treatments-xml/data/3A/01/87/3A0187C5FFFF627FCAE254F6FD74FAAF.xml

<document id="49CBBF4ECDA51510371BB86405762153" ID-CLB-Dataset="189442" ID-DOI="10.11646/zootaxa.5209.4.2" ID-GBIF-Dataset="27ec26b5-9527-49ac-9fa5-1f441c2e0f7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7333828" ID-ZooBank="96EA6905-9FC6-4D27-80EE-372725585540" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668767704168" checkinUser="plazi" docAuthor="Almón, Bruno, Cuesta, Jose A. &amp; García-Raso, J. Enrique" docDate="2022" docId="3A0187C5FFFF627FCAE254F6FD74FAAF" docLanguage="en" docName="zootaxa.5209.4.2.pdf" docOrigin="Zootaxa 5209 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pagurus pectinidactylus Almón &amp; Cuesta &amp; García-Raso 2022, n. sp." docType="treatment" docUuid="E87A7D6A-9982-43A6-B2D3-AE1D821153FF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="436" masterDocId="C638FFBDFFFB6275CA75572EFFD6FFFA" masterDocTitle="A new species of hermit crab of the genus Pagurus Fabricius, 1775 (Crustacea: Anomura: Paguridae) from north-western Africa" masterLastPageNumber="440" masterPageNumber="426" pageNumber="430" updateTime="1702000383839" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="00B4907871F2894123F41B5ACAC9E935">A new species of hermit crab of the genus Pagurus Fabricius, 1775 (Crustacea: Anomura: Paguridae) from north-western Africa</mods:title>
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<mods:affiliation id="447C70274E118E887F7EBC8435E9CED5">Instituto de Ciencias Marinas de Andalucía, ICMAN-CSIC, Avenida República Saharaui, 2, 11519 Puerto Real, Cádiz, Spain. jose. cuesta @ icman. csic. es; https: // orcid. org / 0000 - 0001 - 9482 - 2336</mods:affiliation>
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<mods:affiliation id="866625DF88BA79D277A7864679561BDE">Universidad de Málaga, Fac. de Ciencias, Dep. Biología Animal, Campus Teatinos, s / n, 29071, Málaga, Spain. garciaraso @ uma. es; https: // orcid. org / 0000 - 0003 - 3092 - 9518</mods:affiliation>
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(
<figureCitation id="2A932A56FFFF6271CAEA54D2FEDCFBEC" box="[159,266,1020,1046]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="7.[151,250,1874,1898]" captionStartId-1="8.[151,250,1900,1924]" captionStartId-2="9.[151,250,1866,1890]" captionTargetBox-0="[193,1395,193,1836]" captionTargetBox-1="[205,1391,191,1870]" captionTargetBox-2="[222,1380,187,1836]" captionTargetId-0="figure-24@7.[181,1407,181,1850]" captionTargetId-1="figure-18@8.[195,1393,181,1876]" captionTargetId-2="figure-24@9.[180,1407,181,1842]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." captionText-1="FIGURE 3. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm)." captionText-2="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.7333832" figureDoi-1="http://doi.org/10.5281/zenodo.7333834" figureDoi-2="http://doi.org/10.5281/zenodo.7333836" httpUri-0="https://zenodo.org/record/7333832/files/figure.png" httpUri-1="https://zenodo.org/record/7333834/files/figure.png" httpUri-2="https://zenodo.org/record/7333836/files/figure.png" pageId="4" pageNumber="430">Figs. 2–4</figureCitation>
)
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lsid:zoobank.org:act: 
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<emphasis id="80DCEAC1FFFF6271CAE25346FE94FB78" bold="true" box="[151,322,1128,1154]" pageId="4" pageNumber="430">Type material.</emphasis>
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: 
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sl. 
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(
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), 
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, off 
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, 
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, sand, 
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depth
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, 
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. 
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: 2 ovigerous 
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, sl. 3.9 and 
<quantity id="75509B36FFFF6271C9EE53A2FC26FB5D" box="[923,1008,1164,1191]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="430" unit="mm" value="2.9">2.9 mm</quantity>
(
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), 
<collectingCountry id="CABF7643FFFF6271CEAC53A2FA81FB5C" box="[1241,1367,1164,1190]" name="Mauritania" pageId="4" pageNumber="430">Mauritania</collectingCountry>
, north of 
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, 
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<geoCoordinate id="D79C5014FFFF6271CB92539FFD5EFB36" box="[487,648,1201,1228]" degrees="16" direction="west" minutes="18" orientation="longitude" pageId="4" pageNumber="430" precision="1" seconds="32.0" value="-16.308887">16°18′32.0″W</geoCoordinate>
, sand, 
<elevation id="3985D1E0FFFF6271C8A0539EFC83FB30" box="[725,853,1200,1227]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="430" unit="m" value="30.0">
<quantity id="75509B36FFFF6271C8A0539EFCD8FB31" box="[725,782,1200,1227]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="430" unit="m" value="30.0">30 m</quantity>
depth
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, 
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<collectingDate id="D652E9FBFFFF6271C914539EFC2CFB30" box="[865,1018,1200,1227]" pageId="4" pageNumber="430" value="2012-06-03">03 June 2012</collectingDate>
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.
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<paragraph id="B21736D3FFFF6271CAB253FAFD7FFAA0" blockId="4.[151,1437,1128,2019]" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB253FAFE8FFB14" bold="true" box="[199,345,1236,1262]" pageId="4" pageNumber="430">Description.</emphasis>
<emphasis id="80DCEAC1FFFF6271CB1153FAFE7CFB14" box="[356,426,1236,1262]" italics="true" pageId="4" pageNumber="430">Shield</emphasis>
(
<figureCitation id="2A932A56FFFF6271CBCB53FAFDCDFB14" box="[446,539,1236,1262]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="4" pageNumber="430">Fig. 2A</figureCitation>
) flattened dorso-ventrally, broader than long, with tufts of short setae along shallow paragastric grooves and numerous low blister-like tubercles only visible when stained with methylene blue. Eleven pairs of phyllobranchia (
<figureCitation id="2A932A56FFFF6271C8775232FD8FFACC" box="[514,601,1308,1334]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="4" pageNumber="430">Fig. 2B</figureCitation>
) with arthrobranchia on Mxp3, on cheliped and pereiopods 2–4, with one well-developed pleurobranchia on pereiopod 4.
