416 lines
61 KiB
XML
416 lines
61 KiB
XML
<document ID-CLB-Dataset="291768" ID-DOI="http://dx.doi.org/10.3897/asp.82.e114849" ID-GBIF-Dataset="9b17f188-eb79-4162-a1a3-1f076d362b55" ID-Pensoft-Pub="1864-8312-82-183" ID-Pensoft-UUID="05355FC72CC05232AC03F5909266BD42" ID-ZooBank="E45565A395804B06AB8B093CB0692937" ModsDocID="1864-8312-82-183" checkinTime="1711187908290" checkinUser="pensoft" docAuthor="Alvarez-Parra, Sergio, Nel, Andre, Perrichot, Vincent & Jouault, Corentin" docDate="2024" docId="3BCCD16D2A8C58B4ACCE16F23B210918" docLanguage="en" docName="ArthropodSystPhyl 82: 183-199" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e114849" docTitle="Santonipsocus mimeticus Álvarez-Parra & Nel & Perrichot & Jouault 2024, sp. nov." docType="treatment" docUuid="EB1D823F-33FF-4939-AFD6-1C8AFD81C503" docUuidSource="ZooBank" docVersion="4" id="05355FC72CC05232AC03F5909266BD42" lastPageNumber="183" masterDocId="05355FC72CC05232AC03F5909266BD42" masterDocTitle="Unravelling the mishmash: A new phylogeny for the family Empheriidae (Psocodea, Trogiomorpha) with a new genus and species from Cretaceous Charentese amber" masterLastPageNumber="199" masterPageNumber="183" pageNumber="183" updateTime="1732778970021" updateUser="ExternalLinkService">
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<mods:title id="D531E18EADA64BAE413DDC53F3073482">Unravelling the mishmash: A new phylogeny for the family Empheriidae (Psocodea, Trogiomorpha) with a new genus and species from Cretaceous Charentese amber</mods:title>
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<mods:namePart id="04DEB031F5656F0E672A324215646B21">Alvarez-Parra, Sergio</mods:namePart>
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<mods:affiliation id="0A1C922990A35C7C7F83D239AB0D5EA2">State Key Laboratory of Palaeobiology and Petroleum Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Departament de Dinamica de la Terra i de l'Ocea, Facultat de Ciencies de la Terra, Universitat de Barcelona, c / Marti i Franques s / n, 08028, Barcelona, Spain</mods:affiliation>
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<mods:nameIdentifier id="7ADA723145169648D91BA1603962B078" type="email">sergio.alvarez-parra@ub.edu</mods:nameIdentifier>
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<mods:namePart id="5517A714E631480A907B48B767217DF2">Nel, Andre</mods:namePart>
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<mods:affiliation id="9D7EC0FD958B05C5BF5BDA3A331BC0C5">Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona, Barcelona, Spain</mods:affiliation>
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<mods:namePart id="F73B789582473526DC8628C56277E548">Perrichot, Vincent</mods:namePart>
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<mods:affiliation id="5B8D9F42ABEF1ACBABA43671EF7CDCC0">Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 45 rue Buffon 75005 Paris, France</mods:affiliation>
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<mods:namePart id="0575AA4A5E620F1D95BE93421C93D380">Jouault, Corentin</mods:namePart>
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<mods:affiliation id="0936A5A13A860105A545021DB3B3DDB1">Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona, Barcelona, Spain & Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 45 rue Buffon 75005 Paris, France & Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, F- 35000 Rennes, France</mods:affiliation>
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<mods:title id="F5349C0A63829C8D0A722A3B0B7DBED9">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="3B5466CA47DB3563E9448EB695E63BE1">2024</mods:date>
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<subSection id="A629F72AFB7F0DF32691F1D220CDE6DD" pageId="0" pageNumber="183" type="3. systematic palaeontology">
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<treatment id="3BCCD16D2A8C58B4ACCE16F23B210918" ID-GBIF-Taxon="224057630" LSID="urn:lsid:zoobank.org:act:EB1D823F-33FF-4939-AFD6-1C8AFD81C503" httpUri="http://treatment.plazi.org/id/3BCCD16D2A8C58B4ACCE16F23B210918" lastPageNumber="183" pageId="0" pageNumber="183" scope_family="Empheriidae" scope_infraOrder="Atropetae" scope_order="Psocodea" scope_subOrder="Trogiomorpha">
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<taxonomicName id="DF633C371FF620F6E56134B3672CB968" LSID="https://zoobank.org/EB1D823F-33FF-4939-AFD6-1C8AFD81C503" authority="Álvarez-Parra & Nel & Perrichot & Jouault, 2024" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" family="Empheriidae" genus="Santonipsocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Santonipsocus mimeticus" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="mimeticus" status="sp. nov.">Santonipsocus mimeticus</taxonomicName>
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<taxonomicNameLabel id="7195359734B2BC3CF43D0F04FE32DFA7" pageId="0" pageNumber="183">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="765E31B62DAFFA65640029EABD032ED3" pageId="0" pageNumber="183" type="description">
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<paragraph id="FBEE5FD7B4F9BFAA1EA15B992F4B55DC" pageId="0" pageNumber="183">
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<figureCitation id="B8BE51B25B00A4C56DAF67FC6C430009" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view. Both at same scale." figureDoi="10.3897/asp.82.e114849.figure1" httpUri="https://binary.pensoft.net/fig/1007221" pageId="0" pageNumber="183">Figures 1</figureCitation>
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<figureCitation id="90C33DDCECDEC3CFAD9F49797D61CC63" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">, 2</figureCitation>
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<figureCitation id="753222C02F60F3D6277CD4E1EBBC6660" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">, 3</figureCitation>
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<figureCitation id="7B013388FA9BDDEBAAF6232317B74230" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), specimen IGR. ARC- 169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view; C photograph of wings; D schematic drawing of forewing; E photograph of the tarsi of right foreleg and midleg." figureDoi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="10CCD3CE084729F693DC15793017AAD9" pageId="0" pageNumber="183" type="holotype">
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<paragraph id="2A292528A407558944900E038EF44D43" pageId="0" pageNumber="183">Holotype.</paragraph>
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<paragraph id="F6E7183BEDC6101DE5FEDB5F15BCCB8F" pageId="0" pageNumber="183">
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MNHN.F.A30180 (ARC-186.7), adult specimen, probably male (Figs
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<figureCitation id="F287AC20AFB05ACB6C9E06CEC20D3C52" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view. Both at same scale." figureDoi="10.3897/asp.82.e114849.figure1" httpUri="https://binary.pensoft.net/fig/1007221" pageId="0" pageNumber="183">1</figureCitation>
