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<document id="25A447FECF92A011CCF5E71C90816C7A" ID-CLB-Dataset="6525" ID-DOI="10.11646/zootaxa.4966.3.11" ID-GBIF-Dataset="78e853f4-59a8-4716-ae9a-662cc2ada33b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4736618" ID-ZooBank="DBC63623-352A-41DF-A4FE-209BEC83FCBA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620119574882" checkinUser="plazi" docAuthor="Archibald, S. Bruce &amp; Cannings, Robert A." docDate="2021" docId="3A4187FEFFA2FF9CADB1EBB68441FAEB" docLanguage="en" docName="zootaxa.4966.3.11.pdf" docOrigin="Zootaxa 4966 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Republica Archibald &amp; Cannings 2021, new genus" docType="treatment" docVersion="8" lastPageNumber="396" masterDocId="C678FF86FFA1FF98AD26EB21872EFFBE" masterDocTitle="A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U. S. A." masterLastPageNumber="400" masterPageNumber="392" pageNumber="395" updateTime="1698938302332" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="738994C34DCBA7FCD5F855719516745B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E52B889A368AD7EF557A2420583C5836">
<mods:title id="3E2E065DFDA0E141A0EE3F4D50A4367D">A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U. S. A.</mods:title>
</mods:titleInfo>
<mods:name id="62A4E7CD4DE4A48AD8D733D3D9AFC935" type="personal">
<mods:role id="D752B886723D66A59C785B0AB9547D7D">
<mods:roleTerm id="071567598DEE7F6AC95CE97EDAD9BC87">Author</mods:roleTerm>
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<mods:namePart id="69A275A022942A9B9D9A1E3108520165">Archibald, S. Bruce</mods:namePart>
<mods:affiliation id="273DDA4A13601B6DC7072243F22D9375">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada. Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America. &amp; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada.</mods:affiliation>
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<mods:roleTerm id="DA5A47294F12257AC9656C698CE388AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E1F2D0643BC3B20D708ACF3E1ADF616">Cannings, Robert A.</mods:namePart>
<mods:affiliation id="C8EF02B25155FB63A1AFAF099EBCEEB1">Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada.</mods:affiliation>
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<mods:title id="FEC18D5CD500AEFBB17D7B11EA15BA14">Zootaxa</mods:title>
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<mods:part id="A3D2539D0946470797525A23C254525D">
<mods:date id="1329C750A25FA170134E5CCE27F34C37">2021</mods:date>
<mods:detail id="873E9A720F50807F730BFCA79897257D" type="pubDate">
<mods:number id="A09217EB6E9D7B043C0F89DB998832EC">2021-05-04</mods:number>
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<mods:detail id="E5E673058D3B7D63D6FF008FDE61FE6D" type="volume">
<mods:number id="C481DEF482D513CE8631CAE618256B56">4966</mods:number>
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<mods:detail id="DFFEC83C6EE7E262A5CA639A4FFC7C62" type="issue">
<mods:number id="28713FE3F39DEFCF0ECCA7441E159B43">3</mods:number>
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<mods:classification id="09822730D3CC9A6A553E44DF6533BFD5">journal article</mods:classification>
<mods:identifier id="4799C4B37B6E984A92F24797E46B3141" type="CLB-Dataset">6525</mods:identifier>
<mods:identifier id="94463A853B33084EBFE072CCC09DF17B" type="DOI">10.11646/zootaxa.4966.3.11</mods:identifier>
<mods:identifier id="1EB216C795C9F381D3655CDC43A4D8A4" type="GBIF-Dataset">78e853f4-59a8-4716-ae9a-662cc2ada33b</mods:identifier>
<mods:identifier id="FF0E176FE86C8EB28EBBCF543F8DD145" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB8B95A101EA0701FA7D9BA3AA52015F" type="Zenodo-Dep">4736618</mods:identifier>
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<treatment id="3A4187FEFFA2FF9CADB1EBB68441FAEB" ID-DOI="http://doi.org/10.5281/zenodo.4784029" ID-GBIF-Taxon="182384030" ID-Zenodo-Dep="4784029" LSID="urn:lsid:plazi:treatment:3A4187FEFFA2FF9CADB1EBB68441FAEB" httpUri="http://treatment.plazi.org/id/3A4187FEFFA2FF9CADB1EBB68441FAEB" lastPageId="4" lastPageNumber="396" pageId="3" pageNumber="395">
<subSubSection id="FAF26563FFA2FF9BADB1EBB685EAFF0F" box="[151,708,151,178]" pageId="3" pageNumber="395" type="nomenclature">
<paragraph id="B25736E8FFA2FF9BADB1EBB685EAFF0F" blockId="3.[151,708,151,213]" box="[151,708,151,178]" pageId="3" pageNumber="395">
<heading id="E91F8184FFA2FF9BADB1EBB685EAFF0F" bold="true" box="[151,708,151,178]" fontSize="11" level="1" pageId="3" pageNumber="395" reason="1">
<emphasis id="809CEAFAFFA2FF9BADB1EBB685EAFF0F" bold="true" box="[151,708,151,178]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1EBB68518FF0F" authority="Archibald &amp; Cannings" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[151,566,151,178]" class="Magnoliopsida" family="Euphaeidae" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="809CEAFAFFA2FF9BADB1EBB6863FFF0C" bold="true" box="[151,273,151,178]" italics="true" pageId="3" pageNumber="395">Republica</emphasis>
Archibald &amp; Cannings
</taxonomicName>
,
<taxonomicNameLabel id="9BAF5781FFA2FF9BAF64EBB685EAFF0F" box="[578,708,151,177]" pageId="3" pageNumber="395" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FAF26563FFA2FF9BADB1EB9A87F7FF6B" box="[151,217,187,213]" pageId="3" pageNumber="395" type="description">
<paragraph id="B25736E8FFA2FF9BADB1EB9A87F7FF6B" blockId="3.[151,708,151,213]" box="[151,217,187,213]" pageId="3" pageNumber="395">
<figureCitation id="2AD32A6DFFA2FF9BADB1EB9A87F7FF6B" box="[151,217,187,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1306,1330]" captionTargetBox="[151,1428,498,1270]" captionTargetId="figure-168@5.[151,1436,489,1282]" captionTargetPageId="5" captionText="FIGURE 1. Republica weatbrooki Holotype SR 06-59-08. A, photograph; B, drawing. Both to scale, 5 mm; db, discoidal bracket, see text." figureDoi="http://doi.org/10.5281/zenodo.4736622" httpUri="https://zenodo.org/record/4736622/files/figure.png" pageId="3" pageNumber="395">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FAF26563FFA2FF9BADB1EA228361F9B4" pageId="3" pageNumber="395" type="diagnosis">
<paragraph id="B25736E8FFA2FF9BADB1EA228502FEFF" blockId="3.[151,1437,259,321]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADB1EA22863CFEA3" bold="true" box="[151,274,259,285]" pageId="3" pageNumber="395">Diagnosis.</emphasis>
The following character states of the
<taxonomicName id="75E84D6BFFA2FF9BAF92EA228408FEA3" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[692,806,259,285]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA2FF9BAF92EA228408FEA3" box="[692,806,259,285]" italics="true" pageId="3" pageNumber="395">Republica</emphasis>
</taxonomicName>
forewing (preseumed, see below) most easily separate it from those of other