</paragraph>
<paragraph id="B21736D3FFFF6271CAB2524BFD69FA3C" blockId="4.[151,1437,1128,2019]" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB2524BFEF0FA84" box="[199,294,1381,1406]" italics="true" pageId="4" pageNumber="430">Rostrum</emphasis>
broadly triangular, tip slightly sloping downwards, with small tubercle distally and produced to or slightly beyond level of obtusely triangular lateral projections; frontal margin between lateral projections thickened; lateral projections each armed with minute spine.
</paragraph>
<paragraph id="B21736D3FFFF6271CAB252FFFB49F9F5" blockId="4.[151,1437,1128,2019]" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB252FFFE76FA10" box="[199,416,1489,1514]" italics="true" pageId="4" pageNumber="430">Posterior carapace</emphasis>
(
<figureCitation id="2A932A56FFFF6271CBDA52FEFDD1FA10" box="[431,519,1488,1514]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="4" pageNumber="430">Fig. 2D</figureCitation>
) membranous except for weakly calcified median and submedian parts, both with low blister-like tubercles and tufts of short setae; branchial region with tufts of simple setae.
</paragraph>
<paragraph id="B21736D3FFFF6271CAB25136FAE1F964" blockId="4.[151,1437,1128,2019]" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB25136FE42F9C8" box="[199,404,1560,1586]" italics="true" pageId="4" pageNumber="430">Ocular peduncles</emphasis>
0.7–0.8 length of shield, slightly swollen medially, with row of short setae medially; conspicuous oblique calcified scale covering the latero-proximal part; corneas slightly dilated; ocular acicles ovate, separated basally by 0.5 width of 1 acicle, terminating subacutely, with small submarginal spine and some setae on distal margin; dorsal surface shallowly concave medially; lateral extensions subquadrate, well-developed.
</paragraph>
<paragraph id="B21736D3FFFF6271CAB25186FE93F88C" blockId="4.[151,1437,1128,2019]" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB25186FE17F938" box="[199,449,1704,1730]" italics="true" pageId="4" pageNumber="430">Antennular peduncles</emphasis>
(
<figureCitation id="2A932A56FFFF6271CBA05186FD80F938" box="[469,598,1704,1730]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="4" pageNumber="430">Fig. 2A, C</figureCitation>
) overreaching ocular peduncles by 0.3–0.4 length of ultimate segment; ultimate segment almost twice as long as penultimate segment, slightly widened distally, with tufts of long setae, one near distal margin, second medially; additional simple setae on proximal 1/3, one dorsal and one lateral; penultimate segment with few short setae; basal segment with lateral margin produced into blunt projection with narrow, blunt distal end; ventromesial distal margin produced as rounded projection; statocyst lobe unarmed; ventromesial distal angle unarmed.
</paragraph>
<paragraph id="B21736D3FFFF6270CAB250AEFB28FF2F" blockId="4.[151,1437,1128,2019]" lastBlockId="5.[151,1437,150,2013]" lastPageId="5" lastPageNumber="431" pageId="4" pageNumber="430">
<emphasis id="80DCEAC1FFFF6271CAB250AEFE75F860" box="[199,419,1920,1946]" italics="true" pageId="4" pageNumber="430">Antennal peduncles</emphasis>
(
<figureCitation id="2A932A56FFFF6271CBC450AEFDDFF860" box="[433,521,1920,1946]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="4" pageNumber="430">Fig. 2A</figureCitation>
) overreaching ocular peduncles by approximately 0.3–0.4 length of fifth segment, and slightly longer than antennular peduncles. Second segment with dorsolateral distal angle strongly produced, reaching to 3/4 of fourth peduncular segment, terminating in strong spine, mesial margin with at least 10 additional small spines; dorsomesial distal angle with acute spine. Antennal acicles arcuate, reaching to proximal 0.4–0.5 of ultimate peduncular segment, with small terminal spine; mesial margin with row of rather stiff setae.
</paragraph>
<paragraph id="B21736D3FFFE6270CAB257F1FA4DFEBB" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="431">
<emphasis id="80DCEAC1FFFE6270CAB257F1FEAFFF03" box="[199,377,223,249]" italics="true" pageId="5" pageNumber="431">Third maxiliped</emphasis>
(
<figureCitation id="2A932A56FFFE6270CBFD57F1FE0AFF03" box="[392,476,223,249]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="5" pageNumber="431">Fig. 2E</figureCitation>
) moderately stout; dactylus about 0.8 length of propodus; carpus and merus unarmed; ischium with crista dentata with about 8 corneous teeth increasing in size and more widely spaced proximally, with 1 strong accessory tooth; basis with 2 minute denticles on mesial margin; exopod reaching distal margin of carpus.
</paragraph>
<paragraph id="B21736D3FFFE6270CAB25665FBC4FB13" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="431">
<emphasis id="80DCEAC1FFFE6270CAB25665FEE1FE9F" box="[199,311,331,357]" italics="true" pageId="5" pageNumber="431">Chelipeds</emphasis>
unequal and dissimilar, right appreciably larger than left. 
