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<mods:title id="F0812DE8BD260B80C8E9934C368BBB20">Review of some little-known benthic hydroids (Cnidaria, Hydrozoa) from the Southern Ocean</mods:title>
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<taxonomicName id="72B94D1FFF93B44210905850FD9E3A85" ID-CoL="76RNY" authority="Jaderholm, 1905" authorityName="Jaderholm" authorityYear="1905" box="[151,591,1390,1416]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="9" pageNumber="378" phylum="Cnidaria" rank="species" species="clavata">
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<emphasis id="87CDEA8EFF93B44210905850FEB53A85" bold="true" box="[151,356,1390,1416]" italics="true" pageId="9" pageNumber="378">Perarella clavata</emphasis>
(
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)
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<paragraph id="B506369CFF93B442109058AFFEF13AA7" blockId="9.[151,591,1390,1450]" box="[151,288,1425,1450]" pageId="9" pageNumber="378">
(
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AC)
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<taxonomicName id="72B94D1FFF93B442109058ECFDFC3AE5" authority="Jaderholm, 1905: 5" authorityName="Jaderholm" authorityPageNumber="5" authorityYear="1905" box="[151,557,1490,1512]" class="Hydrozoa" family="Hydractiniidae" genus="Hydractinia" kingdom="Animalia" order="Anthoathecata" pageId="9" pageNumber="378" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF93B442109058ECFEB23AE5" box="[151,355,1490,1512]" italics="true" pageId="9" pageNumber="378">Hydractinia clavata</emphasis>
<treatmentCitation id="3418108DFF93B442116D58ECFDFC3AE5" author="Jaderholm" box="[362,557,1490,1512]" page="5" pageId="9" pageNumber="378" year="1905">
<bibRefCitation id="D1284B6DFF93B442116D58ECFDC63AE5" author="Jaderholm" box="[362,535,1490,1512]" pageId="9" pageNumber="378" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm, 1905</bibRefCitation>
: 5
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6, pl. 3 figs 68;
<treatmentCitation id="3418108DFF93B44212EC58ECFC193AE5" author="Stepanjants" box="[747,968,1490,1512]" page="15" pageId="9" pageNumber="378" year="1979">
<bibRefCitation id="D1284B6DFF93B44212EC58ECFC733AE5" author="Stepanjants" box="[747,930,1490,1512]" pageId="9" pageNumber="378" refString="Stepanjants, S. D. (1979) Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expedition, 6. Issledovaniya Fauny Morei, 20 (30), 1 - 200, pls 1 - 25. [in Russian]" type="journal article" year="1979">Stepanjants, 1979</bibRefCitation>
: 15
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, pl. 1 fig. 9;
<treatmentCitation id="3418108DFF93B442144B58ECFAC73AE5" author="Blanco" box="[1100,1302,1490,1512]" page="152" pageId="9" pageNumber="378" year="1994">
<bibRefCitation id="D1284B6DFF93B442144B58ECFB323AE5" author="Blanco" box="[1100,1251,1490,1512]" pageId="9" pageNumber="378" refString="Blanco, O. M. (1994 a) Claves de familias y generos para facilitar el reconocimiento de los Hydroida (Leptolina) Athecata, Thecata y Limnomedusae Argentinos (generacion polipoide exclusivamente). Revista del Museo de La Plata, Zoologia, 14 (160), 147 - 179." type="journal article" year="1994" yearSuffix="a">Blanco, 1994a</bibRefCitation>
: 152
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; 1994b: 184;
