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<document id="9731A59222A01B8B60794CC1C1F3D38E" ID-DOI="10.37828/em.2024.75.1" ID-ISSN="2336-9744" ID-Zenodo-Dep="13249519" ID-ZooBank="urn:lsid:zoobank.org:pub:C0DB2C3E-017A-4676-AFF3-0D40121D5CEE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722975030520" checkinUser="felipe" docAuthor="Melnitsky, Stanislav I., Ivanov, Vladimir D., Perkovsky, Evgeny E. &amp; Legalov, Andrei A." docDate="2024" docId="3A0BEA25B052AF03AAE46A8BC93A8A90" docLanguage="en" docName="EcolMontenegrina.75.1-11.pdf" docOrigin="Ecologica Montenegrina 75" docSource="http://dx.doi.org/10.37828/em.2024.75.1" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Holocentropus totuttotam Melnitsky, Ivanov &amp; Perkovsky 2024, sp. n." docType="treatment" docUuid="0DCDCB33-E8FB-4995-B45D-BE9D80F7A65F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="3" masterDocId="C632925DB053AF01AA596F1ECC4D8E74" masterDocTitle="The new species and two new records of Polycentropodidae (Insecta: Trichoptera) from Eocene Rovno amber" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1722984628581" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="56AAEF69C810F559B34E02C547B30289">The new species and two new records of Polycentropodidae (Insecta: Trichoptera) from Eocene Rovno amber</mods:title>
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<mods:namePart id="2EC26E398BC6363C114EA95B00ED934C">Melnitsky, Stanislav I.</mods:namePart>
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<mods:affiliation id="870F5DB2C6FD8B7D77117F9F8402B56F">Department of Entomology, Faculty of Biology, St. Petersburg State University, Universitetskaya nab. 7 / 9, St. Petersburg 199034, Russia; https: // orcid. org / 0000 - 0003 - 4535 - 1886; https: // orcid. org / 0000 - 0003 - 3294 - 4965; e-mail: simelnitsky @ yandex. ru</mods:affiliation>
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<mods:namePart id="1AC3229000B4FBA590A4D49AB62104A7">Ivanov, Vladimir D.</mods:namePart>
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<mods:affiliation id="3FCDCFBB0C94C4DF20662DE5709B5C16">Department of Entomology, Faculty of Biology, St. Petersburg State University, Universitetskaya nab. 7 / 9, St. Petersburg 199034, Russia; https: // orcid. org / 0000 - 0003 - 4535 - 1886; https: // orcid. org / 0000 - 0003 - 3294 - 4965; e-mail: simelnitsky @ yandex. ru</mods:affiliation>
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<mods:namePart id="636032B78F0BC101E10DD76A5C7F358F">Perkovsky, Evgeny E.</mods:namePart>
<mods:nameIdentifier id="D0F52B1049445712F03F206299B50A0F" type="ORCID">0000-0002-7959-4379</mods:nameIdentifier>
<mods:affiliation id="98FA7DF1BA2377CB95FF2A0F8B74371A">Natural History Museum of Denmark, 2100 Copenhagen, Denmark; https: // orcid. org / 0000 - 0002 - 7959 - 4379</mods:affiliation>
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<mods:namePart id="7366F03D71FD05156EC6AA029D3827F0">Legalov, Andrei A.</mods:namePart>
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<mods:affiliation id="F0263E7221CE2994FB4E8E39963D1930">Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, 630091 Novosibirsk, Russia; https: // orcid. org / 0000 - 0001 - 7347 - 8169 &amp; Department of Ecology, Biochemistry and Biotechnology, Altai State University, 656049 Barnaul, Russia; &amp; Department of Forestry and Landscape Construction, Tomsk State University, 634050 Tomsk, Russia. * Corresponding author. E-mail: fossilweevils @ gmail. com</mods:affiliation>