<emphasis id="80DCEAC1FFFE6270C9C75665FB5AFE9F" box="[946,1164,331,357]" italics="true" pageId="5" pageNumber="431">Male right cheliped</emphasis>
(
<figureCitation id="2A932A56FFFE6270CEEF5665FAC7FE9F" box="[1178,1297,331,357]" captionStart="FIGURE 3" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[205,1391,191,1870]" captionTargetId="figure-18@8.[195,1393,181,1876]" captionTargetPageId="8" captionText="FIGURE 3. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333834" httpUri="https://zenodo.org/record/7333834/files/figure.png" pageId="5" pageNumber="431">Fig. 3A–C</figureCitation>
) moderately stout, appreciably elongated. Chela subquadrate in dorsal view, about twice as long as wide. 
<emphasis id="80DCEAC1FFFE6270CEE45641FB23FE73" box="[1169,1269,367,393]" italics="true" pageId="5" pageNumber="431">Dactylus</emphasis>
approximately same length as palm, not overlapped by fixed finger when closed; dorsal surface not elevated, smooth except for short row of small spines proximally; longitudinal rows of tufts of setae on dorsal surface and longitudinal row of stiff setae along cutting edge; dorsomesial margin defined by single row of small, obtuse tubercles, with associated tufts of moderately long stout setae; ventral surface smooth, with 3 rows of tufts of long stout setae; cutting edge with row of 7 rounded calcareous teeth, smallest medially, terminating in small corneous claw. 
<emphasis id="80DCEAC1FFFE6270CE98550DFAFEFDC7" box="[1261,1320,547,573]" italics="true" pageId="5" pageNumber="431">Palm</emphasis>
including fixed finger 1.7 times longer than carpus; with row of blunt spines on dorso-mesial margin; dorsal surface flat, smooth, with clumps of stout short setae forming circular tufts arranged in longitudinal rows (
<figureCitation id="2A932A56FFFE6270CEDB5545FAD5FD7F" box="[1198,1283,619,645]" captionStart="FIGURE 3" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[205,1391,191,1870]" captionTargetId="figure-18@8.[195,1393,181,1876]" captionTargetPageId="8" captionText="FIGURE 3. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333834" httpUri="https://zenodo.org/record/7333834/files/figure.png" pageId="5" pageNumber="431">Fig. 3F</figureCitation>
); dorsolateral margin elevated, clearly delimited by row of closely-spaced acute to subacute spines with associated short setae; space between spines widening distally, disappearing near tip; dorsomesial margin elevated, delimited by single row of spines with associated tufts of long setae; ventral surface slightly convex, smooth, with sparse tufts of long setae arranged in longitudinal rows. Cutting edge of fixed finger with 8 or 9 calcareous teeth, smallest proximally, terminating in small corneous claw. 
<emphasis id="80DCEAC1FFFE6270C8425431FD5FFCC3" box="[567,649,799,825]" italics="true" pageId="5" pageNumber="431">Carpus</emphasis>
elongated, widened distally, about twice as long as maximum width and similar in length to merus; dorsal surface smooth, with tufts of stout short setae arranged in rows more dense on lateral and mesial margins; dorsomesial margin slightly elevated and clearly delimited with row of oblique moderately strong blunt or acute spines; dorsodistal margin unarmed, with rows of setae of different size; dorsolateral margin not well-delimited, with several rows of stout short setae; lateral surface slightly convex, perpendicular, smooth and almost glabrous; mesial face slightly concave, with scattered tufts of long simple setae, ventromesial margin forming a wind-like protuberance with spines continuing the dorsomesial row; ventral surface convex, with scattered tufts of moderately long setae. 
<emphasis id="80DCEAC1FFFE6270C8235335FD48FBCE" box="[598,670,1051,1076]" italics="true" pageId="5" pageNumber="431">Merus</emphasis>
subtriangular; dorsal surface with low protuberances with associated short setae; dorsodistal margin with 2 large and 1 smaller acute spines with associated short setae; lateral surface slightly convex, smooth, with few very short scattered setae; mesial surface slightly concave with scattered long setae; ventromesial margin smooth except for small spinulose area distally; ventrolateral margin not well-defined, rounded; ventral surface smooth, with scattered long setae. Ischium unarmed. 
<emphasis id="80DCEAC1FFFE6270CE675385FADAFB3F" box="[1042,1292,1195,1221]" italics="true" pageId="5" pageNumber="431">Female right cheliped</emphasis>
differs from males in length, being less elongated but similar in shape and armature to male.
</paragraph>
<paragraph id="B21736D3FFFE6270CAB253DDFB44F963" blockId="5.[151,1437,150,2013]" pageId="5" pageNumber="431">
<emphasis id="80DCEAC1FFFE6270CAB253DDFE5BFAF7" box="[199,397,1267,1293]" italics="true" pageId="5" pageNumber="431">Male left cheliped</emphasis>
(
<figureCitation id="2A932A56FFFE6270CBEF53DDFDDAFAF7" box="[410,524,1267,1293]" captionStart="FIGURE 3" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[205,1391,191,1870]" captionTargetId="figure-18@8.[195,1393,181,1876]" captionTargetPageId="8" captionText="FIGURE 3. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333834" httpUri="https://zenodo.org/record/7333834/files/figure.png" pageId="5" pageNumber="431">Fig. 3D, E</figureCitation>
) generally similar to right. 
<emphasis id="80DCEAC1FFFE6270C94153DDFC4EFAF7" box="[820,920,1267,1293]" italics="true" pageId="5" pageNumber="431">Dactylus</emphasis>
approximately 1.5 length of palm, dorsomesial margin with row of closely-spaced acute tubercles; dorsal surface smooth, with rows of tufts of moderately long setae; masticatory border toothless, except some very small proximally; rest of masticatory border bearing closelyspaced stout setae, forming a well-developed brush-like structure (
<figureCitation id="2A932A56FFFE6270C9085271FC05FA83" box="[893,979,1375,1401]" captionStart="FIGURE 3" captionStartId="8.[151,250,1900,1924]" captionTargetBox="[205,1391,191,1870]" captionTargetId="figure-18@8.[195,1393,181,1876]" captionTargetPageId="8" captionText="FIGURE 3. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333834" httpUri="https://zenodo.org/record/7333834/files/figure.png" pageId="5" pageNumber="431">Fig. 3D</figureCitation>
inset). Palm with row of subacute spines on dorsomesial margin, ending distally in two conspicuous spines; dorsal surface almost flat, unarmed; dorsolateral margin slightly raised and with row of closely-spaced subacute tubercles not extending to tip of fixed finger. 