<treatmentCitation id="3418108DFF93B44210C058CCFE15390A" author="Pena" box="[199,452,1521,1543]" page="768" pageId="9" pageNumber="378" year="2004">
<bibRefCitation id="D1284B6DFF93B44210C058CCFE43390A" author="Pena" box="[199,402,1521,1543]" pageId="9" pageNumber="378" refString="Pena Cantero, A. L. (2004) How rich is the deep-sea Antarctic benthic hydroid fauna? Polar Biology, 27, 767 - 774. http: // dx. doi. org / 10.1007 / s 00300 - 004 - 0654 - 9" type="journal article" year="2004">Peña Cantero, 2004</bibRefCitation>
: 768
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;
<treatmentCitation id="3418108DFF93B44211C958CFFD15390A" author="Stampar" box="[462,708,1521,1543]" page="58" pageId="9" pageNumber="378" year="2006">
<bibRefCitation id="D1284B6DFF93B44211C958CFFD4F390A" author="Stampar" box="[462,670,1521,1543]" pageId="9" pageNumber="378" refString="Stampar, S. N., Tronolone, V. B. &amp; Morandini, A. C. (2006) Description and life cycle of the hydrozoan Hydractinia uniformis sp. nov. (Cnidaria: Hydrozoa: Hydractiniidae), from the coast of southeastern Brazil. Zootaxa, 1200, 43 - 60." type="journal article" year="2006">
Stampar
<emphasis id="87CDEA8EFF93B442122E58CCFD84390A" box="[553,597,1521,1543]" italics="true" pageId="9" pageNumber="378">et al</emphasis>
., 2006
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: 58
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.
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</paragraph>
<paragraph id="B506369CFF93B44210905B2EFCF6392B" blockId="9.[151,1436,1490,1635]" box="[151,807,1552,1574]" pageId="9" pageNumber="378">
<treatmentCitationGroup id="95A911B2FF93B44210905B2EFCF6392B" box="[151,807,1552,1574]" pageId="9" pageNumber="378">
<taxonomicName id="72B94D1FFF93B44210905B2EFE83392B" box="[151,338,1552,1574]" class="Hydrozoa" family="Cytaeididae" genus="Stylactella" higherTaxonomySource="CoL" kingdom="Animalia" order="Anthoathecata" pageId="9" pageNumber="378" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF93B44210905B2EFE83392B" box="[151,338,1552,1574]" italics="true" pageId="9" pageNumber="378">Stylactella clavata</emphasis>
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<treatmentCitation id="3418108DFF93B442116B5B2EFDE7392B" author="Stechow" box="[364,566,1552,1574]" page="401" pageId="9" pageNumber="378" year="1925">
<bibRefCitation id="D1284B6DFF93B442116B5B2EFDD2392B" author="Stechow" box="[364,515,1552,1574]" pageId="9" pageNumber="378" refString="Stechow, E. (1925) Hydroiden der Deutschen Tiefsee Expedition. Wissenschaftliche Ergebnisse de Deutschen Tiefsee- Expedition auf dem Dampfer ' Valdivia' 1898 - 1899, 17, 383 - 546." type="journal article" year="1925">Stechow, 1925</bibRefCitation>
: 401
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;
<treatmentCitation id="3418108DFF93B44212475B2FFD21392B" author="Iwasa" box="[576,752,1552,1574]" page="262" pageId="9" pageNumber="378" year="1934">
<bibRefCitation id="D1284B6DFF93B44212475B2FFD6F392B" author="Iwasa" box="[576,702,1552,1574]" pageId="9" pageNumber="378" refString="Iwasa, M. (1934) Revision of Stylactis and its allied genera, with description of Stylactella (Stylactis) yerii n. sp. Journal for the Faculty of Science Hokkaido Imperial University, 2, 241 - 277." type="journal article" year="1934">Iwasa, 1934</bibRefCitation>
: 262
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, 269.