<taxonomicName id="75E84D6BFFA2FF9BAC50EA068507FEFF" authorityName="Cockerell" authorityYear="1923" box="[374,553,295,321]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="tribe" tribe="Eodichromatini">Eodichromatini</taxonomicName>
:
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EA4E827CFE37" blockId="3.[151,1437,367,825]" box="[151,1362,367,394]" pageId="3" pageNumber="395">1- narrow, 3.9 times longer than maximum width (L/W); nodus to apex length 2.5 maximum width (N/W);</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EAB28515FE13" blockId="3.[151,1437,367,825]" box="[151,571,403,430]" pageId="3" pageNumber="395">2- arculus near to, just basal to Ax2;</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EA968660FE48" blockId="3.[151,1437,367,825]" pageId="3" pageNumber="395">3- many (at least 28) antenodal crossveins preserved in first row, (at least 26) Ax2 to nodus; not aligned with second row;</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EADE8441FDA7" blockId="3.[151,1437,367,825]" box="[151,879,511,538]" pageId="3" pageNumber="395">4- costal space distal to pterostigma maximum three cells wide;</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E902866CFDDF" blockId="3.[151,1437,367,825]" pageId="3" pageNumber="395">5- both rows of antenodal, postnodal crossveins with numerous branching, some connected by horizontal crossveins;</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E94A8404FD3B" blockId="3.[151,1437,367,825]" box="[151,810,619,646]" pageId="3" pageNumber="395">6- one secondary antenodal crossvein between Ax1, Ax2.</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E9B18638FD70" blockId="3.[151,1437,367,825]" pageId="3" pageNumber="395">7- six long secondary longitudinal veins in CuA–A space (region not well preserved in part, but six appears most likely);</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E9F68395FD4F" blockId="3.[151,1437,367,825]" box="[151,1211,727,754]" pageId="3" pageNumber="395">8- MA smoothly, sinuately curved twice toward anterior margin and MP, CuA deeply curved;</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E9DA8421FC87" blockId="3.[151,1437,367,825]" pageId="3" pageNumber="395">9- MP, CuA sub-parallel, remain close at termination on margin (where preserved, highly likely throughout) without secondary longitudinal veins between them.</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E8468534FC3F" blockId="3.[151,538,871,897]" box="[151,538,871,897]" pageId="3" pageNumber="395">
These distinguish
<taxonomicName id="75E84D6BFFA2FF9BAC45E84686FBFC3F" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[355,469,871,897]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA2FF9BAC45E84686FBFC3F" box="[355,469,871,897]" italics="true" pageId="3" pageNumber="395">Republica</emphasis>
</taxonomicName>
from:
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E88E8354FC77" blockId="3.[151,1436,942,1293]" box="[151,1146,942,970]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1E88E86BBFC77" authority="Cockerell" authorityName="Cockerell" authorityYear="1923" box="[151,405,943,969]" class="Insecta" family="Epallagidae" genus="Eodichroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BADB1E88E860EFC77" box="[151,288,943,969]" italics="true" pageId="3" pageNumber="395">Eodichroma</emphasis>
Cockerell
</taxonomicName>
by 3 [13 Ax2 to nodus], 5 [simple], 7, [less, apparently 3–4], 8, 9;
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1E8F28418FBAF" blockId="3.[151,1436,942,1293]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1E8F28644FC53" authority="Nel" authorityName="Nel" authorityYear="1988" box="[151,362,979,1005]" class="Insecta" family="Epallagidae" genus="Parazacallites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BADB1E8F28617FC53" box="[151,313,979,1005]" italics="true" pageId="3" pageNumber="395">Parazacallites</emphasis>
Nel
</taxonomicName>
by 2 (near Ax1), 3 [11, aligned], 5 [simple], 6 [2], 7 [less: 3 preserved, perhaps another?], 9 [diverge, but no secondary longitudinal veins between];
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EF3A84D6FBE7" blockId="3.[151,1436,942,1293]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1EF3A867EFB8B" authority="Zessin" authorityName="Zessin" authorityYear="2011" box="[151,336,1051,1077]" class="Insecta" family="Epallagidae" genus="Solveigia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BADB1EF3A87D1FB8B" box="[151,255,1051,1077]" italics="true" pageId="3" pageNumber="395">Solveigia</emphasis>
<materialsCitation id="02803CB5FFA2FF9BAC21EF3A867EFB8B" ID-GBIF-Occurrence="3117081305" box="[263,336,1051,1077]" collectionCode="L, W, N" longitude="-2.0" pageId="3" pageNumber="395" specimenCount="1">Zessin</materialsCitation>
</taxonomicName>
by 1 [forewing L/W ~3.0, N/
<geoCoordinate id="D7DC502FFFA2FF9BAFBAEF3A85CBFB8B" box="[668,741,1050,1077]" degrees="2.0" direction="west" orientation="longitude" pageId="3" pageNumber="395" precision="5555" value="-2.0">W 2.0</geoCoordinate>
], 3 [7 preserved, probably 1–2 more, second row 2 preserved, aligned], 4 [1 cell wide], 5 [simple], 6 [none], 7 [less: apparently 3], 8, 9;
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EF4284F0FBC3" blockId="3.[151,1436,942,1293]" box="[151,990,1122,1150]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1EF428677FBC3" authority="Zessin" authorityName="Zessin" authorityYear="2011" box="[151,345,1123,1149]" class="Insecta" family="Epallagidae" genus="Ejerslevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BADB1EF428627FBC3" box="[151,265,1123,1149]" italics="true" pageId="3" pageNumber="395">Ejerslevia</emphasis>
Zessin
</taxonomicName>
by 4 [1 cell wide], 5 [simple], 7 [less, apparently 4], 8, 9;
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EFA68394FB1F" blockId="3.[151,1436,942,1293]" box="[151,1210,1158,1186]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADB1EFA6864EFB1F" box="[151,352,1159,1185]" italics="true" pageId="3" pageNumber="395">Wolfgangeuphaea</emphasis>
Ferwer &amp; Nel: 2 [near Ax1], 4 [1 cell wide], 5 [simple], 9 [diverge distally];
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EF8A83A6FB7B" blockId="3.[151,1436,942,1293]" box="[151,1160,1194,1222]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADB1EF8A8526FB78" authority="Petrulevicius et al." authorityName="Petrulevicius" box="[151,520,1195,1222]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BADB1EF8A8601FB7B" box="[151,303,1195,1221]" italics="true" pageId="3" pageNumber="395">Labandeiraia</emphasis>
Petrulevičius
<emphasis id="809CEAFAFFA2FF9BACF6EF8D852FFB7B" box="[464,513,1195,1221]" italics="true" pageId="3" pageNumber="395">et al</emphasis>
.