<emphasis id="80DCEAC1FFFE6270CF3F5289FA4AFA3B" box="[1354,1436,1447,1473]" italics="true" pageId="5" pageNumber="431">Carpus</emphasis>
with row of subacute oblique spines on dorsomesial margin; dorsal surface smooth, with longitudinal ridges of tufts of stiff setae; dorsodistal margin with several small spines. 
<emphasis id="80DCEAC1FFFE6270C95352C1FCB8F9F2" box="[806,878,1519,1544]" italics="true" pageId="5" pageNumber="431">Merus</emphasis>
with 2 spines on dorsodistal margin; ventromesial and ventrolateral margins each with row, or only few, very small blunt spinules or tubercles. 
<emphasis id="80DCEAC1FFFE6270CEC0513DFA4AF9D7" box="[1205,1436,1555,1581]" italics="true" pageId="5" pageNumber="431">Female left cheliped</emphasis>
similar in shape to male, with more tubercles especially evident at dorsolateral surface of carpus, which is slightly elevated bearing some low tridentate tubercles with associated stiff setae, and merus, with dorsal surface crossed by rows of spinose sulcus, continuing along lateral and mesial surfaces, especially distal ones.
</paragraph>
<paragraph id="B21736D3FFFE6273CAB2518DFE33FEBB" blockId="5.[151,1437,150,2013]" lastBlockId="6.[151,1437,151,1905]" lastPageId="6" lastPageNumber="432" pageId="5" pageNumber="431">
<emphasis id="80DCEAC1FFFE6270CAB2518DFE54F947" box="[199,386,1699,1725]" italics="true" pageId="5" pageNumber="431">Ambulatory legs</emphasis>
(
<figureCitation id="2A932A56FFFE6270CBE6518DFDD9F947" box="[403,527,1699,1725]" captionStart="FIGURE 4" captionStartId="9.[151,250,1866,1890]" captionTargetBox="[222,1380,187,1836]" captionTargetId="figure-24@9.[180,1407,181,1842]" captionTargetPageId="9" captionText="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="5" pageNumber="431">Fig. 4A, B</figureCitation>
) short and stout. 
<emphasis id="80DCEAC1FFFE6270C8AE518DFCE9F947" box="[731,831,1699,1725]" italics="true" pageId="5" pageNumber="431">Dactylus</emphasis>
about same length (second) to 1.2 times longer than propodus (third) and about 5 times longer than broad, flattened laterally and gently curved ventrally terminating in strong corneous claw; lateral surface smooth, with very shallow longitudinal sulcus and few tufts of short setae on lower half; dorsal margin with irregular rows of tufts of short, thick setae, ending in row of long setae occupying distal 1/5; ventro-inner margins each with row of 5 or 6 (second) to 7 or 8 (third) strong corneous spines increasing in size distally, evenly distributed except most proximal of second pereiopod, which is more separated, and associated tufts of long setae. 
<emphasis id="80DCEAC1FFFE6270CB4C5055FE73F86F" box="[313,421,1915,1941]" italics="true" pageId="5" pageNumber="431">Propodus</emphasis>
approximately 1.4 times longer than carpus (second and third); dorsal surfaces unarmed, each with multiple rows of short thick setae; lateral surface smooth, with longitudinal rows of short setae; ventral margins each with 3 corneous spines and tufts of short setae, denser near distal margin. 
<emphasis id="80DCEAC1FFFE6270CE1A50EDFB17F827" box="[1135,1217,1987,2013]" italics="true" pageId="5" pageNumber="431">Carpus</emphasis>
with dorsal margin defined by row of small spines and associate tufts of short setae, with strong spine at dorsodistal margin; several rows of tufts of setae on lateral faces, short at upper half and near lower margin, longer at midline; ventral margin unarmed and glabrous. 
<emphasis id="80DCEAC1FFFD6273CBE857F1FE33FF02" box="[413,485,223,248]" italics="true" pageId="6" pageNumber="432">Merus</emphasis>
with dentate dorsal margin, with associated rows of setae; lateral surface smooth and glabrous; ventral margins unarmed or with low protuberances or spinules, particularly on second pereiopods, and row of moderately long setae.
</paragraph>
<paragraph id="B21736D3FFFD6273CAB25665FB30FE0F" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB25665FE41FE9F" box="[199,407,331,357]" italics="true" pageId="6" pageNumber="432">Fourth pereiopods</emphasis>
(
<figureCitation id="2A932A56FFFD6273CBD05665FE25FE9F" box="[421,499,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1866,1890]" captionTargetBox="[222,1380,187,1836]" captionTargetId="figure-24@9.[180,1407,181,1842]" captionTargetPageId="9" captionText="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="6" pageNumber="432">Fig 4E</figureCitation>
) semichelate, subequal in length from left to right; dactylus with subterminal tuft of short setae on dorsal margin; carpus ending in conspicuous spine at distodorsal margin; stiff long setae and tufts of setae on dorsal margin of propodus to carpus and distodorsal and ventromesial margin of merus. Dactylus nearly straight, with convex dorsal margin, terminating in small, strongly-curved corneous claw; ventral margin with closely-set, microscopic corneous teeth; propodal rasp consisting of single row of corneous scales.
</paragraph>
<paragraph id="B21736D3FFFD6273CAB256D1FC2CFDE3" blockId="6.[151,1437,151,1905]" box="[199,1018,511,537]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB256D1FE56FDE3" box="[199,384,511,537]" italics="true" pageId="6" pageNumber="432">Fifth pereiopods</emphasis>
chelate, propodal rasp occupying distal half of segment.