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</paragraph>
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<treatmentCitationGroup id="95A911B2FF93B44210905B11FD893949" box="[151,600,1582,1605]" pageId="9" pageNumber="378">
<taxonomicName id="72B94D1FFF93B44210905B11FEE53949" box="[151,308,1582,1604]" class="Hydrozoa" family="Cytaeididae" genus="Cytaeis" kingdom="Animalia" order="Anthoathecata" pageId="9" pageNumber="378" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF93B44210905B11FEE53949" box="[151,308,1582,1604]" italics="true" pageId="9" pageNumber="378">Cytaeis clavata</emphasis>
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<treatmentCitation id="3418108DFF93B44211495B10FD823949" author="Bouillon" box="[334,595,1582,1605]" page="145" pageId="9" pageNumber="378" year="2006">
<bibRefCitation id="D1284B6DFF93B44211495B10FDF03949" author="Bouillon" box="[334,545,1582,1605]" pageId="9" pageNumber="378" refString="Bouillon, J., Gravili, C., Pages, F., Gili, J. M. &amp; Boero, F. (2006) An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle, 194, 1 - 591." type="journal article" year="2006">
Bouillon
<emphasis id="87CDEA8EFF93B44211AB5B0EFE093949" box="[428,472,1582,1605]" italics="true" pageId="9" pageNumber="378">et al</emphasis>
., 2006
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: 145
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.
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<paragraph id="B506369CFF93B44210905B73FCB0396E" blockId="9.[151,1436,1490,1635]" box="[151,865,1613,1635]" pageId="9" pageNumber="378">
<treatmentCitationGroup id="95A911B2FF93B44210905B73FCB0396E" box="[151,865,1613,1635]" pageId="9" pageNumber="378">
<taxonomicName id="72B94D1FFF93B44210905B73FE99396E" ID-CoL="76RNY" box="[151,328,1613,1635]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="9" pageNumber="378" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF93B44210905B73FE99396E" box="[151,328,1613,1635]" italics="true" pageId="9" pageNumber="378">Perarella clavata</emphasis>
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<treatmentCitation id="3418108DFF93B44211645B70FDD9396E" author="Rees" box="[355,520,1613,1635]" page="394" pageId="9" pageNumber="378" year="1962">
<bibRefCitation id="D1284B6DFF93B44211645B70FE07396E" author="Rees" box="[355,470,1613,1635]" pageId="9" pageNumber="378" refString="Rees, W. J. (1962) Hydroids of the family Cytaeidae L. Agassiz, 1862. Bulletin of the British Museum (Natural History), Zoology Series, 8, 381 - 400." type="journal article" year="1962">Rees, 1962</bibRefCitation>
: 394
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, fig. 11;
<treatmentCitation id="3418108DFF93B44212675B70FC8C396E" author="Pena" box="[608,861,1613,1635]" page="453" pageId="9" pageNumber="378" year="2008">
<bibRefCitation id="D1284B6DFF93B44212675B70FCFA396E" author="Pena" box="[608,811,1613,1635]" pageId="9" pageNumber="378" refString="Pena Cantero, A. L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451 - 464. http: // dx. doi. org / 10.1007 / s 00300 - 007 - 0371 - 2" type="journal article" year="2008">Peña Cantero, 2008</bibRefCitation>
: 453
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.
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<subSubSection id="FDA36517FF93B44210905BB1FE2D39FD" pageId="9" pageNumber="378" type="materials_examined">
<paragraph id="B506369CFF93B44210905BB1FE2D39FD" blockId="9.[151,1436,1679,2029]" pageId="9" pageNumber="378">
<emphasis id="87CDEA8EFF93B44210905BB1FE5139A5" bold="true" box="[151,384,1679,1704]" pageId="9" pageNumber="378">Material examined.</emphasis>
<collectingCountry id="CDAE760CFF93B44211815BAEFE3F39AA" box="[390,494,1680,1703]" name="Antarctica" pageId="9" pageNumber="378">
<emphasis id="87CDEA8EFF93B44211815BAEFE3F39AA" box="[390,494,1680,1703]" italics="true" pageId="9" pageNumber="378">Antarctic</emphasis>
</collectingCountry>
Swedish South Polar Expedition.