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-
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<figureCitation id="9086BA5C9BCBAE9C9E1F5BB46B619284" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3</figureCitation>
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), erroneously figured as a paratype of
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<taxonomicName id="DBC15E15DB4A1B1FA137DFFA1C99AFF0" authorityName="Perrichot, Azar, Neraudeau & Nel" authorityYear="2003" class="Insecta" family="Archaeatropidae" genus="Proprionoglaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proprionoglaris guyoti" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="guyoti">
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<emphasis id="0A4B88C89F2BD9FFD7B4CE74D7D9EC4A" italics="true" pageId="0" pageNumber="183">Proprionoglaris guyoti</emphasis>
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</taxonomicName>
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in the PhD dissertation of
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<bibRefCitation id="B1AA0173AD3ACD533AF38C0411CAFD99" author="Perrichot, V" journalOrPublisher="Memoires de Geosciences Rennes" pageId="0" pageNumber="183" pagination="1 - 310" refId="B59" refString="Perrichot, V, 2005. Environnements paraliques a ambre et a vegetaux du Cretace nord-aquitain (Charentes, Sud-Ouest de la France). Memoires de Geosciences Rennes 118: 1 - 310" title="Environnements paraliques a ambre et a vegetaux du Cretace nord-aquitain (Charentes, Sud-Ouest de la France)." volume="118" year="2005">Perrichot (2005</bibRefCitation>
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, fig. 31b). Amber fragment cut from a larger amber piece (
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<normalizedToken id="32E28AD08EF3BC3012822BCC756771DD" originalValue="‘ARC-186’">'ARC-186'</normalizedToken>
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) and prepared in Canada balsam. Originally in syninclusion with the holotype of
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<taxonomicName id="41DD493C21C7C28AED57E29B727B0612" authorityName="Perrichot, Azar, Neraudeau & Nel" authorityYear="2003" class="Insecta" family="Archaeatropidae" genus="Prospeleketor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prospeleketor albianensis" order="Pseudoscorpionida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="albianensis">
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<emphasis id="3C3CDCEE32CA5556FA42C7EE3F68E249" italics="true" pageId="0" pageNumber="183">Prospeleketor albianensis</emphasis>
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</taxonomicName>
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Perrichot, Azar,
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<normalizedToken id="F551CF71183E015E57014B7DC347161C" originalValue="Néraudeau">Neraudeau</normalizedToken>
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and Nel, 2003 (specimen MNHN.F.A30111 [ARC-186.10]) and nine other arthropods (
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<taxonomicName id="3E8EBEE7BE3959A6AF5425BC33AA8900" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
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,
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<taxonomicName id="1C3C658F5A0B7F829D36881BD0C1206F" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
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,
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<taxonomicName id="8201CD8A0F44C6552A2F4BB8061120B9" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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, and
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<taxonomicName id="05B7BC7C171226197BAD582201C99BC2" lsidName="" order="Pseudoscorpionida" pageId="0" pageNumber="183" rank="order">Pseudoscorpionida</taxonomicName>
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). In the article by
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<bibRefCitation id="9B3F0CB4B0F4EB062F936B8D8442CFEC" DOI="https://doi.org/10.1017/S0016756803008355" author="Perrichot, V" journalOrPublisher="Geological Magazine" pageId="0" pageNumber="183" pagination="669 - 683" refId="B60" refString="Perrichot, V, Azar, D, Neraudeau, D, Nel, A, 2003. New Psocoptera in the Early Cretaceous amber of SW France and Lebanon (Insecta: Psocoptera: Trogiomorpha). Geological Magazine 140 (6): 669 - 683, DOI: https://doi.org/10.1017/S0016756803008355" title="New Psocoptera in the Early Cretaceous amber of SW France and Lebanon (Insecta: Psocoptera: Trogiomorpha)." url="https://doi.org/10.1017/S0016756803008355" volume="140" year="2003">Perrichot et al. (2003)</bibRefCitation>
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||
, it is indicated that the holotype of
|
||
<taxonomicName id="4655845736DFBC435085F585F1C9D25A" lsidName="P. albianensis" pageId="0" pageNumber="183" rank="species" species="albianensis">
|
||
<emphasis id="5EBF64A3E08D1F8A4736597B20A18DAC" italics="true" pageId="0" pageNumber="183">P. albianensis</emphasis>
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||
</taxonomicName>
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||
is together with the female paratype of
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<taxonomicName id="941FD760D342EBD48D23AF8DDCCC5F23" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
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<emphasis id="8215B2D6A08855DDE046BABE23EF657B" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
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||
</taxonomicName>
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as syninclusions. However,
|
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<taxonomicName id="E3AAA058C4E040CC8CE4E497479253BF" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
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||
<emphasis id="8E42522DCACE6B8C7FF57CF071F1349C" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
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</taxonomicName>
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was described from the female holotype (MNHN.F.A30108 [ARC-58.2]) and two male paratypes (MNHN.F.A30109 [ARC-50.1] and MNHN.F.A30110 [ARC-201.2]), all in other amber pieces. All the above-mentioned material is housed in the palaeontological collection of the
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<normalizedToken id="E000959FC1CAAF87AAA6818CFF6A5392" originalValue="Muséum">Museum</normalizedToken>
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National
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<normalizedToken id="EE0521C8B875F3F903A258455AF4C5FC" originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle, Paris, France.