</paragraph>
<paragraph id="B21736D3FFFD6273CAB2550DFE5BFCEF" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB2550DFE64FDC6" box="[199,434,547,573]" italics="true" pageId="6" pageNumber="432">Sternite of somite XII</emphasis>
in males (thoracomere 6, third pereiopods) with roundly subrectangular anterior lobe (
<figureCitation id="2A932A56FFFD6273CF1A550DFF6EFD9B" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[193,1395,193,1836]" captionTargetId="figure-24@7.[181,1407,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="6" pageNumber="432">Fig. 2F</figureCitation>
), bearing setae of different sizes on anterior margin. 
<emphasis id="80DCEAC1FFFD6273C96E5569FBCFFD9A" box="[795,1049,583,609]" italics="true" pageId="6" pageNumber="432">Sternite of somite XIV</emphasis>
(thoracomere 8, fifth pereiopods) with widely separated lobes in males, each with moderately long setae (
<figureCitation id="2A932A56FFFD6273C9C65545FBDDFD7F" box="[947,1035,619,645]" captionStart="FIGURE 4" captionStartId="9.[151,250,1866,1890]" captionTargetBox="[222,1380,187,1836]" captionTargetId="figure-24@9.[180,1407,181,1842]" captionTargetPageId="9" captionText="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="6" pageNumber="432">Fig. 4C</figureCitation>
); coxae of male each with gonopore partially masked by tuft of setae. 
<emphasis id="80DCEAC1FFFD6273C86E55A1FCC7FD52" box="[539,785,655,681]" italics="true" pageId="6" pageNumber="432">Sternite of somite XII</emphasis>
of females, with subquadrate single central lobe bearing moderately long simple setae (
<figureCitation id="2A932A56FFFD6273CB87559DFD98FD37" box="[498,590,691,717]" captionStart="FIGURE 4" captionStartId="9.[151,250,1866,1890]" captionTargetBox="[222,1380,187,1836]" captionTargetId="figure-24@9.[180,1407,181,1842]" captionTargetPageId="9" captionText="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="6" pageNumber="432">Fig. 4D</figureCitation>
); coxa of females each with gonopore, and associated tufts of setae above, without masking them. Males with 3 uniramous unpaired left 
<emphasis id="80DCEAC1FFFD6273C92155F9FC6FFD0B" box="[852,953,727,753]" italics="true" pageId="6" pageNumber="432">pleopods</emphasis>
; females with slightly unequally biramous 2-4, fifth uniramous..
</paragraph>
<paragraph id="B21736D3FFFD6273CAB25431FABAFC5F" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB25431FED9FCC3" box="[199,271,799,825]" italics="true" pageId="6" pageNumber="432">Telson</emphasis>
(
<figureCitation id="2A932A56FFFD6273CB535431FE56FCC3" box="[294,384,799,825]" captionStart="FIGURE 4" captionStartId="9.[151,250,1866,1890]" captionTargetBox="[222,1380,187,1836]" captionTargetId="figure-24@9.[180,1407,181,1842]" captionTargetPageId="9" captionText="FIGURE 4. Pagurus pectinidactylus n. sp., Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. Pagurus pectinidactylus n. sp. Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) Telson, dorsal view. (Scales: A–F = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="6" pageNumber="432">Fig. 4F</figureCitation>
) with lateral incision indicating anterior and posterior portions; posterior lobes slightly asymmetrical, subquadrate, left slightly larger than right, separated by very small median cleft; terminal margins each with row of calcareous large teeth alternating with small ones, extending into lateral margin on left lobe and to posterior half of right; few short setae associated with spines and some tufts of longer setae on lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="FAB26558FFFD6273CAB25480FE96FC17" pageId="6" pageNumber="432" type="distribution">
<paragraph id="B21736D3FFFD6273CAB25480FE96FC17" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB25480FEB7FC32" bold="true" box="[199,353,942,968]" pageId="6" pageNumber="432">Distribution.</emphasis>
So far only know from two off-shore stations, 
<quantity id="75509B36FFFD6273C9065481FC00FC33" box="[883,982,943,969]" metricMagnitude="1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="6" pageNumber="432" unit="m" value="25.5" valueMax="30.0" valueMin="21.0">21–30 m</quantity>
depth, within Mauritanian waters, north of 
<collectingRegion id="706CF831FFFD6273CAC354FDFEEAFC17" box="[182,316,979,1005]" country="Mauritania" name="Nouakchott" pageId="6" pageNumber="432">Nouakchott</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FAB26558FFFD6273CAB254D9FE25FBA3" pageId="6" pageNumber="432" type="etymology">
<paragraph id="B21736D3FFFD6273CAB254D9FE25FBA3" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB254D9FE9EFBEB" bold="true" box="[199,328,1015,1041]" pageId="6" pageNumber="432">Etymology</emphasis>
. Derived from Latin pecten (= comb) and Greek δάχτμλος (dáctilos = finger), referring to the characteristic row of closely-spaced stout setae on the masticatory border of left cheliped dactylus, forming a welldeveloped brush-like structure.
</paragraph>
</subSubSection>
<subSubSection id="FAB26558FFFD627FCAB2534CFD74FAAF" lastPageId="10" lastPageNumber="436" pageId="6" pageNumber="432" type="discussion">
<paragraph id="B21736D3FFFD6273CAB2534CFB5FF9F3" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
<emphasis id="80DCEAC1FFFD6273CAB2534CFE16FB87" bold="true" box="[199,448,1122,1149]" pageId="6" pageNumber="432">Taxonomic remarks.</emphasis>
The new species is assignable to the informal 
<taxonomicName id="75A84D50FFFD6273C995534DFB04FB87" authorityName="Risso" authorityYear="1827" box="[992,1234,1123,1149]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="80DCEAC1FFFD6273C995534DFB04FB87" box="[992,1234,1123,1149]" italics="true" pageId="6" pageNumber="432">Pagurus anachoretus</emphasis>
</taxonomicName>
group (cf. 