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<mods:date id="8A4E629C3CF42FED78007181C971FE39">2024</mods:date>
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<paragraph id="B21D5B33B052AF00AAE46A8BCF9E8BDB" blockId="1.[189,1199,1429,1519]" box="[189,979,1429,1455]" pageId="1" pageNumber="2">
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<emphasis id="80D68721B052AF00AAE46A8BCF9E8BDB" bold="true" box="[189,979,1429,1455]" pageId="1" pageNumber="2">
<taxonomicName id="75A220B0B052AF00AAE46A8BCFC48BDB" authority="Melnitsky, Ivanov &amp; Perkovsky" authorityName="Melnitsky, Ivanov &amp; Perkovsky" authorityYear="2024" box="[189,905,1429,1455]" class="Insecta" family="Polycentropodidae" genus="Holocentropus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="totuttotam" status="sp. nov.">
<emphasis id="80D68721B052AF00AAE46A8BCDB98BDB" bold="true" box="[189,500,1429,1455]" italics="true" pageId="1" pageNumber="2">Holocentropus totuttotam</emphasis>
Melnitsky, Ivanov &amp; Perkovsky
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<taxonomicNameLabel id="9BE53A5AB052AF00A9C86A8BCF9E8BDB" box="[913,979,1429,1455]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="FAB808B8B052AF00AAE46ACACD0B8B9B" box="[189,326,1492,1519]" pageId="1" pageNumber="2" type="description">
<paragraph id="B21D5B33B052AF00AAE46ACACD0B8B9B" blockId="1.[189,1199,1429,1519]" box="[189,326,1492,1519]" pageId="1" pageNumber="2">
(
<figureCitation id="2A9947B6B052AF00AA9C6ACACD718B9B" box="[197,316,1492,1519]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[189,264,1025,1049]" captionStartId-1="2.[189,264,1999,2023]" captionTargetBox-0="[224,1363,189,1007]" captionTargetBox-1="[420,1168,1276,1972]" captionTargetId-0="figure-118@2.[224,1363,189,1007]" captionTargetId-1="figure-120@2.[410,1178,1264,1979]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="Figure 1. Holocentropus totuttotam sp. nov. Male type, general view, ventral. Scale bar = 0.5 mm." captionText-1="Figure 2. Holocentropus totuttotam sp. nov. Male type, genitalia, ventral view. Abbreviations: X 10th segment, aed aedeagus, ia inferior appendages, pa preanal appendages." figureDoi-0="http://doi.org/10.5281/zenodo.13249523" figureDoi-1="http://doi.org/10.5281/zenodo.13249525" httpUri-0="https://zenodo.org/record/13249523/files/figure.png" httpUri-1="https://zenodo.org/record/13249525/files/figure.png" pageId="1" pageNumber="2">Figs 1 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FAB808B8B052AF00AAE46906C8C18847" box="[189,1164,1560,1587]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="B21D5B33B052AF00AAE46906C8C18847" blockId="1.[189,1399,1560,1992]" box="[189,1164,1560,1587]" pageId="1" pageNumber="2">
<emphasis id="80D68721B052AF00AAE46906CD358846" bold="true" box="[189,376,1560,1586]" pageId="1" pageNumber="2">Type material:</emphasis>
<materialsCitation id="02CA516EB052AF00AB266906C8C18847" box="[383,1164,1560,1587]" collectionCode="SIZK" location="Klesov" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
<typeStatus id="6D19E591B052AF00AB266906CDA18847" box="[383,492,1560,1587]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
. Male.