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</paragraph>
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<paragraph id="3B2BE5238A304C0C15EF518587B7AC9D" pageId="0" pageNumber="183">
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<emphasis id="758844AB239EF5DBCE24A3467EE4EA88" bold="true" pageId="0" pageNumber="183">Figure 1.</emphasis>
|
||
<taxonomicName id="131B6E74D6B14A5F45AB53600C22837A" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" family="Empheriidae" genus="Santonipsocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Santonipsocus mimeticus" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="mimeticus">
|
||
<emphasis id="49C11CA0E1D9D906B332429ABB2ADEF2" italics="true" pageId="0" pageNumber="183">Santonipsocus mimeticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2EC0E3DE742188629C72BC0A5AFD2F05" bold="true" pageId="0" pageNumber="183">gen. et sp. nov.</emphasis>
|
||
(
|
||
<taxonomicName id="DC587041766B8D4A971CAE98BCEA2973" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
,
|
||
<taxonomicName id="FC46638D9C0B3D0AFA049D76B85A3294" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="33AFE83F17702094E863F548049A73EC" authorityName="Baz & Ortuno" authorityYear="2000" class="Insecta" family="Empheriidae" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Empheriidae</taxonomicName>
|
||
), holotype MNHN.F.A30180 (ARC-186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France).
|
||
<emphasis id="DCB6861BBD95112DD668BDAF62F2684E" bold="true" pageId="0" pageNumber="183">A</emphasis>
|
||
habitus from dorsal view;
|
||
<emphasis id="7853F1746964B1086B9DFAE0D9D0C90E" bold="true" pageId="0" pageNumber="183">B</emphasis>
|
||
habitus from ventral view. Both at same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="44A988817970996B9A77F7EF04CE262E" doi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183" start="Figure 2" startId="F2">
|
||
<paragraph id="D80BA8DE02AD2B3AA084A158B998BD41" pageId="0" pageNumber="183">
|
||
<emphasis id="7DF07F15411BF0F2389859847AE39763" bold="true" pageId="0" pageNumber="183">Figure 2.</emphasis>
|
||
<taxonomicName id="27078F76E7C9CAAF231443418625F9A3" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" family="Empheriidae" genus="Santonipsocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Santonipsocus mimeticus" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="mimeticus">
|
||
<emphasis id="077B00EF5350A88A586D78773EC5F5FF" italics="true" pageId="0" pageNumber="183">Santonipsocus mimeticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="487EF0473ABA9F3F1CB2E8F005D22C93" bold="true" pageId="0" pageNumber="183">gen. et sp. nov.</emphasis>
|
||
(
|
||
<taxonomicName id="4A7C176E420D131304EEDDDF795B40F3" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
,
|
||
<taxonomicName id="C807B378D6B835D31CA466AA6381100D" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="9718CA66D63EB46C81E1EF3741354D99" authorityName="Baz & Ortuno" authorityYear="2000" class="Insecta" family="Empheriidae" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Empheriidae</taxonomicName>
|
||
), holotype MNHN.F.A30180 (ARC-186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France).
|
||
<emphasis id="EC23AB6F7615538AD1CEB77746E4916B" bold="true" pageId="0" pageNumber="183">A</emphasis>
|
||
photograph of right fore- and hind wings, overlapped;
|
||
<emphasis id="7F6F08930904A7431331AE5987940DC0" bold="true" pageId="0" pageNumber="183">B</emphasis>
|
||
schematic drawing of forewing;
|
||
<emphasis id="9BEB0D66C985C4E3B369A63F8D8E7060" bold="true" pageId="0" pageNumber="183">C</emphasis>
|
||
schematic drawing of hind wing. B and C at same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="06A82D40DFBE229000C64E69A30F0366" doi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183" start="Figure 3" startId="F3">
|
||
<paragraph id="BE7132F6530DD6531B3C9BD456DE9888" pageId="0" pageNumber="183">
|
||
<emphasis id="C2A17C699253D5D075E337708A6B2028" bold="true" pageId="0" pageNumber="183">Figure 3.</emphasis>
|
||
<taxonomicName id="DD2FA476B037D1B0807DAFECAC4B752B" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" family="Empheriidae" genus="Santonipsocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Santonipsocus mimeticus" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="mimeticus">
|
||
<emphasis id="62FDCB00D14F2FFB0859C5AE1C237950" italics="true" pageId="0" pageNumber="183">Santonipsocus mimeticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="62DE7A81FC304ACC7D94B3C0ED951587" bold="true" pageId="0" pageNumber="183">gen. et sp. nov.</emphasis>
|
||
(
|
||
<taxonomicName id="D89A37AACA62079E7E137052CFAB20D7" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0BAF3D742463912108D2FAEF032D7A6" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="B3FAA37562542B6E0AE18F5E2E6EF1B1" authorityName="Baz & Ortuno" authorityYear="2000" class="Insecta" family="Empheriidae" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Empheriidae</taxonomicName>
|
||
), holotype MNHN.F.A30180 (ARC-186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France).
|
||
<emphasis id="0D243A5275BF454B2C04FFC755533948" bold="true" pageId="0" pageNumber="183">A</emphasis>
|
||
photograph of left antenna;
|
||
<emphasis id="866601FB978C70461DD76021071C73D2" bold="true" pageId="0" pageNumber="183">B</emphasis>
|
||
detail of flagellomeres of left antenna;
|
||
<emphasis id="64CEAA9A4C6001D50C0D1DFE251564E3" bold="true" pageId="0" pageNumber="183">C</emphasis>
|
||
head from ventral view;
|
||
<emphasis id="D87014417BAEF7EE60904E9B4F09478B" bold="true" pageId="0" pageNumber="183">D</emphasis>
|
||
,
|
||
<emphasis id="7F6D498A5B7F7DEC5478E8C51F58C292" bold="true" pageId="0" pageNumber="183">E</emphasis>
|
||
photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu2 and 1A with forewing margin, red arrowhead: emergence of R1 in relation to basal cell in hind wing;
|
||
<emphasis id="E1FADEA050DE7115952520AAB9FA943C" bold="true" pageId="0" pageNumber="183">F</emphasis>
|
||
photograph of left midleg and hind leg;
|
||
<emphasis id="17A01523DE82D15ED4F571C0ED74DCBE" bold="true" pageId="0" pageNumber="183">G</emphasis>
|
||
photograph of tarsus of right hind leg;
|
||
<emphasis id="3F82BF437CCAF7C77C854DE31113BE9D" bold="true" pageId="0" pageNumber="183">H</emphasis>
|
||
genitalia in ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="1DC5FFC6B876612B9DAE7FD931E91540" pageId="0" pageNumber="183" type="materials_examined">
|
||
<paragraph id="A05B4FD57FB9395F7DA90EDA0E51F351" pageId="0" pageNumber="183">Other material.</paragraph>
|
||
<paragraph id="45D6A6CD178897EA8820D7DCD5F373CD" pageId="0" pageNumber="183">
|
||
IGR.ARC-169, adult specimen, sex unknown (Fig.