<bibRefCitation id="D6394B22FFFD6273CF23534DFF06FB5B" author="Forest J." pageId="6" pageNumber="432" pagination="525 - 538" refId="ref8388" refString="Forest J. (1978) Sur deux pagurides nouveaux de l'Atlantique tropical africain: Pagurus laurentae et Paguristes cyanops spp. nov. Bulletin du Museium national d'histoire naturelle, Paris, 3 e serie Zoologie, 356, 525 - 538." type="journal article" year="1978">Forest 1978</bibRefCitation>
) as it concurs with the general definition of the group (
<bibRefCitation id="D6394B22FFFD6273C94853A9FB82FB5B" author="Forest, J. &amp; Ngoc-Ho, N." box="[829,1108,1159,1185]" pageId="6" pageNumber="432" pagination="217 - 227" refId="ref8432" refString="Forest, J. &amp; Ngoc-Ho, N. (1992) Desription de Pagurus dartevellei (Forest, 1958) (Crustacea, Decapoda, Paguridae). Bulletin du Museum national d'histoire naturelle Section A Zoologie, biologie et ecologie animales, 14, 217 - 227." type="journal article" year="1992">Forest &amp; Ngoc-Ho 1992</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273CE1553A9FA41FB5B" author="McLaughlin, P. A. &amp; Forest, J." box="[1120,1431,1159,1185]" pageId="6" pageNumber="432" pagination="297 - 344" refId="ref9806" refString="McLaughlin, P. A. &amp; Forest, J. (1999) Hermit crabs of the genus Pagurus Fabricius (Crustacea, Decapoda, Paguridae) from south-eastern South Africa. Annals of the South African Museum, 105, 297 - 344." type="journal article" year="1999">McLaughlin &amp; Forest 1999</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273CAE25385FE4BFB3F" author="Komai, T. &amp; Rahayu, D. L." box="[151,413,1195,1221]" pageId="6" pageNumber="432" pagination="183 - 200" refId="ref9228" refString="Komai, T. &amp; Rahayu, D. L. (2004) Redescription of Pagurus moluccensis Haig &amp; Ball, 1988, with description of a new species of Pagurus from Indonesia, and taxonomic notes on the Pagurus anachoretus group (Crustacea: Decapoda: Anomura: Paguridae). Raffles Bulletin of Zoology, 52, 183 - 200." type="journal article" year="2004">Komai &amp; Rahayu 2004</bibRefCitation>
). This group includes 12 species at present: 
<taxonomicName id="75A84D50FFFD6273C9F55385FBBBFB3F" authorityName="Risso" authorityYear="1827" box="[896,1133,1195,1221]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="80DCEAC1FFFD6273C9F55385FBBBFB3F" box="[896,1133,1195,1221]" italics="true" pageId="6" pageNumber="432">Pagurus anachoretus</emphasis>
</taxonomicName>
from the Mediterranean, 
<taxonomicName id="75A84D50FFFD6273CFFD5385FE95FB13" authorityName="Forest" authorityYear="1966" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="anachoretoides">
<emphasis id="80DCEAC1FFFD6273CFFD5385FE95FB13" italics="true" pageId="6" pageNumber="432">P. anachoretoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273CB3B53E1FE06FB13" baseAuthorityName="Forest" baseAuthorityYear="1956" box="[334,464,1231,1257]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="gordonae">
<emphasis id="80DCEAC1FFFD6273CB3B53E1FE06FB13" box="[334,464,1231,1257]" italics="true" pageId="6" pageNumber="432">P.gordonae</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273CBAE53E1FD8BFB13" authorityName="Forest" authorityYear="1978" box="[475,605,1231,1257]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="laurentae">
<emphasis id="80DCEAC1FFFD6273CBAE53E1FD8BFB13" box="[475,605,1231,1257]" italics="true" pageId="6" pageNumber="432">P.laurentae</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFFD6273C8FB53E1FD22FB12" baseAuthorityName="Forest" baseAuthorityYear="1952" box="[654,756,1231,1256]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="souriei">
<emphasis id="80DCEAC1FFFD6273C8FB53E1FD22FB12" box="[654,756,1231,1256]" italics="true" pageId="6" pageNumber="432">P.souriei</emphasis>
</taxonomicName>
from the tropical eastern Atlantic, 
<taxonomicName id="75A84D50FFFD6273CE1D53E1FB2DFB12" authorityName="McLaughlin &amp; Forest" authorityYear="1999" box="[1128,1275,1231,1256]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="emmersoni">
<emphasis id="80DCEAC1FFFD6273CE1D53E1FB2DFB12" box="[1128,1275,1231,1256]" italics="true" pageId="6" pageNumber="432">P.emmersoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFFD6273CF5E53E1FA4BFB13" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[1323,1437,1231,1257]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFFD6273CF5E53E1FA4BFB13" box="[1323,1437,1231,1257]" italics="true" pageId="6" pageNumber="432">P.liochele</emphasis>
</taxonomicName>
from 
<collectingCountry id="CABF7643FFFD6273CAA053DDFEB0FAF7" box="[213,358,1267,1293]" name="South Africa" pageId="6" pageNumber="432">South Africa</collectingCountry>
, and the Indo-West Pacific 
<taxonomicName id="75A84D50FFFD6273C8E853DDFBA9FAF7" authority="Komai &amp; Osawa, 2001" authorityName="Komai &amp; Osawa" authorityYear="2001" box="[669,1151,1267,1293]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="decimbranchiae">
<emphasis id="80DCEAC1FFFD6273C8E853DDFD78FAF6" box="[669,686,1267,1292]" italics="true" pageId="6" pageNumber="432">P</emphasis>
. 