<date id="C107105CFF93B442136B5BAEFC1C39A5" box="[876,973,1680,1704]" pageId="9" pageNumber="378" value="1901" valueMax="1903">1901-03</date>
:
<typeStatus id="6A02883EFF93B44213D05BAEFBDE39A5" box="[983,1039,1680,1704]" pageId="9" pageNumber="378">Type</typeStatus>
(SMNH—7946), Stn 8,
<date id="C107105CFF93B44215265BAEFA4639A5" box="[1313,1431,1680,1704]" pageId="9" pageNumber="378" value="1902-02-11">11.II.1902</date>
,
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,
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(Erebus and Terror Gulf, Graham Region),
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, soft clay, colony with numerous polyps and gonophores, on gastropod shell.
</paragraph>
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<emphasis id="87CDEA8EFF93B44210C05BC2FE883818" bold="true" box="[199,345,1788,1813]" pageId="9" pageNumber="378">Description.</emphasis>
Stolonal colony growing on gastropod shell. Hydrorhiza consisting of a mesh of anastomosing tubes of perisarc, not covered by a coenosarc layer. With gastrozooids and gonophores. Neither spines nor dactylozooids. Gastrozooids relatively large (
<figureCitation id="2D822A19FF93B44212A85A7AFD293850" box="[687,760,1860,1885]" captionStart="FIGURE 6" captionStartId="10.[151,250,1898,1920]" captionTargetBox="[164,1419,309,1865]" captionTargetId="figure@10.[151,1436,301,1876]" captionTargetPageId="10" captionText="FIGURE 6. Perarella clavata (Jäderholm, 1905): A, polyp showing basal cup of perisarc (arrow); B, distal part of polyp; C, female gonophore showing basal cup of perisarc (arrow). Scale bar: 500 µm (A C)." httpUri="https://zenodo.org/record/236737/files/figure.png" pageId="9" pageNumber="378">Fig. 6</figureCitation>
A) [
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long and
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wide, according to
<bibRefCitation id="D1284B6DFF93B44210905A59FEBB388D" author="Jaderholm" box="[151,362,1895,1920]" pageId="9" pageNumber="378" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm (1905)</bibRefCitation>
], with conical hypostome and a distal crown of 1517 filiform tentacles (
<figureCitation id="2D822A19FF93B44215045A59FA9F388D" box="[1283,1358,1895,1920]" captionStart="FIGURE 6" captionStartId="10.[151,250,1898,1920]" captionTargetBox="[164,1419,309,1865]" captionTargetId="figure@10.[151,1436,301,1876]" captionTargetPageId="10" captionText="FIGURE 6. Perarella clavata (Jäderholm, 1905): A, polyp showing basal cup of perisarc (arrow); B, distal part of polyp; C, female gonophore showing basal cup of perisarc (arrow). Scale bar: 500 µm (A C)." httpUri="https://zenodo.org/record/236737/files/figure.png" pageId="9" pageNumber="378">Fig. 6</figureCitation>
A, B). Gonozooids absent; gonophores directly arising from hydrorhiza, club-shaped (
<figureCitation id="2D822A19FF93B442142A5AB2FBA238A8" box="[1069,1139,1932,1957]" captionStart="FIGURE 6" captionStartId="10.[151,250,1898,1920]" captionTargetBox="[164,1419,309,1865]" captionTargetId="figure@10.[151,1436,301,1876]" captionTargetPageId="10" captionText="FIGURE 6. Perarella clavata (Jäderholm, 1905): A, polyp showing basal cup of perisarc (arrow); B, distal part of polyp; C, female gonophore showing basal cup of perisarc (arrow). Scale bar: 500 µm (A C)." httpUri="https://zenodo.org/record/236737/files/figure.png" pageId="9" pageNumber="378">Fig. 6</figureCitation>
C), about
<quantity id="72419B79FF93B44214EE5AB2FAFC38A9" box="[1257,1325,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="378" unit="mm" value="2.0">2 mm</quantity>
long and
<quantity id="72419B79FF93B44210905A8EFEED38C5" box="[151,316,1968,1993]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="9" pageNumber="378" unit="mm" value="0.625" valueMax="0.75" valueMin="0.5">0.500.75 mm</quantity>
wide. Both gastrozooids and gonophores with a short, basal cup of perisarc (
<figureCitation id="2D822A19FF93B44214B25A8EFB2838C4" box="[1205,1273,1968,1993]" captionStart="FIGURE 6" captionStartId="10.[151,250,1898,1920]" captionTargetBox="[164,1419,309,1865]" captionTargetId="figure@10.[151,1436,301,1876]" captionTargetPageId="10" captionText="FIGURE 6. Perarella clavata (Jäderholm, 1905): A, polyp showing basal cup of perisarc (arrow); B, distal part of polyp; C, female gonophore showing basal cup of perisarc (arrow). Scale bar: 500 µm (A C)." httpUri="https://zenodo.org/record/236737/files/figure.png" pageId="9" pageNumber="378">Fig. 6</figureCitation>
A, C). Female gonophores with a very large number of eggs.