|
||
<figureCitation id="61073B6DCD4C8CD2410220B3A29729DA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), specimen IGR. ARC- 169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view; C photograph of wings; D schematic drawing of forewing; E photograph of the tarsi of right foreleg and midleg." figureDoi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183">4</figureCitation>
|
||
). From Archingeay-Les Nouillers amber deposit. Amber piece prepared in
|
||
<collectingCountry id="62D92BE38627EE0327D375BA5C07112C" name="Canada">Canada</collectingCountry>
|
||
balsam; the forewings are separated from the rest of the body. Undetermined arthropod leg as syninclusion. Housed in the Geological Department and Museum of the University of Rennes,
|
||
<collectingCountry id="B9B7320E161A38B5178011163DB9E015" name="France">France</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="B153AEF5A0ACEC8C4FF4D66B09550E65" doi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183" start="Figure 4" startId="F4">
|
||
<paragraph id="FC8119141B55B132D76C2044244124B6" pageId="0" pageNumber="183">
|
||
<emphasis id="1C889273FF2E8527D931953F41D264F5" bold="true" pageId="0" pageNumber="183">Figure 4.</emphasis>
|
||
<taxonomicName id="FD6C5135892E8112AFED01E0D5F3D89B" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" family="Empheriidae" genus="Santonipsocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Santonipsocus mimeticus" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="mimeticus">
|
||
<emphasis id="BF3FE8FF6D115EDBC14E4CD95AD42F64" italics="true" pageId="0" pageNumber="183">Santonipsocus mimeticus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="ED88A6620838F09AD79EF3301BD8E05E" bold="true" pageId="0" pageNumber="183">gen. et sp. nov.</emphasis>
|
||
(
|
||
<taxonomicName id="A01DC284231746C4F7525CA98CAD0478" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
,
|
||
<taxonomicName id="9A0E8E27F5D04D512BC51B4519696668" authorityName="Álvarez-Parra & Nel & Perrichot & Jouault" authorityYear="2024" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="634701E444CC721F841619491AF9344C" authorityName="Baz & Ortuno" authorityYear="2000" class="Insecta" family="Empheriidae" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Empheriidae</taxonomicName>
|
||
), specimen IGR.ARC-169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (
|
||
<collectingCountry id="349226A5FDF9BBB30DB1886EC5437C59" name="France">France</collectingCountry>
|
||
).
|
||
<emphasis id="28DC1432C02C94E989977A9EB4265D19" bold="true" pageId="0" pageNumber="183">A</emphasis>
|
||
habitus from dorsal view;
|
||
<emphasis id="B4C9820A51CB2425F40D0DB772923948" bold="true" pageId="0" pageNumber="183">B</emphasis>
|
||
habitus from ventral view;
|
||
<emphasis id="370B77A59C4F08D1387C8E9F8FFDCE26" bold="true" pageId="0" pageNumber="183">C</emphasis>
|
||
photograph of wings;
|
||
<emphasis id="E5299ADC99CCD72373673C9FBB5F5424" bold="true" pageId="0" pageNumber="183">D</emphasis>
|
||
schematic drawing of forewing;
|
||
<emphasis id="34BD3208FF2BB47E68D379BCAFCC5DDD" bold="true" pageId="0" pageNumber="183">E</emphasis>
|
||
photograph of the tarsi of right foreleg and midleg.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="98B25AE9CA3732946815875E2C0F79A6" pageId="0" pageNumber="183" type="locality and horizon">
|
||
<paragraph id="80497572BA8EC75DA904CC4FF04108C4" pageId="0" pageNumber="183">Locality and horizon.</paragraph>
|
||
<paragraph id="4038194DF327ABDEBE0B3FB1F70C7BEB" pageId="0" pageNumber="183">
|
||
Font-de-Benon quarry, Archingeay-Les Nouillers, Charente-Maritime Department (Nouvelle-Aquitaine, France); level A1sl, uppermost Albian-lowermost Cenomanian, Cretaceous (
|
||
<bibRefCitation id="867F2049543D70EAF2D1BBA656ADBDD3" DOI="https://doi.org/10.1016/S0016-6995(02)00024-4" author="Neraudeau, D" journalOrPublisher="Geobios" pageId="0" pageNumber="183" pagination="233 - 240" refId="B51" refString="Neraudeau, D, Perrichot, V, Dejax, J, Masure, E, Nel, A, Philippe, M, Moreau, P, Guillocheau, F, Guyot, T, 2002. Un nouveau gisement a ambre insectifere et a vegetaux (Albien terminal probable): Archingeay (Charente-Maritime, France). Geobios 35 (2): 233 - 240, DOI: https://doi.org/10.1016/S0016-6995(02)00024-4" title="Un nouveau gisement a ambre insectifere et a vegetaux (Albien terminal probable): Archingeay (Charente-Maritime, France)." url="https://doi.org/10.1016/S0016-6995(02)00024-4" volume="35" year="2002">
|
||
<normalizedToken id="6E7F2120FC036DCA53A4C2DED78692D1" originalValue="Néraudeau">Neraudeau</normalizedToken>
|
||
et al. 2002
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B9B5AB2177CC8676F358C9B5F3E51602" pageId="0" pageNumber="183" type="etymology">
|
||
<paragraph id="2CF3893B9D014713E3DCF243776E3729" pageId="0" pageNumber="183">Etymology.</paragraph>
|
||
<paragraph id="61981751393DD6AE25CFB79AC4A67DA5" pageId="0" pageNumber="183">
|
||
From the Greek
|
||
<normalizedToken id="FF2496F055734E1F28E1C2F30E1610E8" originalValue="μιμητικός">μιμητικός</normalizedToken>
|
||
, meaning
|
||
<normalizedToken id="C1B269C067E51CADF93872973C03813F" originalValue="‘imitator’">'imitator'</normalizedToken>
|
||
, based on the resemblance with other Cretaceous barklice species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A88E7C2E578C48B062B22C50867A9C0A" pageId="0" pageNumber="183" type="diagnosis">
|
||
<paragraph id="54D325E069B80F1119B4841645D614C4" pageId="0" pageNumber="183">Diagnosis.</paragraph>
|
||
<paragraph id="59608D1CB04EA909D954153EE311E938" pageId="0" pageNumber="183">As for the genus (vide supra).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CBA8FA4E945E5598269007D3A1ED7EB" pageId="0" pageNumber="183" type="description">
|
||
<paragraph id="84C025F08722D1776E4E1BAA2F8C5C9E" pageId="0" pageNumber="183">Description.</paragraph>
|
||
<paragraph id="3D94D3774CB89BF71624CB9D6285666F" pageId="0" pageNumber="183">
|
||
Probably male. Body length 1.75 mm from clypeus to genitalia (Fig.
