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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="3B4D1DDD1672500C89696130F7AE9AEB" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria tiberii subsp. alcamoensis Brandt 1961, comb. nov." docType="treatment" docVersion="5" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1732485878053" updateUser="ExternalLinkService">
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<mods:title id="00016E9C0C025242912EDF4CF2A6805C">Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:namePart id="3C17812B5196C8027A49E113BA9264A6">De Mattia, Willy</mods:namePart>
<mods:nameIdentifier id="5078905C48804C5C6CD4D03A7BDB8D2D" type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation id="2A403E1952859D06D00F12382F689035">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier id="7B997F5D1F52200612569E6C7AB2D642" type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:namePart id="DC4C8F038F54831B6158195B0D2E2619">Reier, Susanne</mods:namePart>
<mods:affiliation id="0687266A15558CF9F887E8D11D9CCC56">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:namePart id="AAEEE331F44A1708F518C60E9F7856B8">Haring, Elisabeth</mods:namePart>
<mods:affiliation id="6E24B355ADF696D2A8B13430B97BBBC5">Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:typeOfResource id="A942BE193F5F9F5A277A840178E31BD5">text</mods:typeOfResource>
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<mods:title id="D1C5999FFB8C9ACF2E311294EA1AE327">ZooKeys</mods:title>
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<mods:date id="F69544751F68703FF62413EEB0379962">2021</mods:date>
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<mods:number id="2B71136E8ED6C00526879CE817295E71">2021-12-14</mods:number>
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<mods:number id="FDA25827B4CE7516A177684E2AA44DFB">1077</mods:number>
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<mods:start id="8A2A7339390B95389C315042B7796BB4">1</mods:start>
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<mods:identifier id="F56385436A3D60ED9C864D1397E530F3" type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
<mods:identifier id="9F5DD1BB8AF7DC736DC81F03CD82899F" type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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<treatment id="3B4D1DDD1672500C89696130F7AE9AEB" ID-GBIF-Taxon="191645972" LSID="urn:lsid:plazi:treatment:3B4D1DDD1672500C89696130F7AE9AEB" httpUri="http://treatment.plazi.org/id/3B4D1DDD1672500C89696130F7AE9AEB" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Clausiliidae" scope_infraOrder="Clausilioidei" scope_order="Stylommatophora" scope_phylum="Mollusca" scope_superFamily="Clausilioidea">
<subSubSection id="B5637C2EABF25C791161D693C6DCE6F0" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="C0DC3365E1657CD50FE2284C86F254A9" pageId="0" pageNumber="1">
<taxonomicName id="28BD3706FFE4C1F2230DE0E56C3077E4" ID-CoL="8TH4S" LSID="3B4D1DDD-1672-500C-8969-6130F7AE9AEB" authority="Brandt, 1961" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" status="comb. nov." subSpecies="alcamoensis">Siciliaria tiberii alcamoensis Brandt, 1961</taxonomicName>
<taxonomicNameLabel id="F8248E43531105DDA65813BC9D8C68B3" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C4911399ED712C3C5C1A31F18B8E4116" pageId="0" pageNumber="1" type="description">
<paragraph id="0D97FBCC9ED0E577F003276C15F36FD3" pageId="0" pageNumber="1">
<figureCitation id="BBB6F0BF31C5C2484E3BAE1379C0DB0D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 25.3-25.5, 26.6-26.9, 29.4, 29.5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BC0FCB35DCEE1D9B504D70AB84B53723" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="96DA85152B0462DD51DB84429E6AC10F" pageId="0" pageNumber="1">
<taxonomicName id="A5B0AD28356CE65C804ECACEB38466CA" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" status="comb. nov." subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
<bibRefCitation id="E9D6A4AB51D8D643A61A68AA68D2C2A5" author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt 1961</bibRefCitation>
: 9.
</paragraph>
<paragraph id="160416B89A5ED95B3D293102A7598C3A" pageId="0" pageNumber="1">
<taxonomicName id="B1A4C7D7737D682F445E2C0BC024FF95" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subGenus="Siciliaria" subSpecies="alcamoensis">Siciliaria (Siciliaria) septemplicata alcamoensis</taxonomicName>
-
<bibRefCitation id="862A14F37CD1D22485D6410337582457" author="Manganelli, G" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" refId="B43" refString="Manganelli, G, Bodon, M, Favilli, L, Giusti, F, 1995. Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1-60. Calderini, Bologna." title="Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1 - 60. Calderini, Bologna." year="1995">Manganelli et al. 1995</bibRefCitation>
: 25.