<emphasis id="80DCEAC1FFFD6273C8C953DDFCA6FAF7" box="[700,880,1267,1293]" italics="true" pageId="6" pageNumber="432">decimbranchiae</emphasis>
<bibRefCitation id="D6394B22FFFD6273C90D53DDFBA9FAF7" author="Komai, T. &amp; Osawa, M." box="[888,1151,1267,1293]" pageId="6" pageNumber="432" pagination="1291 - 1301" refId="ref9177" refString="Komai, T. &amp; Osawa, M. (2001) A new distinctive species of pagurid hermit crab (Crustacea: Decapoda: Anomura) from Japan. Zoological Science, 18, 1291 - 1301. https: // doi. org / 10.2108 / zsj. 18.1291" type="journal article" year="2001">Komai &amp; Osawa, 2001</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273CEFF53DDFEE0FACB" authority="Komai &amp; Rahayu, 2004" authorityName="Komai &amp; Rahayu" authorityYear="2004" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="fungiformis">
<emphasis id="80DCEAC1FFFD6273CEFF53DDFB4DFAF6" box="[1162,1179,1267,1292]" italics="true" pageId="6" pageNumber="432">P</emphasis>
. 
<emphasis id="80DCEAC1FFFD6273CEDF53DDFAFDFAF7" box="[1194,1323,1267,1293]" italics="true" pageId="6" pageNumber="432">fungiformis</emphasis>
<bibRefCitation id="D6394B22FFFD6273CF4653DDFEE0FACB" author="Komai, T. &amp; Rahayu, D. L." pageId="6" pageNumber="432" pagination="183 - 200" refId="ref9228" refString="Komai, T. &amp; Rahayu, D. L. (2004) Redescription of Pagurus moluccensis Haig &amp; Ball, 1988, with description of a new species of Pagurus from Indonesia, and taxonomic notes on the Pagurus anachoretus group (Crustacea: Decapoda: Anomura: Paguridae). Raffles Bulletin of Zoology, 52, 183 - 200." type="journal article" year="2004">Komai &amp; Rahayu, 2004</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273CB345239FD24FACB" authority="(Grant &amp; McCulloch, 1906)" baseAuthorityName="Grant &amp; McCulloch" baseAuthorityYear="1906" box="[321,754,1303,1329]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="hedleyi">
<emphasis id="80DCEAC1FFFD6273CB345239FE78FACB" box="[321,430,1303,1329]" italics="true" pageId="6" pageNumber="432">P. hedleyi</emphasis>
(
<bibRefCitation id="D6394B22FFFD6273CBC95239FD3CFACB" author="Grant, F. E. &amp; McCulloch, A. R." box="[444,746,1303,1329]" pageId="6" pageNumber="432" pagination="2 - 53" refId="ref8900" refString="Grant, F. E. &amp; McCulloch, A. R. (1906) On a collection of Crustacea from the Port Curtis district, Queensland. Proceedings of the Linnean Society of New South Wales, 31 (1), 2 - 53." type="journal article" year="1906">Grant &amp; McCulloch, 1906</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273C88B5239FBFFFACB" authority="Sankolli, 1961" authorityName="Sankolli" authorityYear="1961" box="[766,1065,1303,1329]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="kulkarnii">
<emphasis id="80DCEAC1FFFD6273C88B5239FCA9FACB" box="[766,895,1303,1329]" italics="true" pageId="6" pageNumber="432">P. kulkarnii</emphasis>
Sankolli, 1961
</taxonomicName>
and 
<taxonomicName id="75A84D50FFFD6273CE155239FF19FAAF" authority="Haig &amp; Ball, 1998" authorityName="Haig &amp; Ball" authorityYear="1998" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="moluccensis">
<emphasis id="80DCEAC1FFFD6273CE155239FAD3FACB" box="[1120,1285,1303,1329]" italics="true" pageId="6" pageNumber="432">P. moluccensis</emphasis>
Haig &amp; Ball, 1998
</taxonomicName>
(see 
<bibRefCitation id="D6394B22FFFD6273CB7E5215FE45FAAF" author="Forest J." box="[267,403,1339,1365]" pageId="6" pageNumber="432" pagination="525 - 538" refId="ref8388" refString="Forest J. (1978) Sur deux pagurides nouveaux de l'Atlantique tropical africain: Pagurus laurentae et Paguristes cyanops spp. nov. Bulletin du Museium national d'histoire naturelle, Paris, 3 e serie Zoologie, 356, 525 - 538." type="journal article" year="1978">Forest 1978</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273CBEA5215FD65FAAF" author="Forest, J. &amp; Ngoc-Ho, N." box="[415,691,1339,1365]" pageId="6" pageNumber="432" pagination="217 - 227" refId="ref8432" refString="Forest, J. &amp; Ngoc-Ho, N. (1992) Desription de Pagurus dartevellei (Forest, 1958) (Crustacea, Decapoda, Paguridae). Bulletin du Museum national d'histoire naturelle Section A Zoologie, biologie et ecologie animales, 14, 217 - 227." type="journal article" year="1992">Forest &amp; Ngoc-Ho 1992</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273C8CA5215FC21FAAF" author="McLaughlin, P. A. &amp; Forest, J." box="[703,1015,1339,1365]" pageId="6" pageNumber="432" pagination="297 - 344" refId="ref9806" refString="McLaughlin, P. A. &amp; Forest, J. (1999) Hermit crabs of the genus Pagurus Fabricius (Crustacea, Decapoda, Paguridae) from south-eastern South Africa. Annals of the South African Museum, 105, 297 - 344." type="journal article" year="1999">McLaughlin &amp; Forest 1999</bibRefCitation>
). Four other species from the tropical eastern Atlantic, 