</taxonomicName>
by 5 [see discussion], 9 [
<taxonomicName id="75E84D6BFFA2FF9BAE03EF8D84AFFB7B" box="[805,897,1196,1221]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA2FF9BAE03EF8D84AFFB7B" box="[805,897,1196,1221]" italics="true" pageId="3" pageNumber="395">L. riveri</emphasis>
</taxonomicName>
, see notes, below] and:
</paragraph>
<paragraph id="B25736E8FFA2FF9BADE0EFF18331FB57" blockId="3.[151,1436,942,1293]" box="[198,1055,1230,1258]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADE0EFF1855CFB54" authority="Petrulevicius et al." authorityName="Petrulevicius" box="[198,626,1231,1258]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="americaborealis">
<emphasis id="809CEAFAFFA2FF9BADE0EFF186B4FB57" box="[198,410,1231,1257]" italics="true" pageId="3" pageNumber="395">L. americaborealis</emphasis>
Petrulevičius
<emphasis id="809CEAFAFFA2FF9BAF1CEFF18545FB57" box="[570,619,1231,1257]" italics="true" pageId="3" pageNumber="395">et al</emphasis>
.
</taxonomicName>
and
<taxonomicName id="75E84D6BFFA2FF9BAF8FEFF18484FB54" authority="Bechly et al." authorityName="Bechly" box="[681,938,1231,1258]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="809CEAFAFFA2FF9BAF8FEFF1843AFB57" box="[681,788,1231,1257]" italics="true" pageId="3" pageNumber="395">L. browni</emphasis>
Bechly
<emphasis id="809CEAFAFFA2FF9BAE54EFF1848DFB57" box="[882,931,1231,1257]" italics="true" pageId="3" pageNumber="395">et al</emphasis>
.
</taxonomicName>
: 6 [none];
</paragraph>
<paragraph id="B25736E8FFA2FF9BADE0EFD5840DFAB3" blockId="3.[151,1436,942,1293]" box="[198,803,1266,1293]" pageId="3" pageNumber="395">
<taxonomicName id="75E84D6BFFA2FF9BADE0EFD58698FAB3" authority="Bechly et al." authorityName="Bechly" box="[198,438,1267,1293]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA2FF9BADE0EFD5860FFAB3" box="[198,289,1268,1293]" italics="true" pageId="3" pageNumber="395">L. riveri</emphasis>
<materialsCitation id="02803CB5FFA2FF9BAC0FEFD28698FAB3" ID-GBIF-Occurrence="3117081302" box="[297,438,1267,1293]" collectionCode="L" longitude="-3.4" pageId="3" pageNumber="395" specimenCount="1">
Bechly
<emphasis id="809CEAFAFFA2FF9BAC59EFD5869EFAB3" box="[383,432,1267,1293]" italics="true" pageId="3" pageNumber="395">et al</emphasis>
.
</materialsCitation>
</taxonomicName>
:1 [forewing L/
<geoCoordinate id="D7DC502FFFA2FF9BAF44EFD28585FAB3" box="[610,683,1266,1293]" degrees="3.4" direction="west" orientation="longitude" pageId="3" pageNumber="395" precision="5555" value="-3.4">W 3.4</geoCoordinate>
], 6 [none].