|
||
<figureCitation id="069EF449CBEB4614E54B3060465C614F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view. Both at same scale." figureDoi="10.3897/asp.82.e114849.figure1" httpUri="https://binary.pensoft.net/fig/1007221" pageId="0" pageNumber="183">1</figureCitation>
|
||
). -
|
||
<emphasis id="6103F5DAF01AFD2C35A566447465C17E" bold="true" italics="true" pageId="0" pageNumber="183">Head</emphasis>
|
||
: almost twice wider than long, 0.26 mm long, 0.44 mm wide; vertex covered by setae; marked epicranial suture with indistinct anterior arms; no ocelli; compound eye not prominent and small; left antenna complete with 17 flagellomeres (Fig.
|
||
<figureCitation id="151EB009B57994BD30F3BA789DF21EF9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3A</figureCitation>
|
||
), right antenna with eight flagellomeres preserved; short setae on proximal flagellomeres; scape 0.04 mm long, pedicel 0.05 mm long; lengths of flagellomeres of left antenna: f1 0.04 mm, f2 0.04 mm, f3 0.05 mm, f4 0.07 mm, f5 0.07 mm, f6 0.06 mm, f7 0.06 mm, f8 0.07 mm, f9 0.06 mm, f10 0.07 mm, f11 0.07 mm, f12 0.04 mm, f13 0.06 mm, f14 0.06 mm, f15 0.05 mm, f16 0.04 mm, f17 0.04 mm; flagellomeres lacking secondary annulations (Fig.
|
||
<figureCitation id="61BFD4F7A6BFE3C7F9A8A3B2E6EBDEDE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3B</figureCitation>
|
||
); clypeus bulging; maxillary palps four-segmented, covered by short setae (Fig.
|
||
<figureCitation id="595DD38B5116F533721A2A6E8A6D6D24" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3C</figureCitation>
|
||
); lengths of maxillary palpomeres: I 0.02 mm, II 0.08 mm, III 0.02 mm, IV 0.07 mm; sensillum not present on second maxillary palpomere; distal maxillary palpomere globose and rounded; labial palps seemingly two-segmented, with the distal labial palpomere showing an elongate shape 0.06 mm long (Fig, 3C); a structure might correspond to lacinia. -
|
||
<emphasis id="59F954868A6F5433326DE74A06A0E433" bold="true" italics="true" pageId="0" pageNumber="183">Thorax</emphasis>
|
||
: 0.46 mm long; pronotum bulging, covered by a few setae; macropterous (Fig.
|
||
<figureCitation id="1C342F4944C6EE69E9C5BFB0A2823E06" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">2</figureCitation>
|
||
). -
|
||
<emphasis id="BCC84280E5F91578C2D936D9292A427D" bold="true" italics="true" pageId="0" pageNumber="183">Wings</emphasis>
|
||
: membrane hyaline with setae; forewing and hind wing with nearly the same size, surpassing the distal part of the abdomen only slightly (Fig.
|
||
<figureCitation id="7A942CA91FF8F1B1B19946DC57F090F5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view. Both at same scale." figureDoi="10.3897/asp.82.e114849.figure1" httpUri="https://binary.pensoft.net/fig/1007221" pageId="0" pageNumber="183">1</figureCitation>
|
||
); forewing with margin covered by setae, veins with two rows of setae (Fig.
|
||
<figureCitation id="5517F135DDB037EE56E7DDD35DD26C2D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">2A</figureCitation>
|
||
); hind wing glabrous (Fig.
|
||
<figureCitation id="4FECC3D6B7C7FB38541FFD8858918BE0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">2B</figureCitation>
|
||
). Forewing 3
|
||
<normalizedToken id="BDD897A1A503B4AA503082C5CB6A36D1" originalValue="×">x</normalizedToken>
|
||
longer than wide, 1.39 mm long and 0.46 mm wide (Fig.