</paragraph>
<paragraph id="B506369CFF90B44110C05DA6FD093FF5" blockId="10.[151,1436,151,248]" pageId="10" pageNumber="379">
Measurements (in µm).
<emphasis id="87CDEA8EFF90B44111D15DA9FDEA3FBD" box="[470,571,151,176]" italics="true" pageId="10" pageNumber="379">Cnidome</emphasis>
: microbasic mastigophore [range 8.09.5 x 3.54.5, mean 9.2±0.5 x 4.2±0.3 (n=10); ratio, range 2.02.4, mean 2.2±0.1 (n=10)], desmonemes [range 5.56.5 x 3.54.5, mean 6.1±0.3 x 4.2±0.3 (n=10); ratio, range 1.31.6, mean 1.5±0.1 (n=10)].
</paragraph>
</subSubSection>
<caption id="E1C66614FF90B44110905A54FC363893" httpUri="https://zenodo.org/record/236737/files/figure.png" pageId="10" pageNumber="379" targetBox="[164,1419,309,1865]" targetPageId="10">
<paragraph id="B506369CFF90B44110905A54FC363893" blockId="10.[151,1435,1898,1950]" pageId="10" pageNumber="379">
<emphasis id="87CDEA8EFF90B44110905A54FEC53872" bold="true" box="[151,276,1898,1920]" pageId="10" pageNumber="379">FIGURE 6.</emphasis>
<taxonomicName id="72B94D1FFF90B441111C5A54FD45388D" authority="Jaderholm, 1905" authorityName="Jaderholm" authorityYear="1905" box="[283,660,1898,1920]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="10" pageNumber="379" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF90B441111C5A54FE1F388D" box="[283,462,1898,1920]" italics="true" pageId="10" pageNumber="379">Perarella clavata</emphasis>
(Jäderholm, 1905)
</taxonomicName>
: A, polyp showing basal cup of perisarc (arrow); B, distal part of polyp; C, female gonophore showing basal cup of perisarc (arrow). Scale bar: 500 µm (AC).
</paragraph>
</caption>
<subSubSection id="FDA36517FF91B44010C05DA9FB0C3DA5" pageId="11" pageNumber="380" type="discussion">
<paragraph id="B506369CFF91B44010C05DA9FE8E3E4D" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="380">
<emphasis id="87CDEA8EFF91B44010C05DA9FEEA3FBD" bold="true" box="[199,315,151,176]" pageId="11" pageNumber="380">Remarks.</emphasis>
<bibRefCitation id="D1284B6DFF91B44011435DA6FD4B3FBD" author="Rees" box="[324,666,151,176]" pageId="11" pageNumber="380" refString="Rees, W. J. (1962) Hydroids of the family Cytaeidae L. Agassiz, 1862. Bulletin of the British Museum (Natural History), Zoology Series, 8, 381 - 400." type="journal article" year="1962">Rees (1962: 394395, fig. 11)</bibRefCitation>
also provided a description of the species after examining the
<typeStatus id="6A02883EFF91B440156B5DA6FA4D3FBD" box="[1388,1436,152,176]" pageId="11" pageNumber="380">type</typeStatus>
material.