</paragraph>
<paragraph id="B25736E8FFA2FF9BADB1EE1A8361F9B4" blockId="3.[151,1436,1339,1546]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADB1EE1A87F6FAEB" bold="true" box="[151,216,1339,1365]" pageId="3" pageNumber="395">Note:</emphasis>
some character states cannot be compared when only the hind wing is known due to forewing-hind wing differences or where the relevant region is missing,
<emphasis id="809CEAFAFFA2FF9BAFCEEE418427FAC7" box="[744,777,1376,1401]" italics="true" pageId="3" pageNumber="395">e.g</emphasis>
., the distal half of the
<taxonomicName id="75E84D6BFFA2FF9BA928EE7E839EFAC7" authorityName="Nel" authorityYear="1988" box="[1038,1200,1375,1401]" class="Insecta" family="Epallagidae" genus="Parazacallites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA2FF9BA928EE7E839EFAC7" box="[1038,1200,1375,1401]" italics="true" pageId="3" pageNumber="395">Parazacallites</emphasis>
</taxonomicName>
wing; others that do not appear in descriptions and were not illustrated cannot be assessed,
<emphasis id="809CEAFAFFA2FF9BAE91EEA584F7FA23" box="[951,985,1412,1437]" italics="true" pageId="3" pageNumber="395">e.g</emphasis>
., character state
<quantity id="75109B0DFFA2FF9BA9BEEEA283EAFA23" box="[1176,1220,1411,1437]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="3" pageNumber="395" unit="in" value="8.0">8 in</quantity>
<emphasis id="809CEAFAFFA2FF9BA9EAEEA282BBFA23" box="[1228,1429,1411,1437]" italics="true" pageId="3" pageNumber="395">Wolfgangeuphaea</emphasis>
, character state
<quantity id="75109B0DFFA2FF9BAC1BEE868649FA7F" box="[317,359,1447,1474]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="395" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="75E84D6BFFA2FF9BAC4BEE8986E9FA7F" box="[365,455,1448,1473]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA2FF9BAC4BEE8986E9FA7F" box="[365,455,1448,1473]" italics="true" pageId="3" pageNumber="395">L. riveri</emphasis>
</taxonomicName>
, and the presence of secondary crossveins between Ax1 and Ax2 and the general density of secondary longitudinal veins in
<taxonomicName id="75E84D6BFFA2FF9BAF30EEED85AFFA5B" box="[534,641,1483,1509]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="809CEAFAFFA2FF9BAF30EEED85AFFA5B" box="[534,641,1483,1509]" italics="true" pageId="3" pageNumber="395">L. browni</emphasis>
</taxonomicName>
and
<taxonomicName id="75E84D6BFFA2FF9BAF93EEED843EFA5B" box="[693,784,1484,1509]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA2FF9BAF93EEED843EFA5B" box="[693,784,1484,1509]" italics="true" pageId="3" pageNumber="395">L. riveri</emphasis>
</taxonomicName>
(
<taxonomicName id="75E84D6BFFA2FF9BAE39EEED84A4FA5B" box="[799,906,1483,1509]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="browni">
<emphasis id="809CEAFAFFA2FF9BAE39EEED84A4FA5B" box="[799,906,1483,1509]" italics="true" pageId="3" pageNumber="395">L. browni</emphasis>
</taxonomicName>
contradicts character state 9 from the published photograph; however, the published photograph of
<taxonomicName id="75E84D6BFFA2FF9BAFF3EED1841EF9B7" box="[725,816,1520,1545]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA2FF9BAFF3EED1841EF9B7" box="[725,816,1520,1545]" italics="true" pageId="3" pageNumber="395">L. riveri</emphasis>
</taxonomicName>
is unclear in this regard).
</paragraph>
</subSubSection>
<subSubSection id="FAF26563FFA2FF9BADE1ED1684BBF9EC" box="[199,917,1591,1618]" pageId="3" pageNumber="395" type="type_taxon">
<paragraph id="B25736E8FFA2FF9BADE1ED1684BBF9EC" blockId="3.[151,1437,1591,2014]" box="[199,917,1591,1618]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADE1ED168675F9EF" bold="true" box="[199,347,1591,1617]" pageId="3" pageNumber="395">
<typeStatus id="6D53884AFFA2FF9BADE1ED16862CF9EF" box="[199,258,1591,1617]" pageId="3" pageNumber="395">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="75E84D6BFFA2FF9BAC4FED168571F9EF" authority="Archibald &amp; Cannings, 2021" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[361,607,1591,1617]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="species" species="weatbrooki" status="sp. nov.">
<emphasis id="809CEAFAFFA2FF9BAC4FED168571F9EF" box="[361,607,1591,1617]" italics="true" pageId="3" pageNumber="395">Republica weatbrooki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9BAF5781FFA2FF9BAF40ED1985C1F9EF" box="[614,751,1591,1618]" pageId="3" pageNumber="395" rank="species">new species</taxonomicNameLabel>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="FAF26563FFA2FF9BADE1ED7A8571F9CB" box="[199,607,1627,1653]" pageId="3" pageNumber="395" type="description">
<paragraph id="B25736E8FFA2FF9BADE1ED7A8571F9CB" blockId="3.[151,1437,1591,2014]" box="[199,607,1627,1653]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADE1ED7A8677F9CB" bold="true" box="[199,345,1627,1653]" pageId="3" pageNumber="395">Description.</emphasis>
As for its only species.
</paragraph>
</subSubSection>
<subSubSection id="FAF26563FFA2FF9CADE1ED5E8441FAEB" lastPageId="4" lastPageNumber="396" pageId="3" pageNumber="395" type="etymology">
<paragraph id="B25736E8FFA2FF9BADE1ED5E869FF903" blockId="3.[151,1437,1591,2014]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADE1ED5E8666F927" bold="true" box="[199,328,1663,1689]" pageId="3" pageNumber="395">Etymology</emphasis>
. The genus name is the Latinized form of
<emphasis id="809CEAFAFFA2FF9BAE03ED5E84A7F927" box="[805,905,1663,1689]" italics="true" pageId="3" pageNumber="395">Republic</emphasis>
, the town where the only known specimen was found. Gender, feminine.
</paragraph>
<paragraph id="B25736E8FFA2FF9BADE1EDE6854DF8BB" blockId="3.[151,1437,1591,2014]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADE1EDE68652F95F" bold="true" box="[199,380,1735,1761]" pageId="3" pageNumber="395">Range and age</emphasis>
.
<materialsCitation id="02803CB5FFA2FF9BACABEDE98571F8BB" ID-GBIF-Occurrence="3117081304" collectionCode="B" country="United States of America" county="Republic" location="Tom Thumb Tuff Member" municipality="Mountain Formation" pageId="3" pageNumber="395" specimenCode="B4131" specimenCount="1" stateProvince="Washington">
<location id="B7376033FFA2FF9BACABEDE98593F95F" LSID="urn:lsid:plazi:treatment:3A4187FEFFA2FF9CADB1EBB68441FAEB:B7376033FFA2FF9BACABEDE98593F95F" box="[397,701,1735,1762]" country="United States of America" county="Republic" municipality="Mountain Formation" name="Tom Thumb Tuff Member" pageId="3" pageNumber="395" stateProvince="Washington">Tom Thumb Tuff Member</location>
of the
<location id="B7376033FFA2FF9BAE33EDE6835DF95F" LSID="urn:lsid:plazi:treatment:3A4187FEFFA2FF9CADB1EBB68441FAEB:B7376033FFA2FF9BAE33EDE6835DF95F" box="[789,1139,1735,1761]" country="United States of America" county="Republic" municipality="Mountain Formation" name="Klondike Mountain Formation" pageId="3" pageNumber="395" stateProvince="Washington">
<location id="B7376033FFA2FF9BAE33EDE684DAF95F" LSID="urn:lsid:plazi:treatment:3A4187FEFFA2FF9CADB1EBB68441FAEB:B7376033FFA2FF9BAE33EDE684DAF95F" box="[789,1012,1735,1761]" country="United States of America" county="Republic" municipality="Mountain Formation" name="Klondike Mountain" pageId="3" pageNumber="395" stateProvince="Washington">
Klondike
<collectingMunicipality id="5233AC92FFA2FF9BAEA1EDE684DAF95F" box="[903,1012,1735,1761]" pageId="3" pageNumber="395">Mountain</collectingMunicipality>
</location>
Formation
</location>
at
<collectingCounty id="5B364E64FFA2FF9BA9BDEDE6822DF95F" box="[1179,1283,1735,1761]" pageId="3" pageNumber="395">Republic</collectingCounty>
,
<collectingRegion id="702CF80AFFA2FF9BA836EDE682B6F95F" box="[1296,1432,1735,1761]" country="United States of America" name="Washington" pageId="3" pageNumber="395">Washington</collectingRegion>
,
<collectingCountry id="CAFF7678FFA2FF9BADB1EDCA87C8F8BB" box="[151,230,1771,1797]" name="United States of America" pageId="3" pageNumber="395">U.S.A.</collectingCountry>
, exposure
<specimenCode id="E24E9E93FFA2FF9BAC78EDCA8683F8BB" box="[350,429,1771,1797]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="395" type="Herbarium">B4131</specimenCode>
; latest Ypresian
</materialsCitation>
.