|
||
<figureCitation id="A59830D449E2D040775EE406AE544B59" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">2A</figureCitation>
|
||
); basal section of Sc curved and joining R1 at 0.60 mm from wing base, forming a narrow cell; a short, curved crossvein emerging from basal section of Sc, very close to the meeting point of Sc and R1, reaching margin at 0.67 mm from wing base; distal bent of basal section of Sc between emerging of crossvein and meeting point with R1 curved, not straight or perpendicular to R1; distal section of Sc separating from R1 and joining wing margin at 1.02 mm from wing base showing a sigmoidal path; R1 long reaching margin at 1.22 mm from wing base; basal section of Rs oblique, with crossvein between R1 and Rs present, forming a six-angled radial cell; vein Rs+M 0.06 mm long; Rs bifurcating at 0.86 mm from wing base; R2+3 and R4+5 reaching margin at 1.34 mm and 1.37 mm from wing base, respectively; bifurcation of M1 and M2 at 0.90 mm from wing base, reaching margin at 1.38 mm and 1.34 mm from wing base, respectively; M3 emerging from M at 0.65 mm from wing base and reaching margin at 1.26 mm from wing base; cells between R2+3 and R4+5, and between M1 and M2, elongate and relatively narrow; bifurcation of Cu1 into Cu1a and Cu1b at 0.48 mm from wing base; elongate and narrow areola postica, with Cu1a extending towards apex; Cu1a and Cu1b reaching margin at 1.11 mm and 0.79 mm from wing base, respectively; evanescent Cu2, without rows of setae, extending straight and reaching margin at 0.65 mm from wing base; vein 1A showing sigmoidal path and reaching margin at 0.56 mm from wing base; no nodulus (Fig.
|
||
<figureCitation id="6C75B2004E3918A90057C19F5FF10DCD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3D, E</figureCitation>
|
||
); joining of Cu2 with margin separated by 0.11 mm from joining of 1A with margin. Hind wing almost 3
|
||
<normalizedToken id="416713E967F09BA1BCC4AD3469F8AC7C" originalValue="×">x</normalizedToken>
|
||
longer than wide, 1.20 mm long and 0.42 mm wide (Fig.
|
||
<figureCitation id="93803773BC85BBF4BDAA861CFC39319D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of right fore- and hind wings, overlapped; B schematic drawing of forewing; C schematic drawing of hind wing. B and C at same scale." figureDoi="10.3897/asp.82.e114849.figure2" httpUri="https://binary.pensoft.net/fig/1007222" pageId="0" pageNumber="183">2B</figureCitation>
|
||
); Sc not visible; basal cell closed, elongate and narrow, with three angles, 0.15 mm long and 0.03 mm wide; R1 not emerging from apex of basal cell; R1 fused to Rs+M for 0.03 mm; R1 emerging from basal cell in specimen IGR.ARC-169; sigmoidal R1 reaching margin at 0.83 mm from wing base; bifurcation of Rs into R2+3 and R4+5 at the same level as R1 reaching margin; R2+3 and R4+5 reaching margin at 1.14 mm and 1.19 mm from wing base, respectively; cell between M1 and M2 elongate and narrow; M1 and M2 reaching margin at 1.12 mm and 0.97 mm from wing base, respectively; Cu1 reaching margin at 0.62 mm from wing base; a vein reaching margin close to Cu1 might correspond to Cu2; anal vein not visible. -
|
||
<emphasis id="97ADF43070B022207D080571E9A31937" bold="true" italics="true" pageId="0" pageNumber="183">Legs</emphasis>
|
||
: femora thick, without setae; tibiae thin, with three to four spines, and covered by setae; tarsi three-segmented, with first tarsomere covered by a few short setae (Fig.
|
||
<figureCitation id="9A2E39375F07E38A72A957BE95955E03" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3F, G</figureCitation>
|
||
); lengths of tarsomeres in foreleg: I 0.10 mm, II 0.04 mm, III 0.03 mm; lengths of tarsomeres in midleg: I 0.13 mm, II 0.03 mm, III 0.03 mm; lengths of tarsomeres in hind leg: I 0.18 mm, II 0.05 mm, III 0.04 mm; pretarsal claws lacking preapical tooth and pulvillus. -
|
||
<emphasis id="51D84B639848E8BE247B219B6085D01C" bold="true" italics="true" pageId="0" pageNumber="183">Abdomen</emphasis>
|
||
: 1.03 mm long; genitalia poorly visible, although male characters, such as hypandrium and paraprocts, are discernible (Fig.
|
||
<figureCitation id="9A5612523FA4357E0566617EDBE1682C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), holotype MNHN. F. A 30180 (ARC- 186.7), adult specimen, probably male; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A photograph of left antenna; B detail of flagellomeres of left antenna; C head from ventral view; D, E photograph and schematic drawing, respectively, of nodulus area of right forewing, both at same scale, blue arrowheads: meeting point of veins Cu 2 and 1 A with forewing margin, red arrowhead: emergence of R 1 in relation to basal cell in hind wing; F photograph of left midleg and hind leg; G photograph of tarsus of right hind leg; H genitalia in ventral view." figureDoi="10.3897/asp.82.e114849.figure3" httpUri="https://binary.pensoft.net/fig/1007223" pageId="0" pageNumber="183">3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55FBC472F581C4E64355F9042256E223" pageId="0" pageNumber="183" type="remarks">
|
||
<paragraph id="9A2ADDF9E595ED27EB00B9FDF6B6F686" pageId="0" pageNumber="183">Remarks.</paragraph>
|
||
<paragraph id="A020A1FBC09734AAC9BD653994E95F4B" pageId="0" pageNumber="183">
|
||
Although the specimen IGR.ARC-169 is poorly preserved (Fig.
|
||
<figureCitation id="15196974E32CBF69A04178155FA7D61B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), specimen IGR. ARC- 169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view; C photograph of wings; D schematic drawing of forewing; E photograph of the tarsi of right foreleg and midleg." figureDoi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183">4A, B</figureCitation>
|
||
), it is preliminary considered conspecific with holotype MNHN.F.A30180 (ARC-186.7) based on coincident diagnostic characters of the new genus: body length from clypeus to genitalia (1.75 mm holotype vs. 1.72 mm); distal maxillary palpomere globose and rounded; forewing with setae on margin, two rows of setae on veins, and very similar venation (Fig.
|
||
<figureCitation id="4F20629FA808DEAE54A0C705371CCF46" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), specimen IGR. ARC- 169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view; C photograph of wings; D schematic drawing of forewing; E photograph of the tarsi of right foreleg and midleg." figureDoi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183">4C, D</figureCitation>
|
||
), particularly presence of a cross-vein between Sc and margin emerging very close to meeting point of Sc with R1, distal bent of basal section of Sc curved between emerging of crossvein and meeting point with R1, not straight or perpendicular to R1, without nodulus; and pretarsal claws lacking preapical tooth and pulvillus (Fig.