</paragraph>
<paragraph id="D23807BB1555B991463EA14537351C9A" pageId="0" pageNumber="1">
<taxonomicName id="5F3D9B5E9F7063F0773EEA7FC2739378" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. hemmeni" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subSpecies="hemmeni">Siciliaria septemplicata hemmeni</taxonomicName>
<bibRefCitation id="4BB3BF4170D60FC0AAC9604E4B48CFE4" DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
: 190.
</paragraph>
<paragraph id="B4039A076ADAA209E2914307C9FB189E" pageId="0" pageNumber="1">
<taxonomicName id="FDB1A6585A437090A82E00C99F2DF9FC" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria (Siciliaria) septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subGenus="Siciliaria" subSpecies="alcamoensis">Charpentieria (Siciliaria) septemplicata alcamoensis</taxonomicName>
-
<bibRefCitation id="15CEDAFE752665D32E32A451BB450DBA" author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck 2007</bibRefCitation>
: 54.
</paragraph>
<paragraph id="4A69B1BB4BB809CD623E165099A71D74" pageId="0" pageNumber="1">
<taxonomicName id="74B7AC1E8BC6E0F09E9DC5D2F8A16B8A" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subSpecies="alcamoensis">Siciliaria septemplicata alcamoensis</taxonomicName>
-
<bibRefCitation id="DCC7467824FA5A202D58F68BF1892B64" author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
: 10.
</paragraph>
</subSubSection>
<subSubSection id="B1B165FFA9A501800E6710845F9E6118" pageId="0" pageNumber="1" type="examined specimens">
<paragraph id="18034AD0A413AFAF7FB170644CB7D233" pageId="0" pageNumber="1">Examined specimens.</paragraph>
<paragraph id="812CA33AD359B3B8E5BB4419E8A18F86" pageId="0" pageNumber="1">
Italy, Sicily, Cinisi, Piano Margi, 670 m asl,
<geoCoordinate id="2F9535921354DEEEE2315616717E683B" degrees="38" direction="north" minutes="08" orientation="latitude" precision="1" seconds="58.56" value="38.1496">38°08'58.56&quot;N</geoCoordinate>
,
<geoCoordinate id="9FC53F4FAE81190D847463E1F01625E6" degrees="13" direction="east" minutes="09" orientation="longitude" precision="1" seconds="25.83" value="13.157175">13°09'25.83&quot;E</geoCoordinate>
, [Lab ID 37_1, COI: MW758928, ITS2: MW757102, MW757101], W. De Mattia and J. Macor leg., 14.iv.2017. 2 dissected spm.
</paragraph>
</subSubSection>
<subSubSection id="B5A3F47D67008492A90BBAC103B6DBCB" pageId="0" pageNumber="1" type="shell">
<paragraph id="B47002CB94995ADE0EC27B33D6A8F34E" pageId="0" pageNumber="1">Shell</paragraph>
<paragraph id="61B4397253B05FECFEB08273A2BE6B5D" pageId="0" pageNumber="1">
<emphasis id="0F64D0CBC8B5DCD908C9C2F6C6257B54" bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation id="F12822708FFBAB751440B3EFE6F231D2" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
.6-26.9, 29.4-29.5).
</emphasis>
Shell decollate; whorls striated to rib-striated; dorsal keel very prominent; inferior lamella high; anterior upper palatal plica present, mostly separated from upper palatal plica, sometimes a second or also third upper palatal plica present; anterior lower palatal plica strong; palatal edge of clausilium plate distally not receding, palatal edge distally more or less strongly bent upwards.
</paragraph>
</subSubSection>
<subSubSection id="366CF6C61165F8A546C7F6F9BD2671A2" pageId="0" pageNumber="1" type="description">
<paragraph id="AAFC3F98DF1850C81C394B62651727BF" pageId="0" pageNumber="1">Measurements</paragraph>
<paragraph id="6560553CE003892645DF8236D85B341A" pageId="0" pageNumber="1">
(n = 18, decollate): shell height 21.5
<normalizedToken id="C2384C52A0A472EEB1BE5274D4B3133F" originalValue="±">+/-</normalizedToken>
1.3, whorl width 5.2
<normalizedToken id="9863279F68A023FD7B4E09C0641C5C4B" originalValue="±">+/-</normalizedToken>
0.3, aperture height 4.6
<normalizedToken id="86DD50E04DE8E45183D031CF83465B67" originalValue="±">+/-</normalizedToken>
0.2, aperture width 3.4
<normalizedToken id="F59AA99584283143DB5BF054672EA352" originalValue="±">+/-</normalizedToken>
0.4.