</paragraph>
<paragraph id="B25736E8FFA2FF9BADE1EC2E86DEF863" blockId="3.[151,1437,1591,2014]" pageId="3" pageNumber="395">
<emphasis id="809CEAFAFFA2FF9BADE1EC2E86FEF897" bold="true" box="[199,464,1807,1834]" pageId="3" pageNumber="395">
Position of
<taxonomicName id="75E84D6BFFA2FF9BAC72EC2E86E7F894" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[340,457,1807,1834]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA2FF9BAC72EC2E86E7F894" bold="true" box="[340,457,1807,1834]" italics="true" pageId="3" pageNumber="395">Republica</emphasis>
</taxonomicName>
.
</emphasis>
<taxonomicName id="75E84D6BFFA2FF9BACFCEC2E8562F897" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[474,588,1807,1833]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="395" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA2FF9BACFCEC2E8562F897" box="[474,588,1807,1833]" italics="true" pageId="3" pageNumber="395">Republica</emphasis>
</taxonomicName>
belongs to the Euphaeida and Epallagoidea of
<bibRefCitation id="D6794B19FFA2FF9BA95EEC2E8232F897" author="Bechly, G." box="[1144,1308,1807,1833]" pageId="3" pageNumber="395" pagination="33 - 63" refId="ref4220" refString="Bechly, G. (1998) New fossil damsel flies from Baltic amber, with description of a new species, a redescription of Litheuphaea carpenteri Fraser, and a discussion on the phylogeny of Epallagidae (Zygoptera: Caloptera). International Journal of Odonatology, 1, 33 - 63. https: // doi. org / 10.1080 / 13887890.1998.9748092" type="journal article" year="1998">Bechly (1998)</bibRefCitation>
and to the
<taxonomicName id="75E84D6BFFA2FF9BADB1EC128634F8F3" box="[151,282,1843,1869]" class="Insecta" family="Euphaeidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="family">Euphaeidae</taxonomicName>
by the combination of (as
<taxonomicName id="75E84D6BFFA2FF9BAF74EC1285F5F8F3" authorityName="Needham" authorityYear="1903" box="[594,731,1843,1869]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="family">Epallagidae</taxonomicName>
:
<bibRefCitation id="D6794B19FFA2FF9BAFC0EC128454F8F3" author="Bechly, G." box="[742,890,1843,1869]" pageId="3" pageNumber="395" pagination="1 - 402" refId="ref4172" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Beruchsichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Boblingen, Special Volume 2, 1 - 402." type="journal article" year="1996">Bechly 1996</bibRefCitation>
,
<bibRefCitation id="D6794B19FFA2FF9BAEA1EC1284ECF8F3" author="Bechly, G." box="[903,962,1843,1869]" pageId="3" pageNumber="395" pagination="33 - 63" refId="ref4220" refString="Bechly, G. (1998) New fossil damsel flies from Baltic amber, with description of a new species, a redescription of Litheuphaea carpenteri Fraser, and a discussion on the phylogeny of Epallagidae (Zygoptera: Caloptera). International Journal of Odonatology, 1, 33 - 63. https: // doi. org / 10.1080 / 13887890.1998.9748092" type="journal article" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="D6794B19FFA2FF9BAEE8EC1283C7F8F3" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[974,1257,1843,1870]" pageId="3" pageNumber="395" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA2FF9BA94CEC1583B3F8F3" box="[1130,1181,1843,1869]" italics="true" pageId="3" pageNumber="395">et al</emphasis>
. 2007
</bibRefCitation>
): the distal side of the quadrangle and sub-quadrangle (discoidal bracket: “db” of
<figureCitation id="2AD32A6DFFA2FF9BAE5BEC768493F8CF" box="[893,957,1879,1905]" captionStart="FIGURE 1" captionStartId="5.[151,250,1306,1330]" captionTargetBox="[151,1428,498,1270]" captionTargetId="figure-168@5.[151,1436,489,1282]" captionTargetPageId="5" captionText="FIGURE 1. Republica weatbrooki Holotype SR 06-59-08. A, photograph; B, drawing. Both to scale, 5 mm; db, discoidal bracket, see text." figureDoi="http://doi.org/10.5281/zenodo.4736622" httpUri="https://zenodo.org/record/4736622/files/figure.png" pageId="3" pageNumber="395">Fig. 1</figureCitation>
) is thick, with reversed obliquity; densely reticulate venation; CuA sigmoidally and smoothly curved; curved intercalary veins between CuA and the hind margin; antenodal crossveins numerous [
<taxonomicName id="75E84D6BFFA2FF9BAF55ECBE85D1F807" authorityName="Cockerell" authorityYear="1928" box="[627,767,1951,1977]" class="Insecta" family="Zacallitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="family">Zacallitidae</taxonomicName>
: less numerous]; and the quadrangle is longer than wide [
<taxonomicName id="75E84D6BFFA2FF9BADB8ECE28604F863" authorityName="Cockerell" authorityYear="1928" box="[158,298,1987,2013]" class="Insecta" family="Zacallitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="family">Zacallitidae</taxonomicName>
: as long as wide].