<collectionCode id="D4B3C3F6B052AF00A81F6906CECA8847" box="[582,647,1560,1587]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="2">SIZK</collectionCode>
K325,
<location id="B77D0DE8B052AF00A8B46906CF0E8847" LSID="urn:lsid:plazi:treatment:3A0BEA25B052AF03AAE46A8BC93A8A90:B77D0DE8B052AF00A8B46906CF0E8847" box="[749,835,1560,1587]" name="Klesov" pageId="1" pageNumber="2" stateProvince="Rivne">Klesov</location>
,
<collectingRegion id="706695D1B052AF00A9166907CFD08847" box="[847,925,1561,1587]" country="Ukraine" name="Rivne" pageId="1" pageNumber="2">Rovno</collectingRegion>
amber, late Eocene.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FAB808B8B052AF00AB456924CD6589BC" pageId="1" pageNumber="2" type="description">
<paragraph id="B21D5B33B052AF00AB456924C8EA8803" blockId="1.[189,1399,1560,1992]" pageId="1" pageNumber="2">
<emphasis id="80D68721B052AF00AB456924CDF98820" bold="true" box="[284,436,1594,1620]" pageId="1" pageNumber="2">Description.</emphasis>
Body length
<quantity id="755AF6D6B052AF00A80E6924CEF88821" box="[599,693,1594,1621]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="2" unit="mm" value="3.6">3.6 mm</quantity>
; forewing length
<quantity id="755AF6D6B052AF00A9DE6924CFAB8821" box="[903,998,1594,1621]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="2" unit="mm" value="4.6">4.6 mm</quantity>
. Head, antennae, thorax, legs and abdomen light brown. Wings brown with numerous light hairs. Head with light hairs.
</paragraph>
<paragraph id="B21D5B33B052AF00AB456963CDD58935" blockId="1.[189,1399,1560,1992]" pageId="1" pageNumber="2">Male genitalia. Inferior appendages of unusual structure: in ventrocaudal view, rectangular bases have two curved teeth: sharp lateral on a small lobe, and robust medial with wider basal part, these tips directed towards each other. Ventrolateral well-sclerotized finger-like projection with setose apical rounded tip is seen in between them. Posterior margin of segment X with two membranous lobes. Preanal appendages (cerci) with extended concave apex covered long hairs. Aedeagus very thin, slightly forked at the apex.</paragraph>
<paragraph id="B21D5B33B052AF00AB456856CD6589BC" blockId="1.[189,1399,1560,1992]" pageId="1" pageNumber="2">
<emphasis id="80D68721B052AF00AB456856CDF38916" bold="true" box="[284,446,1864,1890]" pageId="1" pageNumber="2">Comparison.</emphasis>
The new species differs from
<taxonomicName id="75A220B0B052AF00A9786856C88E8917" authority="Ulmer, 1912" authorityName="Ulmer" authorityYear="1912" box="[801,1219,1864,1891]" class="Insecta" family="Polycentropodidae" genus="Holocentropus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="omega">
<emphasis id="80D68721B052AF00A9786856C8688916" box="[801,1061,1864,1891]" italics="true" pageId="1" pageNumber="2">Holocentropus omega</emphasis>
<bibRefCitation id="D63326C2B052AF00AE756856C88E8917" author="Ulmer, G." box="[1068,1219,1864,1891]" pageId="1" pageNumber="2" pagination="1 - 380" refId="ref5109" refString="Ulmer, G. (1912) Die Trichopteren des Baltischen Bernsteins. Beitrage zur Naturkunde Preussens, 10, 1 - 380." type="journal article" year="1912">Ulmer, 1912</bibRefCitation>
</taxonomicName>
, which appears to be the closest to it, by the shape of the inferior appendages: in
<taxonomicName id="75A220B0B052AF00A99D6877C88489F0" authorityName="Ulmer" authorityYear="1912" box="[964,1225,1897,1924]" class="Insecta" family="Polycentropodidae" genus="Holocentropus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="omega">
<emphasis id="80D68721B052AF00A99D6877C88489F0" box="[964,1225,1897,1924]" italics="true" pageId="1" pageNumber="2">Holocentropus omega</emphasis>
</taxonomicName>
, the basal part longer and have a lateral incision, ventromedial tooth shorter than basal part, ventrolateral part is more elongate.
</paragraph>
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<caption id="E6DD0BBBB051AF03AAE46B1FC8918A6D" ID-DOI="http://doi.org/10.5281/zenodo.13249523" ID-Zenodo-Dep="13249523" box="[189,1244,1025,1050]" httpUri="https://zenodo.org/record/13249523/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,264,1025,1049]" targetBox="[224,1363,189,1007]" targetPageId="2" targetType="figure">
<paragraph id="B21D5B33B051AF03AAE46B1FC8918A6D" blockId="2.[189,1244,1025,1050]" box="[189,1244,1025,1050]" pageId="2" pageNumber="3">
<emphasis id="80D68721B051AF03AAE46B1FCD518A6D" bold="true" box="[189,284,1025,1049]" pageId="2" pageNumber="3">Figure 1</emphasis>
.