|
||
<figureCitation id="3375FFA8A1498BA63A93461346C28966" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Santonipsocus mimeticus gen. et sp. nov. (Psocodea, Trogiomorpha, Empheriidae), specimen IGR. ARC- 169, sex unknown; uppermost Albian-lowermost Cenomanian amber of Archingeay-Les Nouillers (France). A habitus from dorsal view; B habitus from ventral view; C photograph of wings; D schematic drawing of forewing; E photograph of the tarsi of right foreleg and midleg." figureDoi="10.3897/asp.82.e114849.figure4" httpUri="https://binary.pensoft.net/fig/1007224" pageId="0" pageNumber="183">4E</figureCitation>
|
||
). The forewing venation of IGR.ARC-169 is nebulous and difficult to see, but it can be discerned using a combination of transmitted and reflected light under a stereomicroscope. We consider the distal maxillary palpomere globose and rounded and the lack of nodulus as key characters of the new genus. Two differences in wing venation can be related to intraspecific variability or teratism typical in barklice (
|
||
<bibRefCitation id="A0DF4DB430AC48DCD75D9CD45A4C7356" DOI="https://doi.org/10.3853/j.0067-1967.14.1972.424" author="Smithers, CN" journalOrPublisher="Australian Musseum Memoir" pageId="0" pageNumber="183" pagination="1 - 349" refId="B73" refString="Smithers, CN, 1972. The classification and phylogeny of the Psocoptera. Australian Musseum Memoir 14: 1 - 349, DOI: https://doi.org/10.3853/j.0067-1967.14.1972.424" title="The classification and phylogeny of the Psocoptera." url="https://doi.org/10.3853/j.0067-1967.14.1972.424" volume="14" year="1972">Smithers 1972</bibRefCitation>
|
||
): forewing with basal section of Rs placed more proximal in IGR.ARC-169 than in holotype and hind wing with four-angled basal cell in IGR.ARC-169 in comparison with three-angled basal cell in the holotype. Based on the coincident characters and the poor preservation of IGR.ARC-169, we believe that the most parsimonious option is to place this specimen, at least preliminarily, in the same species as the holotype, yet refraining from including it within the type material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="687FE05F8888ABC903218AAF3F8FC357" pageId="0" pageNumber="183" type="notes on proprionoglaris guyoti and proprionoglaris axioperierga">
|
||
<paragraph id="DFFB9923B999FCEE0D7D2F6BB9BADB75" pageId="0" pageNumber="183">
|
||
Notes on
|
||
<taxonomicName id="92777854CCCB84CE2AADA9B552B97AA2" authorityName="Perrichot, Azar, Neraudeau & Nel" authorityYear="2003" class="Insecta" family="Archaeatropidae" genus="Proprionoglaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proprionoglaris guyoti" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="guyoti">
|
||
<emphasis id="DFFE554A141EF23E97AE035C1050C092" italics="true" pageId="0" pageNumber="183">Proprionoglaris guyoti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="647E0D7E7C2755BFE592625EC96DEAB1" authorityName="Azar, Nel & Perrichot" authorityYear="2014" class="Insecta" family="Archaeatropidae" genus="Proprionoglaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proprionoglaris axioperierga" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="axioperierga">
|
||
<emphasis id="2CA00A090F50178214625C916F8519C4" italics="true" pageId="0" pageNumber="183">Proprionoglaris axioperierga</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5123FE4E22401C85A3C3DD2F8ABAA034" pageId="0" pageNumber="183">
|
||
The presence of a nodulus in forewing is confirmed for both species (Fig.
|
||
<figureCitation id="FB316E537F68C7BBE3FB300043B6A8AF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Forewings of Proprionoglaris guyoti and Proprionoglaris axioperierga (Psocodea, Trogiomorpha, Empheriidae). A holotype of P. guyoti, female, MNHN. F. A 30108 (ARC 58.2 R); B paratype of P. guyoti, male, MNHN. F. A 30109 (ARC 201.2); C - E new specimens of P. guyoti, sex unknown, IGR. ARC- 352.1, IGR. ARC- 157, and IGR. ARC- 355, respectively; F holotype of P. axioperierga, female, IGR. GAR- 69; G paratype of P. axioperierga, sex unknown, IGR. GAR- 39. White arrowheads: nodulus." figureDoi="10.3897/asp.82.e114849.figure5" httpUri="https://binary.pensoft.net/fig/1007225" pageId="0" pageNumber="183">5</figureCitation>
|
||
). The species
|
||
<taxonomicName id="726CCC6BD4CEFFBD3231D8D8208953B9" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
|
||
<emphasis id="6E5E398FCB19EE3372BE1E3F72D9C9B1" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
|
||
</taxonomicName>
|
||
was described as possessing only one row of setae along veins in forewing (
|
||
<bibRefCitation id="8396DEC3EDD2FE6655397C92A33387B9" DOI="https://doi.org/10.1017/S0016756803008355" author="Perrichot, V" journalOrPublisher="Geological Magazine" pageId="0" pageNumber="183" pagination="669 - 683" refId="B60" refString="Perrichot, V, Azar, D, Neraudeau, D, Nel, A, 2003. New Psocoptera in the Early Cretaceous amber of SW France and Lebanon (Insecta: Psocoptera: Trogiomorpha). Geological Magazine 140 (6): 669 - 683, DOI: https://doi.org/10.1017/S0016756803008355" title="New Psocoptera in the Early Cretaceous amber of SW France and Lebanon (Insecta: Psocoptera: Trogiomorpha)." url="https://doi.org/10.1017/S0016756803008355" volume="140" year="2003">Perrichot et al. 2003</bibRefCitation>
|
||
). However, a detailed re-examination has allowed us to distinguish two rows of setae along veins in all the type specimens. Therefore, the diagnosis must be emended replacing "setae arranged in one row on veins" with "setae arranged in two rows on veins" in forewing. The species