</paragraph>
</subSubSection>
<subSubSection id="0C4F40F97394B0ADCED3AF601B754A18" pageId="0" pageNumber="1" type="external morphology of the genital organs">
<paragraph id="46DA7CA96FF371937154457866EC00E4" pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
<paragraph id="FD38D4D2D87F3495FEF22328128E6702" pageId="0" pageNumber="1">
<emphasis id="2A56C7AF6E7CE5EF3BEEDA2951F3AFD9" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="207BD0E2CA267348943FA14AE19FFD0D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
.3).
</emphasis>
The FO is longer than the V (FO/V range 1.4-1.6). The VD is thin along its whole course. The FDBC is as long as the BC+SDBC (DBC/BC+SDBC = 1.0). The BC+SDBC is spindle-like to cylindrical and longer than the V (BC+SDBC/V range 1.4-1.5), with no distinction between the SDBC and the BC. The apex is pointed. The D is longer than the V (D/V range 2.5-2.6) and longer that the BC+SDBC (D/BC+SDBC range 1.6-1.8), thinner than the BC+SDBC and with a pointed apex. The V is long, cylindrical and small in diameter. The A is large. The PC is much longer than the V (P+E/V range 2.4-2.6). The PR is long and thin. The ET is clearly visible. There is a clear distinction between P and E, there is a visible ER, and a proximal narrowing. The E is longer than the P (E/P range 2.0-2.1), gradually shrinking and turning into the VD.
</paragraph>
</subSubSection>
<subSubSection id="4DC18ECE7E9AF92B9F0474059C3ED8DF" pageId="0" pageNumber="1" type="internal morphology of the genital organs">
<paragraph id="BF2593CDE7481DC67B64DB958FE3B1F5" pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
<paragraph id="DB6602BAA1DC4609B026BE8D93528C3C" pageId="0" pageNumber="1">
<emphasis id="110BCB9863D829615D6E5D5F8B9B25C8" bold="true" pageId="0" pageNumber="1">
(Figs
<figureCitation id="7D387A39CF44C7A83E78535E8AB501DB" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
.4-25.5).
</emphasis>
The A shows a set of irregular fleshy folds. The P presents two or three main large longitudinal pleats. These pleats are heavily transversely segmented, getting a comb-like shape. The fine structure of the wall is smooth.
</paragraph>
<paragraph id="E32B0E35263AEC042F4B0D22AFD12787" pageId="0" pageNumber="1">The PP is large, elongated, and heavily wrinkled. The P-E transition presents one ER with both the PP and ELP originating from the ER. The epiphallar formula is: 1ER(PP+ELP). The E shows a set of three or four smooth longitudinal pleats. The V is almost completely smooth. The wall shows a fine granulation.</paragraph>
</subSubSection>
<subSubSection id="A68E409C412736CAA89FA6563A7DE485" pageId="0" pageNumber="1" type="ecology">
<paragraph id="DCF7F125F80D1FB1EA7B8A6691F2903A" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="F53C31CF431B80CD5C5F84881A1A1E0C" pageId="0" pageNumber="1">
<taxonomicName id="4B0AB0F288DD2D55B0DA830D50714972" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="E57D8D30B3720237FFD5250B90184974" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
</taxonomicName>
comb. nov. inhabits scattered isolated limestone boulders, and is hiding in rock cracks and crevices. After careful field research in a wide area around Piano Margi, the taxon revealed to be remarkably rare and scattered.
</paragraph>
</subSubSection>
<subSubSection id="D2B27372082CA0DD87486D24876DF627" pageId="0" pageNumber="1" type="distribution">
<paragraph id="45A394D8357749192E2AFA772823AE2D" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="6AD684E0B27F7F727B7571CA9187DDDA" pageId="0" pageNumber="1">
<taxonomicName id="A46277AAFF2011F7B3CA85530A4AC21E" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="AAA4B8B2489181A365514C53C53E1E3C" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
</taxonomicName>
comb. nov. is known only from a small area of less than 1 km2 in the surrounding of the type locality: Piano Margi south of Cinisi on the Montagna Longa. Further field research is needed to define its actual distribution range.
</paragraph>
</subSubSection>
<subSubSection id="6F59C1B4BDDA7EDE872F8AF45CCC5008" pageId="0" pageNumber="1" type="remarks">
<paragraph id="DF810EA43853BAD719DFB94E8490445D" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="BF8D0A91C2C74E93D1CCBC9E5241D8C1" pageId="0" pageNumber="1">
<bibRefCitation id="744342251755ECCC9F0E1D69999A7A3C" author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
: 9) described
<taxonomicName id="6F9893F284BDC4C203BEA757CCF0F9BF" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
based on material from the Senckenberg Museum (SMF163949), provided by Monterosato and labelled as
<taxonomicName id="B9CCE182AEF1147233CA6EA039FFE024" authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
<emphasis id="8B4AFDCA65A06E0D2EAEEE21F545F92B" italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
</taxonomicName>
from &quot;Alcamo in NW-Sizilien, ohne
<normalizedToken id="C6898F13867B11929683C5126F8555C8" originalValue="nähere">naehere</normalizedToken>
Fundortangabe&quot;.
<bibRefCitation id="C9244F5DA6066DB14AB312A6D0364E2F" author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
: 9) introduced the name
<taxonomicName id="D662123EE57617034F08FA28D3299FAA" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis">
<emphasis id="CC865C8AC5B03070BA8A6DB4DABC9F8F" italics="true" pageId="0" pageNumber="1">Siciliaria alcamoensis</emphasis>
</taxonomicName>
as
<taxonomicName id="32D5D6D96FAB4BB09845877034BCA43B" authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
<emphasis id="D23CEFF6DE47AADF35FD8542CFE063FF" italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
</taxonomicName>
sensu Monterosato [non
<taxonomicName id="C2004A0D2F0F568222262F35E12061FB" authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
<emphasis id="18DBC5440D57282D6B1DE2ED9DF23F4D" italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
</taxonomicName>
Retowski, 1889, currently
<taxonomicName id="A0AB07177CA84FD8A22E83FF5AE7AFFA" baseAuthorityName="Retowski" baseAuthorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Roseniella" kingdom="Animalia" lsidName="Roseniella difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
<emphasis id="B940EB558030A9C4B6A64E6D448BFCCE" italics="true" pageId="0" pageNumber="1">Roseniella difficilis</emphasis>
</taxonomicName>
(Retowski, 1889)] is a nomen nudum.
<bibRefCitation id="9BD7DB62B748DC9979DCC4326DE88E7A" author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
: 10) stated that he was not able to find this species again during field collecting nearby Alcamo, but he also lists in the type material: &quot;Paratypoide Coll. Brandt Cl. 2365/2 leg. Belluci&quot;. He did not provide the collecting site of the paratype material so its origin is unknown, but it is conceivable that their shell morphology matches with what he described as
<taxonomicName id="D6406C97C365C043BE019FF865D5E0BC" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
indeed. Undoubtedly, the locality indication provided by Monterosato on the original SMF label is wrong. Despite intensive field research (WDM) all over the Alcamo area and Bosco
<normalizedToken id="F81C3435B726F44CB6EC816FB189005A" originalValue="d’Alcamo">d'Alcamo</normalizedToken>
/Monte Bonifato, no specimens/population was found that matches
<normalizedToken id="5EEEBCC9312C0AAD4544EE075D26B079" originalValue="Brandt’s">Brandt's</normalizedToken>
description of
<taxonomicName id="D32D0DCF384AF3F3B2C00E17E585ABA1" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis">
<emphasis id="A4ABABC617CE23543764ADC427B267A9" italics="true" pageId="0" pageNumber="1">Siciliaria alcamoensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="7141FCB390B997624141E6621A177B72" DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann (2004</bibRefCitation>
: 190) described
<taxonomicName id="FA97ADF8DB11BDFF4DB2BAA761637341" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. hemmeni" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" subSpecies="hemmeni">
<emphasis id="B65F219A6C798C549545B0C98FDE4742" italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata hemmeni</emphasis>
</taxonomicName>
from Piano Margi (Cinisi, Palermo). Its description and the pictures (
<bibRefCitation id="6D838022C0B43ECB66CD9C06793D5585" DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
: 189 and 190) match with
<normalizedToken id="FE262D90FD12DBE8A7CA2656C8B8E240" originalValue="Brandt’s">Brandt's</normalizedToken>
<taxonomicName id="FD1A3D2E422AFD6029EBA4E34C4AB3D8" authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
, as stated by
<bibRefCitation id="FC5296676C7D1C35C032693371867FD1" author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
: 7) and therefore, it is reasonable to assume that the Monterosato material (SMF163949) was collected from the same area of
<normalizedToken id="649E5697E8268C565BE94B72FFEBA754" originalValue="Beckmann’s">Beckmann's</normalizedToken>
species.