<taxonomicName id="75A220B0B051AF03AB736B1FCE718A6D" authorityName="Melnitsky &amp; Ivanov &amp; Perkovsky &amp; Legalov" authorityYear="2024" box="[298,572,1025,1049]" class="Insecta" family="Polycentropodidae" genus="Holocentropus" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="totuttotam" status="sp. nov.">
<emphasis id="80D68721B051AF03AB736B1FCE718A6D" box="[298,572,1025,1049]" italics="true" pageId="2" pageNumber="3">Holocentropus totuttotam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9BE53A5AB051AF03A81D6B1CCED58A6D" box="[580,664,1026,1049]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
Male type, general view, ventral. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="FAB808B8B051AF03AB456B7ACF838AD4" pageId="2" pageNumber="3" type="etymology">
<paragraph id="B21D5B33B051AF03AB456B7ACF838AD4" blockId="2.[189,1399,1123,1252]" pageId="2" pageNumber="3">
<emphasis id="80D68721B051AF03AB456B7ACDE78A0A" bold="true" box="[284,426,1124,1150]" pageId="2" pageNumber="3">Etymology.</emphasis>
The specific name
<taxonomicName id="75A220B0B051AF03A8C86B7BCF478A0A" authorityName="Melnitsky &amp; Ivanov &amp; Perkovsky &amp; Legalov" authorityYear="2024" box="[657,778,1125,1150]" class="Insecta" family="Polycentropodidae" genus="Holocentropus" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="totuttotam">
<emphasis id="80D68721B051AF03A8C86B7BCF478A0A" box="[657,778,1125,1150]" italics="true" pageId="2" pageNumber="3">totuttotam</emphasis>
</taxonomicName>
is indeclinable and must not agree in gender with its generic name; it derives from Russian and means “here and there”.
</paragraph>
</subSubSection>
<subSubSection id="FAB808B8B051AF03AB456BB9CEBC8AB6" box="[284,753,1191,1218]" pageId="2" pageNumber="3" type="distribution">
<paragraph id="B21D5B33B051AF03AB456BB9CEBC8AB6" blockId="2.[189,1399,1123,1252]" box="[284,753,1191,1218]" pageId="2" pageNumber="3">
<emphasis id="80D68721B051AF03AB456BB9CDF18AB5" bold="true" box="[284,444,1191,1217]" pageId="2" pageNumber="3">Distribution.</emphasis>
Priabonian
<collectingRegion id="706695D1B051AF03A8136BB6CED58AB6" box="[586,664,1192,1218]" country="Ukraine" name="Rivne" pageId="2" pageNumber="3">Rovno</collectingRegion>
amber.
</paragraph>
</subSubSection>
<subSubSection id="FAB808B8B051AF03AB456BD7C93A8A90" box="[284,1399,1225,1252]" pageId="2" pageNumber="3" type="discussion">
<paragraph id="B21D5B33B051AF03AB456BD7C93A8A90" blockId="2.[189,1399,1123,1252]" box="[284,1399,1225,1252]" pageId="2" pageNumber="3">
<emphasis id="80D68721B051AF03AB456BD7CDC38A97" bold="true" box="[284,398,1225,1251]" pageId="2" pageNumber="3">Remarks</emphasis>
. The specimen was found in the small piece of amber (
<quantity id="755AF6D6B051AF03AE4D6BD7C8038A90" box="[1044,1102,1225,1252]" metricMagnitude="-3" metricUnit="kg" metricValue="1.1" pageId="2" pageNumber="3" unit="g" value="1.1">1.1 g</quantity>
after primary treatment).
</paragraph>
</subSubSection>
</treatment>
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