|
||
<taxonomicName id="106675A994588D9307AF369E934625C1" lsidName="P. axioperierga" pageId="0" pageNumber="183" rank="species" species="axioperierga">
|
||
<emphasis id="AC39A5954D45D1CD622C55232827330F" italics="true" pageId="0" pageNumber="183">P. axioperierga</emphasis>
|
||
</taxonomicName>
|
||
was originally described with two rows of setae along veins in forewing (
|
||
<bibRefCitation id="DA8403253F74818986756B7123745BAF" DOI="https://doi.org/10.17161/PC.1808.15983" author="Azar, D" journalOrPublisher="Geodiversitas" pageId="0" pageNumber="183" refId="B9" refString="Azar, D, Nel, A, Perrichot, V, 2014. Diverse barklice (Psocodea) fom Late Cretaceous Vendean amber. Paleontological Contributions 10C: 9-15. https://doi.org/10.17161/PC.1808.15983" title="Diverse barklice (Psocodea) fom Late Cretaceous Vendean amber. Paleontological Contributions 10 C: 9 - 15." url="https://doi.org/10.17161/PC.1808.15983" year="2014">Azar et al. 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="BB613DED267BAE0FD7DCE9599462730E" doi="10.3897/asp.82.e114849.figure5" httpUri="https://binary.pensoft.net/fig/1007225" pageId="0" pageNumber="183" start="Figure 5" startId="F5">
|
||
<paragraph id="42273FF249A5C82AE487100F9A47292E" pageId="0" pageNumber="183">
|
||
<emphasis id="612BD4B5A772DBA38EAF3F5D45406C5E" bold="true" pageId="0" pageNumber="183">Figure 5.</emphasis>
|
||
Forewings of
|
||
<taxonomicName id="B171E0FBFD3C668E7289543DAB2344AE" authorityName="Perrichot, Azar, Neraudeau & Nel" authorityYear="2003" class="Insecta" family="Archaeatropidae" genus="Proprionoglaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proprionoglaris guyoti" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="guyoti">
|
||
<emphasis id="79BD3430A71D80DC7F2B51FA697C4486" italics="true" pageId="0" pageNumber="183">Proprionoglaris guyoti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6469DA0924AFCB1170983E8908B517F6" authorityName="Azar, Nel & Perrichot" authorityYear="2014" class="Insecta" family="Archaeatropidae" genus="Proprionoglaris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proprionoglaris axioperierga" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="axioperierga">
|
||
<emphasis id="2FEE0DE6FA51D2E3D9772F0C40CA67E5" italics="true" pageId="0" pageNumber="183">Proprionoglaris axioperierga</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="A2C73D3DAACE2EA3B4C29B5DD4E597D7" class="Insecta" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
,
|
||
<taxonomicName id="9A3FF2E58140D731D30582F49AE216CF" class="Insecta" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="subOrder" subOrder="Trogiomorpha">Trogiomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="BEB6B599006406C728E3115014D5877D" authorityName="Baz & Ortuno" authorityYear="2000" class="Insecta" family="Empheriidae" kingdom="Animalia" lsidName="" order="Psocodea" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Empheriidae</taxonomicName>
|
||
).
|
||
<emphasis id="E49A6ED2D1B59003F1B6B545D88544CC" bold="true" pageId="0" pageNumber="183">A</emphasis>
|
||
holotype of
|
||
<taxonomicName id="EEF5BC79CC57F65AAD54C942FAA10968" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
|
||
<emphasis id="E7EC651EB6D19ED298C7C2CF8D102F03" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
|
||
</taxonomicName>
|
||
, female, MNHN.F.A30108 (ARC 58.2 R);
|
||
<emphasis id="BF90621AAB128EE9CD43F15FF7D494EC" bold="true" pageId="0" pageNumber="183">B</emphasis>
|
||
paratype of
|
||
<taxonomicName id="A128B96E0D5EDE98DBCEE99D2837AE28" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
|
||
<emphasis id="C69F121E75BF127B016182CFBBD682F3" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
|
||
</taxonomicName>
|
||
, male, MNHN.F.A30109 (ARC 201.2);
|
||
<emphasis id="9AC0B736DEEBA11E155FBB246303149A" bold="true" pageId="0" pageNumber="183">C</emphasis>
|
||
-
|
||
<emphasis id="084CAFECCF78860462606FE1D1C62F84" bold="true" pageId="0" pageNumber="183">E</emphasis>
|
||
new specimens of
|
||
<taxonomicName id="42249E3E3F2AC1D9C51A61C4870298AD" lsidName="P. guyoti" pageId="0" pageNumber="183" rank="species" species="guyoti">
|
||
<emphasis id="3270FD4178CBD5277DC6924DBD2B071F" italics="true" pageId="0" pageNumber="183">P. guyoti</emphasis>
|
||
</taxonomicName>
|
||
, sex unknown, IGR.ARC-352.1, IGR.ARC-157, and IGR.ARC-355, respectively;
|
||
<emphasis id="03E6B10C47C75D7B487961AA9C7D2263" bold="true" pageId="0" pageNumber="183">F</emphasis>
|
||
holotype of
|
||
<taxonomicName id="89B72196312D2D34B1BE91462EB48F06" lsidName="P. axioperierga" pageId="0" pageNumber="183" rank="species" species="axioperierga">
|
||
<emphasis id="6C1AE6B70EC12CAD6DCE245C7195A41A" italics="true" pageId="0" pageNumber="183">P. axioperierga</emphasis>
|
||
</taxonomicName>
|
||
, female, IGR.GAR-69;
|
||
<emphasis id="64CEEF9E5CFFFF3DDCBEF97DF20B5E2F" bold="true" pageId="0" pageNumber="183">G</emphasis>
|
||
paratype of
|
||
<taxonomicName id="24E2BA1DEE09D6996ED4BA1652E5F853" lsidName="P. axioperierga" pageId="0" pageNumber="183" rank="species" species="axioperierga">
|
||
<emphasis id="B0FFB35735689D3350A4BBC3B507B1A2" italics="true" pageId="0" pageNumber="183">P. axioperierga</emphasis>
|
||
</taxonomicName>
|
||
, sex unknown, IGR.GAR-39. White arrowheads: nodulus.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |