176F0F">SM</collectionCode>
uncatalogued, 28.1 mm SL; Serian, Sungai Kuhas, 6.9 km from Tebelu Tebakang turnoff, 5.8 km inside right side road (
<geoCoordinate id="FAE84E1F12793504014590DA0C4CCBD6" direction="north" orientation="latitude" value="1.1527778">01º0910.0”N</geoCoordinate>
<geoCoordinate id="108D1903A9645E4594B7C88826E60B84" direction="east" orientation="longitude" value="110.48963">110º2922.7”E</geoCoordinate>
); H. H. Tan et al., 5 Sept. 1995.
</materialsCitation>
</paragraph>
<paragraph id="1CE35D868BB6B1854F9F39EB968D62DF" pageNumber="9">
<materialsCitation id="7672A4AC1F31D126A473B046B1496ABF" ID-GBIF-Occurrence="923893639" country="Malaysia" stateProvince="Sarawak">
<typeStatus id="B3F440CE9E5363974E6735AD38A370B2">PARATYPES</typeStatus>
:
<collectionCode id="87184AD61DE8A2BF0EFA245067193EA5">SM</collectionCode>
uncatalogued, 9 ex., 16.9-24.7 mm SL
</materialsCitation>
;
<materialsCitation id="278320AD19510481A440BD80925D10F8" ID-GBIF-Occurrence="923893646" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="30600DBEBCCA17EE7D980DC8D335FCE6" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
39416, 10 ex., 16.7-28.4 mm SL
</materialsCitation>
;
<materialsCitation id="8C7A3E7A42E086BA6CB52E9B7E2B22CC" ID-GBIF-Occurrence="923893642" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="1750C66324EE46A7A2DE56A18840C031" collectionName="CMK">CMK</collectionCode>
11958, 4 ex., 25.1-28.0 mm SL; preceding lots collected with holotype
</materialsCitation>
. The following
<typeStatus id="841A338804F9B551536C44A3D826BD6F">paratype</typeStatus>
lots are also from the type locality, but with different collectors and dates of collection, as indicated.
<materialsCitation id="FA40C6071DC113713067B9180B647583" ID-GBIF-Occurrence="923893640" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="3CE9EAB48FCF4CDABF4D8CA2A1E64EA5" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
39831, 7 ex., 20.3-29.7 mm SL; H. H Tan et al.,
<date id="FEE306BC8DC26753FC8991D88A0158D0" day="14" month="01" value="1996-01-14" year="1996">14 Jan. 1996</date>
</materialsCitation>
.
<materialsCitation id="D8B4A5BD489D1FB3DCA262A987F20675" ID-GBIF-Occurrence="923893648" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="9B06E63CE1C8492B2996EAC637724F5E" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47075, 6 ex., 18.9-22.4 mm SL; H. H. Tan et al.,
<date id="6A39274B236CCCFEB7D6A5C3CA0D8E82" day="19" month="02" value="1997-02-19" year="1997">19 Feb. 1997</date>
</materialsCitation>
.
<materialsCitation id="0FFAAC603C184FB4FFC7E5E95F60BFCB" ID-GBIF-Occurrence="923893644" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="2FF72C67FAC3C0B968DFB8C2141F02DB" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47076, 24 ex., 18.2-26.8 mm SL; Honours 98/99 Fish Group,
<date id="4CA98103481E5978D3583E5424ED87B4" day="23" month="06" value="1998-06-23" year="1998">23 June 1998</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9C5E5D2AB25CCF7E6FDF1B1E2BFC593" type="materials_examined">
<paragraph id="38A1880EEC5DDE83AED7AA68E5EBFE32" pageNumber="10">
NON-TYPE MATERIAL:
<materialsCitation id="C928EB03DA3AD0E42D71880FC2144A66" ID-GBIF-Occurrence="923893662" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="551BA10F5F86C87821465475710742A3" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47084, 1 ex., 29.0 mm SL;
<collectingRegion id="6FBA9D829E19B7AD283C3C6E18B7EC4C">Sarawak</collectingRegion>
: Tebedu, Sungai Ahi, on road towards Mongkos (
<geoCoordinate id="EEE5E084445ADADC9104275915C74DF6" direction="north" orientation="latitude" value="0.92399997">00°55.44'N</geoCoordinate>
<geoCoordinate id="B74A9B4667F0EA9E45291ACAF386E25D" direction="east" orientation="longitude" value="110.539">110°32.34'E</geoCoordinate>
); H. H. Tan et al.,
<date id="12A9F986F015A0975B3ABAFDA6CA09CD" day="12" month="06" value="1999-06-12" year="1999">12 June 1999</date>
</materialsCitation>
. The following non-type lots are all from the type locality, but with different collectors and dates of collection, as indicated.
<materialsCitation id="511F7E3768304EBEC28A11006792E881" ID-GBIF-Occurrence="923893653" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="A2912865F7C60E11957A6B928689D6F1" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
41213, 16 ex., 18.0-22.2 mm SL; H. H Tan et al.,
<date id="5521D54E7CC649C3DF606FABDB55C91C" day="19" month="02" value="1997-02-19" year="1997">19 Feb. 1997</date>
</materialsCitation>
.
<materialsCitation id="E5C394A70520ABD8E2FDADCBA8B851BD" ID-GBIF-Occurrence="923893650" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="8FD7E41E24B6B03A2E8C50E8138F267C" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47077, 2 ex., 19.8-27.8 mm SL; H. H. Tan et al,
<date id="48E2070FE49D63410A5DBC2489994A0D" day="29" month="10" value="1997-10-29" year="1997">29 Oct. 1997</date>
</materialsCitation>
.
<materialsCitation id="555CEF6248B510AE0F0E2C25B51D4143" ID-GBIF-Occurrence="923893663" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="E3846CADA893E819F04EB12FDA5AF821" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47078, 5 ex., 21.5-24.3 mm SL (site 1, sample 2)
</materialsCitation>
;
<materialsCitation id="595E968058C0DBD6C93C16E9BBD63BCE" ID-GBIF-Occurrence="923893660" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="42ADA6E6614C3F4B77578EDC6272EE8D" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47079, 6 ex., 18.1-26.1 mm SL (site 1, sample 3); Honours 98/99 Fish Group,
<date id="0C15BA1A3EB8E9BCD0FE329F66625AA0" day="24" month="06" value="1998-06-24" year="1998">24 June 1998</date>
</materialsCitation>
.
<materialsCitation id="F8472878EA29E28BC48FB2C295F24921" ID-GBIF-Occurrence="923893659" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="FC406F94A5FEDBE9AD09C7601719C1CF" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47080, 2 ex., 22.7-25.4 mm SL (site 2, sample 2)
</materialsCitation>
;
<materialsCitation id="6FA0DF8F4BB101006954376D60EA98D1" ID-GBIF-Occurrence="923893664" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="BE18819E180ADC8D03138BB9A9D52F69" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47081, 2 ex., 17.3-23.4 mm SL (site 2, sample 3); Honours 98/99 Fish Group,
<date id="86E2ECC432B25CF0F209C2A3A7BA0CA1" day="25" month="06" value="1998-06-25" year="1998">25 June 1998</date>
</materialsCitation>
.
<materialsCitation id="D754FDC06E073AAC4916997EBED332E4" ID-GBIF-Occurrence="923893654" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="6D8AA72FFC59466559CA40B2FC715791" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47082, 2 ex., 16.6-16.9 mm SL (site 3, sample 2); Honours 98/99 Fish Group,
<date id="301B4EE3176A63459B838F897A8C6182" day="26" month="06" value="1998-06-26" year="1998">26 June 1998</date>
</materialsCitation>
.
<materialsCitation id="47E86D6DD74C32DEE32B0CB80926D382" ID-GBIF-Occurrence="923893668" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="890064DBB58FB77E387E608E3A8A07BE" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47083, 14 ex., 14.5-24.0 mm SL; H. H. Tan et al.,
<date id="1794C3AA79C471F3D5F005ED2F516123" day="10" month="06" value="1999-06-10" year="1999">10 June 1999</date>
</materialsCitation>
.
<materialsCitation id="760068EF8C54BFCB201A4EDEBF48D7B7" ID-GBIF-Occurrence="923893674" country="Malaysia" stateProvince="Sarawak">
<collectionCode id="68EB3EAEECA87F01A1C711EEDC2484DD" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
47188, 9 ex., 21.4-25.3 mm SL; native collectors,
<date id="5E777F6947AE3C1E598F95334B091498" day="22" month="06" value="2002-06-22" year="2002">22 June 2002</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8EF5FE85C54840093DAB59AEC08AFA17" type="diagnosis">
<paragraph id="C92168C7E53647013CD7FFE55CFBB6D5" pageNumber="10">
Diagnosis.-
<taxonomicName id="6006AD5A55854A7C2416D3526E96E36D" LSID-ZBK="urn:lsid:zoobank.org:act:896D0D5B-0278-4732-B2AE-55DA94B684AA" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon crenastus Tan &amp; Leh 2006" order="Cypriniformes" pageNumber="10" phylum="Chordata" rank="species" species="crenastus">Gastromyzon crenastus</taxonomicName>
differs from its congeners in having the following unique combination of characters: a black body; dorsum with 6-8 thin cream bars or spots; head black, with cream spots and blotches; sublacrymal groove visible from side; absence of a secondary rostrum; absence of a postoral pouch; absence of a subopercular groove; gill slit vertical; snout truncate when viewed dorsally; abdomen without scales; 60-65 scales in lateral line; pelvic fin not overlapping anal fin origin, adpressed dorsal fin not at level overlapping anal fin origin. Maximum size: 29.7 mm SL (ZRC 39831), the smallest known species of
<taxonomicName id="7DCA223C1AB01C9241358D94CFF7748A" LSID-ZBK="urn:lsid:zoobank.org:act:D6E2004D-AAC6-4A97-9D32-DC5186F09138" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon Günther 1874:454" order="Cypriniformes" pageNumber="10" phylum="Chordata" rank="genus">Gastromyzon</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DD99BBD74017CCAC83BF819B0F548372" type="description">
<paragraph id="56742F33AC1BDC4BC231574BF9A5F9DC" pageNumber="10">Description. General body shape and appearance as in Figs. 3-4. Meristic and morphometric data appear in Table 1. Head truncate in dorsal profile, relatively short (27.5-29.8 % SL) and relatively wide (21.3-24.2 % SL, 74.3-83.3 % HL); head relatively flattened (head depth 12.8-13.9 % SL, 44.3-47.6 % HL); tubercles concentrated at anterior part of snout; snout relatively long (snout length 54.1-58.2 % HL); sublacrymal groove present, distinct when viewed from side; gill slit straight and vertical; length of gill slit about same as eye diameter; subopercular groove absent; postoral pouch absent; belly scales absent; pectoral fin overlapping anterior part of pelvic fin; pelvic fin not overlapping anal fin origin; anteriormost pectoral and pelvic-fin rays with dorsal serrae; dorsal fin situated at about mid body (predorsal length 55.2-59.2 % SL), adpressed dorsal fin not extending past level of anal-fin origin; deepest part of body at dorsal-fin origin (body depth at dorsal-fin origin 16.2-18.5 % SL); anus situated just beyond posterior base of fused pelvic fins; caudal peduncle relatively deep (9.5-10.7 % SL) and relatively short (8.5-10.8 % SL).</paragraph>
<paragraph id="FE183C5879966B3D05D141E081A22FC3" lastPageNumber="11" pageNumber="10">Pigmentation and life coloration.-See Fig. 3. Body black on dorsum and sides; dorsum with 6-8 thin cream bars or spots; side with bars interrupted to form spots and blotches; ventrum cream. Head dorsum black, with cream spots and blotches. Eye with golden iris. Dorsal fin light brown, with 1 thick black bar, hyaline interradial membrane and margin; antero-basal black spot present on dorsal fin. Caudal fin base black, fin rays light brown with subdistal part a large black blotch, area just before blotch with iridescent blue, upper border with red, hyaline interradial membrane and thick margin. Anal-fin rays brown, hyaline edge. Pectoral and pelvic fins brown with 1 black bar. Pelvic axillary flap black with distal cream margin.</paragraph>