</paragraph>
<paragraph id="B25736E8FFA5FF9CADE1EBB686EEFE13" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="396">
It is assigned to the subfamily
<taxonomicName id="75E84D6BFFA5FF9CAF0AEBB685C5FF0F" authorityName="Cockerell" authorityYear="1923" box="[556,747,151,177]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Eodichromatinae">Eodichromatinae</taxonomicName>
by Ax1 and Ax2 being distinctly bolder than the secondary antenodal crossveins (
<bibRefCitation id="D6794B19FFA5FF9CACA9EB9A8589FF6B" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[399,679,187,213]" pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CAF0CEB9A8572FF6B" box="[554,604,187,213]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
); other character states listed by
<bibRefCitation id="D6794B19FFA5FF9CA930EB9A8399FF6B" author="Bechly, G." box="[1046,1207,187,213]" pageId="4" pageNumber="396" pagination="33 - 63" refId="ref4220" refString="Bechly, G. (1998) New fossil damsel flies from Baltic amber, with description of a new species, a redescription of Litheuphaea carpenteri Fraser, and a discussion on the phylogeny of Epallagidae (Zygoptera: Caloptera). International Journal of Odonatology, 1, 33 - 63. https: // doi. org / 10.1080 / 13887890.1998.9748092" type="journal article" year="1998">Bechly (1998)</bibRefCitation>
and
<bibRefCitation id="D6794B19FFA5FF9CA9C8EB9A862BFF47" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CA8AEEB9A8783FF47" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. (2007)
</bibRefCitation>
that separate the subfamilies cannot be evaluated by preservation. We assign it to the tribe
<taxonomicName id="75E84D6BFFA5FF9CA9C8EBFE82B2FF47" authorityName="Cockerell" authorityYear="1923" box="[1262,1436,223,249]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="tribe" tribe="Eodichromatini">Eodichromatini</taxonomicName>
(
<taxonomicName id="75E84D6BFFA5FF9CAD87EA228604FEA3" authorityName="Cockerell" authorityYear="1923" box="[161,298,259,285]" class="Insecta" family="Epallagidae" genus="Eodichroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAD87EA228604FEA3" box="[161,298,259,285]" italics="true" pageId="4" pageNumber="396">Eodichroma</emphasis>
</taxonomicName>
,
<taxonomicName id="75E84D6BFFA5FF9CAC64EA2286CAFEA3" authorityName="Nel" authorityYear="1988" box="[322,484,259,285]" class="Insecta" family="Epallagidae" genus="Parazacallites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAC64EA2286CAFEA3" box="[322,484,259,285]" italics="true" pageId="4" pageNumber="396">Parazacallites</emphasis>
</taxonomicName>
,
<taxonomicName id="75E84D6BFFA5FF9CACDAEA2285BAFEA3" box="[508,660,259,285]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CACDAEA2285BAFEA3" box="[508,660,259,285]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
</taxonomicName>
.,
<taxonomicName id="75E84D6BFFA5FF9CAF95EA228435FEA3" authorityName="Zessin" authorityYear="2011" box="[691,795,259,285]" class="Insecta" family="Epallagidae" genus="Solveigia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAF95EA228435FEA3" box="[691,795,259,285]" italics="true" pageId="4" pageNumber="396">Solveigia</emphasis>
</taxonomicName>
,
<taxonomicName id="75E84D6BFFA5FF9CAE15EA22848BFEA3" authorityName="Zessin" authorityYear="2011" box="[819,933,259,285]" class="Insecta" family="Epallagidae" genus="Ejerslevia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAE15EA22848BFEA3" box="[819,933,259,285]" italics="true" pageId="4" pageNumber="396">Ejerslevia</emphasis>
</taxonomicName>
, and
<emphasis id="809CEAFAFFA5FF9CAED0EA228391FEA3" box="[1014,1215,259,285]" italics="true" pageId="4" pageNumber="396">Wolfgangeuphaea</emphasis>
) by the presence of secondary antenodals between ScP and RA, absent in the other tribe of
<taxonomicName id="75E84D6BFFA5FF9CA925EA0683EBFEFF" authorityName="Cockerell" authorityYear="1923" box="[1027,1221,295,321]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Eodichromatinae">Eodichromatinae</taxonomicName>
, the
<taxonomicName id="75E84D6BFFA5FF9CA827EA0682B3FEFF" authorityName="Bechly" authorityYear="1996" box="[1281,1437,295,321]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="tribe" tribe="Litheuphaeini">Litheuphaeini</taxonomicName>
(
<taxonomicName id="75E84D6BFFA5FF9CAD87EA6A8650FEDB" authority="Fraser" authorityName="Fraser" authorityYear="1955" box="[161,382,331,357]" class="Insecta" family="Epallagidae" genus="Litheuphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAD87EA6A8603FEDB" box="[161,301,331,357]" italics="true" pageId="4" pageNumber="396">Litheuphaea</emphasis>
Fraser
</taxonomicName>
).
<taxonomicName id="75E84D6BFFA5FF9CACB6EA6A8537FEDB" authorityName="Cockerell" authorityYear="1923" box="[400,537,331,357]" class="Insecta" family="Epallagidae" genus="Eodichroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CACB6EA6A8537FEDB" box="[400,537,331,357]" italics="true" pageId="4" pageNumber="396">Eodichroma</emphasis>
</taxonomicName>
is poorly known; a drawing of its single fossil wing was never published, only a poor-quality photo (
<bibRefCitation id="D6794B19FFA5FF9CAC52EA4E850AFE37" author="Cockerell, T. D. A." box="[372,548,367,393]" pageId="4" pageNumber="396" pagination="397 - 400" refId="ref4686" refString="Cockerell, T. D. A. (1923) Fossil insects from the Eocene of Texas. American Journal of Science, Series 5, 5, 397 - 400. https: // doi. org / 10.2475 / ajs. s 5 - 5.29.397" type="journal article" year="1923">Cockerell 1923</bibRefCitation>
, fig. 2; reproduced by
<bibRefCitation id="D6794B19FFA5FF9CAE3AEA4E84B3FE37" author="Berry, E. W." box="[796,925,367,393]" pageId="4" pageNumber="396" refId="ref4437" refString="Berry, E. W. (1924) The middle and upper Eocene floras of southwestern North America. Professional Paper 92, United States Geological Survey, Department of the Interior, Washington, 206 pp. https: // doi. org / 10.3133 / pp 92" type="book" year="1924">Berry 1924</bibRefCitation>
, figure 1 of his plate LXIII), and the specimen is now lost (
<bibRefCitation id="D6794B19FFA5FF9CAC39EAB2869DFE13" author="Bechly, G." box="[287,435,403,429]" pageId="4" pageNumber="396" pagination="33 - 63" refId="ref4220" refString="Bechly, G. (1998) New fossil damsel flies from Baltic amber, with description of a new species, a redescription of Litheuphaea carpenteri Fraser, and a discussion on the phylogeny of Epallagidae (Zygoptera: Caloptera). International Journal of Odonatology, 1, 33 - 63. https: // doi. org / 10.1080 / 13887890.1998.9748092" type="journal article" year="1998">Bechly 1998</bibRefCitation>
).