<taxonomicName id="72B94D1FFF91B44011065D82FE143FD8" box="[257,453,188,213]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF91B44011065D82FE143FD8" box="[257,453,188,213]" italics="true" pageId="11" pageNumber="380">Perarella clavata</emphasis>
</taxonomicName>
is a morphologically well-characterized species, with gonophores arising directly from the hydrorhiza. The only missing relevant information concerned the cnidome. I have tried to fill this gap with the reevaluation of the
<typeStatus id="6A02883EFF91B44011765C3BFE703E10" box="[369,417,261,285]" pageId="11" pageNumber="380">type</typeStatus>
material. The presence of a basal perisarc cup also supports that this species belong to the family
<taxonomicName id="72B94D1FFF91B44010E05C19FE8B3E4D" box="[231,346,295,320]" class="Hydrozoa" family="Cytaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="family">Cytaeidae</taxonomicName>
.
</paragraph>
<paragraph id="B506369CFF91B44010C05C73FE493D30" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="380">
<bibRefCitation id="D1284B6DFF91B44010C05C73FE963E68" author="Rees" box="[199,327,332,357]" pageId="11" pageNumber="380" refString="Rees, W. J. (1956) On three northern species of Hydractinia. Bulletin of the British Museum (Natural History), 3, 351 - 362." type="journal article" year="1956">Rees (1956</bibRefCitation>
,
<bibRefCitation id="D1284B6DFF91B44011545C72FE5D3E68" author="Rees" box="[339,396,332,357]" pageId="11" pageNumber="380" refString="Rees, W. J. (1962) Hydroids of the family Cytaeidae L. Agassiz, 1862. Bulletin of the British Museum (Natural History), Zoology Series, 8, 381 - 400." type="journal article" year="1962">1962</bibRefCitation>
) re-established the genus
<taxonomicName id="72B94D1FFF91B44012AB5C72FCC73E68" box="[684,790,332,357]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="genus">
<emphasis id="87CDEA8EFF91B44012AB5C72FCC73E68" box="[684,790,332,357]" italics="true" pageId="11" pageNumber="380">Perarella</emphasis>
</taxonomicName>
for species with fixed sporosacs or degenerated medusae.
<bibRefCitation id="D1284B6DFF91B44010905C51FE573E85" author="Bouillon" box="[151,390,367,392]" pageId="11" pageNumber="380" refString="Bouillon, J., Medel, M. D., Pages, F., Gili, J. M., Boero, B. &amp; Gravili, C. (2004) Fauna of the Mediterranean Hydrozoa. Scientia Marina, 68 (Supplement 2), 1 - 448. http: // dx. doi. org / 10.3989 / scimar. 2004.68 s 25" type="journal article" year="2004">
Bouillon
<emphasis id="87CDEA8EFF91B44011065C4FFEE23E85" box="[257,307,367,392]" italics="true" pageId="11" pageNumber="380">et al</emphasis>
. (2004
</bibRefCitation>
,
<bibRefCitation id="D1284B6DFF91B44011945C51FE1E3E85" author="Bouillon" box="[403,463,367,392]" pageId="11" pageNumber="380" refString="Bouillon, J., Gravili, C., Pages, F., Gili, J. M. &amp; Boero, F. (2006) An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle, 194, 1 - 591." type="journal article" year="2006">2006</bibRefCitation>
), following the idea that genera should not be delimited based exclusively on medusa reduction, considered, however, this genus congeneric with
<taxonomicName id="72B94D1FFF91B440133E5CABFC5C3EA1" box="[825,909,405,428]" class="Hydrozoa" family="Cytaeididae" genus="Cytaeis" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="genus">
<emphasis id="87CDEA8EFF91B440133E5CABFC5C3EA1" box="[825,909,405,428]" italics="true" pageId="11" pageNumber="380">Cytaeis</emphasis>
</taxonomicName>