<paragraph id="10F895E3D39F204BAA10C5F89F94E031" pageNumber="11">Color in alcohol.-See Fig. 4. Body black on dorsum and sides, dorsum with 6-8 thin cream bars or spots, side with bars interrupted to form spots and blotches, ventrum cream. Head dorsum black with cream spots and blotches. Dorsal fin rays light brown with 1 thick black bar, fin with hyaline interradial membrane and margin, antero-basal black spot present. Caudal fin base black, anterior part with thin crescentic black band, centre of fin with large black blotch and with thin hyaline margin, hyaline interradial membrane and margin. Anal fin black, with hyaline margin. Pectoral and pelvic fins grey with 1 black bar. Pelvic axillary flap black, with distal cream margin. Juveniles with 5-6 cream spots and blotches on black body, head plain, caudal fin base black, with faint central black bar.</paragraph>
</subSubSection>
<subSubSection id="0C4B1AD390A6D3575C80645B06F819EF" type="discussion">
<paragraph id="A72C578109DF3DEB4A5FBE70CEF8113B" pageNumber="12">
Remarks.-
<taxonomicName id="97D482DBEC0A374908BC096F6ABCF762" LSID-ZBK="urn:lsid:zoobank.org:act:896D0D5B-0278-4732-B2AE-55DA94B684AA" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon crenastus Tan &amp; Leh 2006" order="Cypriniformes" pageNumber="12" phylum="Chordata" rank="species" species="crenastus">Gastromyzon crenastus</taxonomicName>
can be further differentiated from other congeners of the
<taxonomicName id="51DF6CC3FE8940E00425E5398E843D5C" LSID-ZBK="urn:lsid:zoobank.org:act:AAFF45C0-3FCB-4745-B516-B7E6595AA76E" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ridens Roberts 1982:515" order="Cypriniformes" pageNumber="12" phylum="Chordata" rank="species" species="ridens">G. ridens group</taxonomicName>
by the following characters: snout truncate in dorsal profile vs. rounded in
<taxonomicName id="2AAB12AE1DC7683C84F2EBF4DEA3E651" LSID-ZBK="urn:lsid:zoobank.org:act:AAFF45C0-3FCB-4745-B516-B7E6595AA76E" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ridens Roberts 1982:515" order="Cypriniformes" pageNumber="12" phylum="Chordata" rank="species" species="ridens">G. ridens</taxonomicName>
; adpressed dorsal fin not overlapping level of anal-fin origin vs. overlapping in
<taxonomicName id="90CECF70E5F9C36C95218F508543CF16" LSID-ZBK="urn:lsid:zoobank.org:act:AAFF45C0-3FCB-4745-B516-B7E6595AA76E" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ridens Roberts 1982:515" order="Cypriniformes" pageNumber="12" phylum="Chordata" rank="species" species="ridens">G. ridens</taxonomicName>
; pelvic fin not overlapping anal fin origin vs. overlapping in
<taxonomicName id="43945B3ECB340CD554C418820C060FA2" LSID-ZBK="urn:lsid:zoobank.org:act:AAFF45C0-3FCB-4745-B516-B7E6595AA76E" family="Balitoridae" genus="Gastromyzon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gastromyzon ridens Roberts 1982:515" order="Cypriniformes" pageNumber="12" phylum="Chordata" rank="species" species="ridens">G. ridens</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B7A210E79BCE23838A05DFF61673DB6F" type="distribution">
<paragraph id="159B2D3CAA1D74E379BF11B6DA3E85D7" pageNumber="12">Distribution.-This species is known only from the hill streams feeding the Sadong River basin in southern Sarawak (Fig. 7).</paragraph>
</subSubSection>
<subSubSection id="7FE9FDAC1A8494F955AC937B725E8F5E" type="etymology">
<paragraph id="8C8D331E12D78A08CD1DE18269FCE0F7" pageNumber="13">Etymology.-The species name is from the Latin crena, meaning notch. This is in reference to the white blotches and bars on the dark coloured body. Used as an adjective.</paragraph>
</subSubSection>
</treatment>
</document>