</paragraph>
<paragraph id="B25736E8FFA5FF9CADE1EA9684EDFD17" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="396">
The wing of
<taxonomicName id="75E84D6BFFA5FF9CAC71EA9686E7FE6F" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[343,457,439,465]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAC71EA9686E7FE6F" box="[343,457,439,465]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
is morphologically closest to those of
<taxonomicName id="75E84D6BFFA5FF9CAE5EEA96833EFE6F" box="[888,1040,439,465]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAE5EEA96833EFE6F" box="[888,1040,439,465]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
</taxonomicName>
as in the diagnosis of
<bibRefCitation id="D6794B19FFA5FF9CA82CEA96860CFE4B" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CADB1EAFA87E9FE4B" box="[151,199,475,501]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. (2007)
</bibRefCitation>
: it has more than 20 antenodal crossveins; the arculus very close to Ax2; an elongate, narrow quadrangle (inferred in
<taxonomicName id="75E84D6BFFA5FF9CAC04EADE86E0FDA7" authority=": MP" authorityName="MP" box="[290,462,511,537]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAC04EADE86BAFDA7" box="[290,404,511,537]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
: MP
</taxonomicName>
+CuA, between the quadrangle and sub-quadrangle is not preserved); most likely six long secondary longitudinal veins in the CuA–A space (this region not well preserved in part in
<taxonomicName id="75E84D6BFFA5FF9CA9F6E902826CFD83" box="[1232,1346,547,573]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CA9F6E902826CFD83" box="[1232,1346,547,573]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
, but six appears reasonable); more than two rows of cells between C and RA distal to the pterostigma and between RA and RP1 below it. The absence of the pterostigmal brace vein cannot be established as region poorly preserved. Further to the diagnosis, as in
<taxonomicName id="75E84D6BFFA5FF9CACB6E9AE854DFD17" authority=", MA" authorityName="MA" box="[400,611,655,681]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CACB6E9AE8506FD17" box="[400,552,655,681]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
, MA
</taxonomicName>
smoothly curves several times.
</paragraph>
<paragraph id="B25736E8FFA5FF9CADE1E99285A3FC77" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="396">
<taxonomicName id="75E84D6BFFA5FF9CADE1E9928617FD73" box="[199,313,691,717]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CADE1E9928617FD73" box="[199,313,691,717]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
differs from the diagnosis of
<taxonomicName id="75E84D6BFFA5FF9CAFBDE9928342FD73" authority="(Petrulevicius et al. 2007)" baseAuthorityName="Petrulevicius" baseAuthorityYear="2007" box="[667,1132,691,717]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAFBDE992841DFD73" box="[667,819,691,717]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
(
<bibRefCitation id="D6794B19FFA5FF9CAE63E992834AFD73" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[837,1124,691,717]" pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CAEC2E9928337FD73" box="[996,1049,691,717]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
)
</taxonomicName>
as the anal area is not so broadened; however, this broadening is based on the wider hind wings. The distinctive transverse cells of the anal region appear present in the forewing of
<taxonomicName id="75E84D6BFFA5FF9CAF45E9DA8591FCAA" box="[611,703,763,788]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="riveri">
<emphasis id="809CEAFAFFA5FF9CAF45E9DA8591FCAA" box="[611,703,763,788]" italics="true" pageId="4" pageNumber="396">L. riveri</emphasis>
</taxonomicName>
in figure 5 of
<bibRefCitation id="D6794B19FFA5FF9CAE44E9DA836DFCAB" author="Bechly, G. &amp; Garrouste, R. &amp; Aase, A. &amp; Karr, J. A. &amp; Grande, L. &amp; Nel, A." box="[866,1091,763,789]" pageId="4" pageNumber="396" pagination="1 - 30" refId="ref4362" refString="Bechly, G., Garrouste, R., Aase, A., Karr, J. A., Grande, L. &amp; Nel, A. (2020) The damselfly palaeofauna from the Eocene of Wyoming and Colorado, USA (Insecta, Odonata, Zygoptera). Papers in Palaeontology, 2020, 1 - 30. https: // doi. org / 10.1002 / spp 2.1346" type="journal article" year="2020">
Bechly
<emphasis id="809CEAFAFFA5FF9CAE9FE9DA84C4FCAB" box="[953,1002,763,789]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. (2020)
</bibRefCitation>
confirming their presence in a narrower forewing of that genus.
<taxonomicName id="75E84D6BFFA5FF9CAF37E83E85ADFC87" box="[529,643,799,825]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAF37E83E85ADFC87" box="[529,643,799,825]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
lacks these elongate cells.
<taxonomicName id="75E84D6BFFA5FF9CAE92E83E8308FC87" box="[948,1062,799,825]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAE92E83E8308FC87" box="[948,1062,799,825]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
also differs by shorter secondary longitudinal veins between main veins. In
<taxonomicName id="75E84D6BFFA5FF9CAF5DE8628466FCE3" authority="MA" authorityName="MA" box="[635,840,835,861]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAF5DE862843DFCE3" box="[635,787,835,861]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
MA
</taxonomicName>
and CuA diverge toward the wing margin with long secondary longitudinal veins between them, while they are sub-parallel, remaining close in
<taxonomicName id="75E84D6BFFA5FF9CA9ABE84683D1FC3F" box="[1165,1279,871,897]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CA9ABE84683D1FC3F" box="[1165,1279,871,897]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
. Although the space between MA and CuA is poorly preserved in the sole specimen of
<taxonomicName id="75E84D6BFFA5FF9CAEE6E8AA831CFC1B" box="[960,1074,907,933]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAEE6E8AA831CFC1B" box="[960,1074,907,933]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
, there appears to be no room for a secondary longitudinal vein between them.