</paragraph>
<paragraph id="AACC45CD3FFC6A21D52740F7704C96F0" pageId="0" pageNumber="1">
<taxonomicName id="FAA81897A701DFFE0C8EA2ADADBCF3BE" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="8B31B8EB5ACE71ADBDA6EE069F66578F" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
</taxonomicName>
comb. nov. (Figs
<figureCitation id="ABDC2381D31271495AD3C162585D2A8D" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
.6-26.9) was treated as a subspecies of
<taxonomicName id="FE463278CAC8F25848AD2541B2087A0C" lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
<emphasis id="AD57906D9FC8C2DB4B9BF9FB69A484BC" italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
</taxonomicName>
(MolluscaBase 2021). This is due to the fact that the typical form of
<taxonomicName id="094B265422B2B687D58E4BBFD9610C05" lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
<emphasis id="EE099982714AB01BA3BC385D81254BBC" italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
</taxonomicName>
(e.g., Monte Gallo populations, Figs
<figureCitation id="8CCCC1BFA3074C36AAE5A4D07E9BE28F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siciliaria septemplicata (Philippi, 1836), Monte Gallo, Sferracavallo 10.1 shell 10.2 detail of the aperture 10.3 clausiliar plate 10.4 shell 10.5 detail of the aperture 10.6 detail of the columellar side of last whorl 10.7 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure10" httpUri="https://binary.pensoft.net/fig/622802" pageId="0" pageNumber="1">10</figureCitation>
.1-10.7), is quite similar regarding the morphology of the clausiliar apparatus, (
<bibRefCitation id="A319F0B6EBC4C62E3DF102F87ABADAA5" author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
), despite they greatly differ in the shell surface, sutures ornamentation, colouration, thickness and dimensions. The genital morphology (Figs
<figureCitation id="773CC43267C13ADBC1038BE4B6BBB931" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
.3-25.4) of
<taxonomicName id="088D07A25F5A832289580DE18B0F9D9C" lsidName="S. tiberii subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="0C88964B1250A62894B9926C8CC857BB" italics="true" pageId="0" pageNumber="1">S. tiberii alcamoensis</emphasis>
</taxonomicName>
gretaly differs from that of
<taxonomicName id="1C976277C7FF15B265CF090183D19F9A" lsidName="S. s. subsp. septemplicata" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="septemplicata">
<emphasis id="14BFD4C028ADA6486A3405712775A2BF" italics="true" pageId="0" pageNumber="1">S. s. septemplicata</emphasis>
</taxonomicName>
(Figs
<figureCitation id="78CB07711B0FFEF2CE99F860DE9E7045" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Siciliaria septemplicata (Philippi, 1836), Monte Gallo, Sferracavallo 9.1 whole distal genital organs 9.2 internal distal part of genital organs 9.3 whole distal genital organs 9.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure9" httpUri="https://binary.pensoft.net/fig/622801" pageId="0" pageNumber="1">9</figureCitation>
.1-9.2). In the COI and concatenated trees,
<taxonomicName id="A790C48488B2E9B5A5D7DC560479DBAC" lsidName="S. t. subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="t." subSpecies="alcamoensis">
<emphasis id="A0F4C639000BA842804672043A21CCF0" italics="true" pageId="0" pageNumber="1">S. t. alcamoensis</emphasis>
</taxonomicName>
is is well embedded within the
<taxonomicName id="7EBF4551C1ABD192FC6D953C86883A4F" authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">
<emphasis id="FFE0D3B205265CB56093B45694C3471F" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii</emphasis>
</taxonomicName>
clade. Since it is found well within the distribution range of
<taxonomicName id="FCFAF8580E7362FA2F0ACBF4B47D60F6" lsidName="S. tiberii" pageId="0" pageNumber="1" rank="species" species="tiberii">
<emphasis id="C39704301AF05F39E08C589CB90575FD" italics="true" pageId="0" pageNumber="1">S. tiberii</emphasis>
</taxonomicName>
s. l., introgression due to hybridisation appears as an unlikely explanation. It might be better explained by incomplete lineage sorting. Possible interspecies hybridisation should be tested on a larger sample size with additional nuclear markers. There is no argument for considering
<taxonomicName id="52011A2A4175C18B4436926535D90C66" lsidName="S. s. subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="alcamoensis">