. Nevertheless,
<bibRefCitation id="D1284B6DFF91B440143D5CAAFB2D3EA1" author="Schuchert" box="[1082,1276,404,429]" pageId="11" pageNumber="380" refString="Schuchert, P. (2007) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera part 2. Revue suisse de Zoologie, 114, 195 - 396." type="journal article" year="2007">Schuchert (2007)</bibRefCitation>
indicated that this approach is valid in cases where a hydroid can be attributed unambiguously to a certain family, but not in the case of
<taxonomicName id="72B94D1FFF91B44010E95CE2FE893EF8" box="[238,344,476,501]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="genus">
<emphasis id="87CDEA8EFF91B44010E95CE2FE893EF8" box="[238,344,476,501]" italics="true" pageId="11" pageNumber="380">Perarella</emphasis>
</taxonomicName>
, which could even belong to the families
<taxonomicName id="72B94D1FFF91B44013305CE2FC233EF8" box="[823,1010,476,501]" class="Hydrozoa" family="Bougainvilliidae" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="family">Bougainvilliidae</taxonomicName>
or
<taxonomicName id="72B94D1FFF91B440141E5CE2FB413EF8" box="[1049,1168,476,501]" class="Hydrozoa" family="Pandeidae" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="family">Pandeidae</taxonomicName>
. He concluded that it is advisable to retain the genus
<taxonomicName id="72B94D1FFF91B44011DB5CC1FD973D15" box="[476,582,511,536]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="genus">
<emphasis id="87CDEA8EFF91B44011DB5CC1FD973D15" box="[476,582,511,536]" italics="true" pageId="11" pageNumber="380">Perarella</emphasis>
</taxonomicName>
until its affinities are clearly revealed by molecular phylogenetic analyses. I agree with his opinion.
</paragraph>
<paragraph id="B506369CFF91B44010C05F79FB0C3DA5" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="380">
<emphasis id="87CDEA8EFF91B44010C05F79FE273D6D" bold="true" box="[199,502,583,608]" pageId="11" pageNumber="380">Ecology and distribution.</emphasis>
<taxonomicName id="72B94D1FFF91B44011FA5F79FD133D6D" box="[509,706,583,608]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF91B44011FA5F79FD133D6D" box="[509,706,583,608]" italics="true" pageId="11" pageNumber="380">Perarella clavata</emphasis>
</taxonomicName>
has been collected at depths from
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(
<bibRefCitation id="D1284B6DFF91B44014995F76FAA63D6D" author="Pena" box="[1182,1399,584,608]" pageId="11" pageNumber="380" refString="Pena Cantero, A. L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451 - 464. http: // dx. doi. org / 10.1007 / s 00300 - 007 - 0371 - 2" type="journal article" year="2008">Peña Cantero 2008</bibRefCitation>
) to
<quantity id="72419B79FF91B44010905F52FF0C3D89" box="[151,221,620,645]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="380" unit="m" value="360.0">360 m</quantity>
(
<bibRefCitation id="D1284B6DFF91B44010EC5F52FE723D88" author="Jaderholm" box="[235,419,620,645]" pageId="11" pageNumber="380" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm 1905</bibRefCitation>
), epibiotic on gastropod shells (
<taxonomicName id="72B94D1FFF91B44013005F52FC143D89" box="[775,965,620,644]" class="Gastropoda" family="Cochlespiridae" genus="Aforia" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="380" phylum="Mollusca" rank="species" species="magnifica">
<emphasis id="87CDEA8EFF91B44013005F52FC143D89" box="[775,965,620,644]" italics="true" pageId="11" pageNumber="380">Aforia magnifica</emphasis>
</taxonomicName>
) (
<bibRefCitation id="D1284B6DFF91B44013DB5F52FB453D88" author="Jaderholm" box="[988,1172,620,645]" pageId="11" pageNumber="380" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm 1905</bibRefCitation>
;
<bibRefCitation id="D1284B6DFF91B44014A75F53FAA83D89" author="Pena" box="[1184,1401,620,644]" pageId="11" pageNumber="380" refString="Pena Cantero, A. L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451 - 464. http: // dx. doi. org / 10.1007 / s 00300 - 007 - 0371 - 2" type="journal article" year="2008">Peña Cantero 2008</bibRefCitation>
). It has been found with gonophores in January (
<bibRefCitation id="D1284B6DFF91B440128B5FAEFCB43DA5" author="Pena" box="[652,869,656,680]" pageId="11" pageNumber="380" refString="Pena Cantero, A. L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451 - 464. http: // dx. doi. org / 10.1007 / s 00300 - 007 - 0371 - 2" type="journal article" year="2008">Peña Cantero 2008</bibRefCitation>
) and February (
<bibRefCitation id="D1284B6DFF91B44014115FB1FB013DA5" author="Jaderholm" box="[1046,1232,655,680]" pageId="11" pageNumber="380" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm 1905</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FDA36517FF91B44010C05F8AFA473DFD" pageId="11" pageNumber="380" type="reference_group">
<paragraph id="B506369CFF91B44010C05F8AFA473DFD" blockId="11.[151,1437,151,752]" pageId="11" pageNumber="380">
<taxonomicName id="72B94D1FFF91B44010C05F8AFE5A3DC0" ID-CoL="76RNY" box="[199,395,692,717]" class="Hydrozoa" family="Cytaeididae" genus="Perarella" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="380" phylum="Cnidaria" rank="species" species="clavata">
<emphasis id="87CDEA8EFF91B44010C05F8AFE5A3DC0" box="[199,395,692,717]" italics="true" pageId="11" pageNumber="380">Perarella clavata</emphasis>
</taxonomicName>
is endemic to West
<taxonomicName id="72B94D1FFF91B44012685F8AFD363DC0" box="[623,743,692,717]" class="Hydrozoa" family="Bougainvilliidae" genus="Gravelya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="377" phylum="Cnidaria" rank="species" species="antarctica">
<collectingCountry id="CDAE760CFF91B44012685F8AFD363DC0" box="[623,743,692,717]" name="Antarctica" pageId="11" pageNumber="380">Antarctica</collectingCountry>
</taxonomicName>
, being only known from the Erebus and Terror Gulf, Graham Land (
<bibRefCitation id="D1284B6DFF91B44010D85FE9FE443DFD" author="Jaderholm" box="[223,405,727,752]" pageId="11" pageNumber="380" refString="Jaderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Sudpolar-Expedition 1901 - 1903, 5 (8), 1 - 41, pls. 1 - 14." type="journal article" year="1905">Jäderholm 1905</bibRefCitation>
) and off the south of Livingston
<collectingCountry id="CDAE760CFF91B44013005FE9FC9F3DFD" box="[775,846,727,752]" name="Iceland" pageId="11" pageNumber="380">Island</collectingCountry>
, in the South
<collectingCountry id="CDAE760CFF91B44013E05FE9FB4E3DFD" box="[999,1183,727,752]" name="United Kingdom" pageId="11" pageNumber="380">Shetland Islands</collectingCountry>
(
<bibRefCitation id="D1284B6DFF91B44014A95FE6FA5B3DFD" author="Pena" box="[1198,1418,728,752]" pageId="11" pageNumber="380" refString="Pena Cantero, A. L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451 - 464. http: // dx. doi. org / 10.1007 / s 00300 - 007 - 0371 - 2" type="journal article" year="2008">Peña Cantero 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>