</paragraph>
<paragraph id="B25736E8FFA5FF9CADE1E8F2862AFBC3" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="396">
<taxonomicName id="75E84D6BFFA5FF9CADE1E8F28617FC53" box="[199,313,979,1005]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CADE1E8F28617FC53" box="[199,313,979,1005]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
is further separated from
<taxonomicName id="75E84D6BFFA5FF9CAF78E8F285D8FC53" box="[606,758,979,1005]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAF78E8F285D8FC53" box="[606,758,979,1005]" italics="true" pageId="4" pageNumber="396">Labandeiraia</emphasis>
</taxonomicName>
by both rows of antenodal, postnodal crossveins with many branching, some connected by horizontal crossveins (diagnostic character state 5). One is, however, branched in
<taxonomicName id="75E84D6BFFA5FF9CADB1EF3A8640FB8B" box="[151,366,1051,1077]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="americaborealis">
<emphasis id="809CEAFAFFA5FF9CADB1EF3A8789FB8A" box="[151,167,1051,1076]" italics="true" pageId="4" pageNumber="396">L</emphasis>
.
<emphasis id="809CEAFAFFA5FF9CAD91EF3A8640FB8B" box="[183,366,1051,1077]" italics="true" pageId="4" pageNumber="396">americaborealis</emphasis>
</taxonomicName>
specimen 31.665A (
<bibRefCitation id="D6794B19FFA5FF9CAF73EF3A845EFB8B" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[597,880,1051,1077]" pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CAFD4EF3A840BFB8B" box="[754,805,1051,1077]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
, fig. 1C), and three in
<taxonomicName id="75E84D6BFFA5FF9CA951EF3A83DFFB8A" box="[1143,1265,1051,1076]" class="Insecta" family="Epallagidae" genus="Labandeiraia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="species" species="europae">
<emphasis id="809CEAFAFFA5FF9CA951EF3A83DFFB8A" box="[1143,1265,1051,1076]" italics="true" pageId="4" pageNumber="396">L. europae</emphasis>
</taxonomicName>
,
<typeStatus id="6D53884AFFA5FF9CA827EF3A824FFB8B" box="[1281,1377,1051,1077]" pageId="4" pageNumber="396" type="paratype">paratype</typeStatus>
MM 1255 (
<bibRefCitation id="D6794B19FFA5FF9CADF8EF1E86DBFBE7" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[222,501,1087,1113]" pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevičius
<emphasis id="809CEAFAFFA5FF9CAC5CEF1E8682FBE7" box="[378,428,1087,1113]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
, fig. 2A)], but we consider these few cases minor, not as in
<taxonomicName id="75E84D6BFFA5FF9CA9BEEF1E8224FBE7" box="[1176,1290,1087,1113]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CA9BEEF1E8224FBE7" box="[1176,1290,1087,1113]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
, where there are many.
</paragraph>
<paragraph id="B25736E8FFA5FF9CADE1EFA68441FAEB" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="396">
We make the assumption that the
<taxonomicName id="75E84D6BFFA5FF9CAF63EFA68599FB1F" box="[581,695,1159,1185]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CAF63EFA68599FB1F" box="[581,695,1159,1185]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
wing is a forewing, as wherever the wings of
<taxonomicName id="75E84D6BFFA5FF9CA9E2EFA6825CFB1F" authorityName="Cockerell" authorityYear="1923" box="[1220,1394,1159,1185]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="tribe" tribe="Eodichromatini">Eodichromatini</taxonomicName>
are known to be forewings or hind wings, the hind wing is broader to much broader and/or more triangular, deeper in the middle than the forewing. Even when the hind wing is not as extremely broad as it is in
<emphasis id="809CEAFAFFA5FF9CA9A9EFEE823CFB57" box="[1167,1298,1231,1257]" italics="true" pageId="4" pageNumber="396">Labandeira</emphasis>
species (see
<bibRefCitation id="D6794B19FFA5FF9CADB1EFD28685FAB3" author="Petrulevicius, J. F. &amp; Nel, A. &amp; Rust, J. &amp; Bechly, G. &amp; Kohls, D." box="[151,427,1267,1293]" pageId="4" pageNumber="396" pagination="15 - 25" refId="ref5718" refString="Petrulevicius, J. F., Nel, A., Rust, J., Bechly, G. &amp; Kohls, D. (2007) New Paleogene Epallagidae (Insecta: Odonata) recorded in North America and Europe. Biogeographic implications. Alavesia, 1, 15 - 25." type="journal article" year="2007">
Petrulevicius
<emphasis id="809CEAFAFFA5FF9CAC16EFD2864FFAB3" box="[304,353,1267,1293]" italics="true" pageId="4" pageNumber="396">et al</emphasis>
. 2007
</bibRefCitation>
), it is triangular, not smoothly curved along the posterior margin as in the
<taxonomicName id="75E84D6BFFA5FF9CA9C3EFD28279FAB3" box="[1253,1367,1267,1293]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CA9C3EFD28279FAB3" box="[1253,1367,1267,1293]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
wing. If the
<taxonomicName id="75E84D6BFFA5FF9CADFEEE368664FA8F" box="[216,330,1303,1329]" class="Magnoliopsida" genus="Republica" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="396" phylum="Tracheophyta" rank="genus">
<emphasis id="809CEAFAFFA5FF9CADFEEE368664FA8F" box="[216,330,1303,1329]" italics="true" pageId="4" pageNumber="396">Republica</emphasis>
</taxonomicName>
wing were a hind wing, it would differ much more distinctly from other
<taxonomicName id="75E84D6BFFA5FF9CA94CEE368236FA8F" authorityName="Cockerell" authorityYear="1923" box="[1130,1304,1303,1329]" class="Insecta" family="Epallagidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="396" phylum="Arthropoda" rank="tribe" tribe="Eodichromatini">Eodichromatini</taxonomicName>
in this way. To be conservative, however, we treat it as a presumed forewing.
</paragraph>
</subSubSection>
</treatment>
</document>
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