<emphasis id="8481B76B29AEE568586A0DE1866FED9E" italics="true" pageId="0" pageNumber="1">S. s. alcamoensis</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="FA67E4F306309DA71218FE07F8DB393F" lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
<emphasis id="74C09A7537AF23C97B2E87FBF1078296" italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
</taxonomicName>
, we here consider it a subspecies
<taxonomicName id="6171700DB48FFA00F63926A5A51303DE" lsidName="S. tiberii" pageId="0" pageNumber="1" rank="species" species="tiberii">
<emphasis id="521803D4C84C207BC05CA6AD0A5CFC1A" italics="true" pageId="0" pageNumber="1">S. tiberii</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="4DB04BF3524640F0A4B183C5005C30FA" doi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
<paragraph id="CC81DFCDF20C7429C515AC9336A9F844" pageId="0" pageNumber="1">
<emphasis id="F9AFCBFB06FB5DBB029BDF7C2F9E089C" bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
<taxonomicName id="34B09FC6A9BE53B020625A81BEC8EE55" authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis id="B67BA9290C4BB8BB85483F9E334C58D3" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
(A. Schmidt, 1868), Capo Rama, Terrasini
<emphasis id="BB308B04C9B0CBC6F8BA07A2B436DBA4" bold="true" pageId="0" pageNumber="1">25.1</emphasis>
whole distal genital organs
<emphasis id="C5DEBA7AFDC724839E2DD5D98FFE94D6" bold="true" pageId="0" pageNumber="1">25.2</emphasis>
internal distal part of genital organs.
<taxonomicName id="5F11EC99652A8165C4D52626B829CB66" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="3E24FBDECA68A788CCC7638E6C49B0FF" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
</taxonomicName>
Brandt, 1961 comb. nov., Piano Margi, Carini
<emphasis id="197D51CABE461FE9FB6753B58DF45B89" bold="true" pageId="0" pageNumber="1">25.3</emphasis>
whole distal genital organs
<emphasis id="BD77E48CAAAA87FD4135746DAA678FBE" bold="true" pageId="0" pageNumber="1">25.4</emphasis>
internal distal part of genital organs
<emphasis id="8F1192F7710C27BA278EFC201F92F433" bold="true" pageId="0" pageNumber="1">25.5</emphasis>
detail of the pseudopapilla.
</paragraph>
</caption>
<caption id="C669D3E3B6FE6B04E08095ADE901881F" doi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
<paragraph id="02DA9497B84EA04610FD245D026F51CA" pageId="0" pageNumber="1">
<emphasis id="9656B278D4C62EAC4155A885405C051F" bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
<taxonomicName id="01C4A92096D9A0A5BD36D299459B17D2" authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
<emphasis id="BA1AD6078D1A5E441DCAEB55BFC0E466" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
</taxonomicName>
(A. Schmidt, 1868), Capo Rama, Terrasini
<emphasis id="001D49CB8EA096B01B549EFDB9D79E23" bold="true" pageId="0" pageNumber="1">26.1</emphasis>
shell
<emphasis id="F7CFCA6AB3A28EDE3476AFBEDBEC6E30" bold="true" pageId="0" pageNumber="1">26.2</emphasis>
clausiliar plate double side
<emphasis id="2BBF20060A4D8758F140C35237D88A53" bold="true" pageId="0" pageNumber="1">26.3</emphasis>
shell
<emphasis id="2DBC84CD80F1ED37770D4CB67F7E2BAC" bold="true" pageId="0" pageNumber="1">26.4</emphasis>
detail of the aperture
<emphasis id="1F08126055EB9A04A65C9F5FEC67FBAD" bold="true" pageId="0" pageNumber="1">26.5</emphasis>
clausiliar plate double side.
<taxonomicName id="659EBCD60DC8A794AA2AC6B72AADE4AE" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia &amp; Reier &amp; Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
<emphasis id="E0402934DF3C3653A02831B277D5575A" italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
</taxonomicName>
Brandt, 1961 comb. nov., Piano Margi, Carini
<emphasis id="18E10BF72ED65897B38AC5686868BDD9" bold="true" pageId="0" pageNumber="1">26.6</emphasis>
shell
<emphasis id="07A72A57D6B3FF5BB834588647BBF080" bold="true" pageId="0" pageNumber="1">26.7</emphasis>
clausiliar plate double side
<emphasis id="2F8FDACB4275A8B667C4609F492358F2" bold="true" pageId="0" pageNumber="1">26.8</emphasis>
detail of the aperture
<emphasis id="E9D7FB0D3C2E567AB335070626A7F177" bold="true" pageId="0" pageNumber="1">26.9</emphasis>
shell.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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