<taxonomicName id="75A84D50FFFD6273CB205271FE14FA83" baseAuthorityName="Balss" baseAuthorityYear="1911" box="[341,450,1375,1401]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="80DCEAC1FFFD6273CB205271FE14FA83" box="[341,450,1375,1401]" italics="true" pageId="6" pageNumber="432">P. alcocki</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273CBA45271FCD8FA83" authority="(Forest, 1958)" baseAuthorityName="Forest" baseAuthorityYear="1958" box="[465,782,1374,1401]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="dartevellei">
<emphasis id="80DCEAC1FFFD6273CBA45271FDB3FA83" box="[465,613,1375,1401]" italics="true" pageId="6" pageNumber="432">P. dartevellei</emphasis>
(
<bibRefCitation id="D6394B22FFFD6273C8015271FCD0FA83" author="Forest, J." box="[628,774,1374,1401]" pageId="6" pageNumber="432" pagination="144 - 168" refId="ref8267" refString="Forest, J. (1958) Les Crustaces Anomoures du Musee royal du Congo belge. Revue de Zoologie et de Botanique Africaines, 58, 144 - 168." type="journal article" year="1958">Forest, 1958</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="75A84D50FFFD6273C96F5271FC7FFA83" authorityName="Forest" authorityYear="1966" box="[794,937,1375,1401]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFFD6273C96F5271FC7FFA83" box="[794,937,1375,1401]" italics="true" pageId="6" pageNumber="432">P. fimbriatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFFD6273C9945271FB55FA83" baseAuthorityName="Chevreux &amp; Bouvier" baseAuthorityYear="1982" box="[993,1155,1375,1401]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="80DCEAC1FFFD6273C9945271FB55FA83" box="[993,1155,1375,1401]" italics="true" pageId="6" pageNumber="432">P. triangularis</emphasis>
</taxonomicName>
, also fit the definition of the 
<taxonomicName id="75A84D50FFFD6273CAB552ADFEB5FA67" authorityName="Risso" authorityYear="1827" box="[192,355,1411,1437]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="80DCEAC1FFFD6273CAB552ADFEB5FA67" box="[192,355,1411,1437]" italics="true" pageId="6" pageNumber="432">P. anachoretus</emphasis>
</taxonomicName>
group except for the cheliped form (
<bibRefCitation id="D6394B22FFFD6273C88C52ADFC54FA67" author="Forest J." box="[761,898,1411,1437]" pageId="6" pageNumber="432" pagination="525 - 538" refId="ref8388" refString="Forest J. (1978) Sur deux pagurides nouveaux de l'Atlantique tropical africain: Pagurus laurentae et Paguristes cyanops spp. nov. Bulletin du Museium national d'histoire naturelle, Paris, 3 e serie Zoologie, 356, 525 - 538." type="journal article" year="1978">Forest 1978</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273C9F852ADFB49FA67" author="Forest, J. &amp; Ngoc-Ho, N." box="[909,1183,1411,1437]" pageId="6" pageNumber="432" pagination="217 - 227" refId="ref8432" refString="Forest, J. &amp; Ngoc-Ho, N. (1992) Desription de Pagurus dartevellei (Forest, 1958) (Crustacea, Decapoda, Paguridae). Bulletin du Museum national d'histoire naturelle Section A Zoologie, biologie et ecologie animales, 14, 217 - 227." type="journal article" year="1992">Forest &amp; Ngoc-Ho 1992</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273CEDF52ADFF04FA3B" author="McLaughlin, P. A. &amp; Forest, J." pageId="6" pageNumber="432" pagination="297 - 344" refId="ref9806" refString="McLaughlin, P. A. &amp; Forest, J. (1999) Hermit crabs of the genus Pagurus Fabricius (Crustacea, Decapoda, Paguridae) from south-eastern South Africa. Annals of the South African Museum, 105, 297 - 344." type="journal article" year="1999">McLaughlin &amp; Forest 1999</bibRefCitation>
; 
<bibRefCitation id="D6394B22FFFD6273CAA85289FE35FA3B" author="Komai, T. &amp; Rahayu, D. L." box="[221,483,1447,1473]" pageId="6" pageNumber="432" pagination="183 - 200" refId="ref9228" refString="Komai, T. &amp; Rahayu, D. L. (2004) Redescription of Pagurus moluccensis Haig &amp; Ball, 1988, with description of a new species of Pagurus from Indonesia, and taxonomic notes on the Pagurus anachoretus group (Crustacea: Decapoda: Anomura: Paguridae). Raffles Bulletin of Zoology, 52, 183 - 200." type="journal article" year="2004">Komai &amp; Rahayu 2004</bibRefCitation>
). 
<taxonomicName id="75A84D50FFFD6273CB805289FD01FA3B" authorityName="Landschoff" authorityYear="2018" box="[501,727,1447,1473]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="fraserorum">
<emphasis id="80DCEAC1FFFD6273CB805289FD01FA3B" box="[501,727,1447,1473]" italics="true" pageId="6" pageNumber="432">Pagurus fraserorum</emphasis>
</taxonomicName>
described by 
<bibRefCitation id="D6394B22FFFD6273C9055289FB96FA3B" author="Landschoff, J. &amp; Komai, T. &amp; du Plessis, A. &amp; Gouws, G. &amp; Griffiths, C. L." box="[880,1088,1447,1473]" pageId="6" pageNumber="432" pagination="0203107" refId="ref9403" refString="Landschoff, J., Komai, T., du Plessis, A., Gouws, G. &amp; Griffiths, C. L. (2018) MicroCT imaging applied to description of a new species of Pagurus Fabricius, 1775 (Crustacea: Decapoda: Anomura: Paguridae), with selection of three-dimensional type data. PLoS ONE, 13 (9), e 0203107. https: // doi. org / 10.1371 / journal. pone. 0203107" type="journal article" year="2018">Landschoff (2018)</bibRefCitation>
also shares the main characters of the group except for the shape of the right cheliped, and based on molecular results, it has been suggested that it may constitute a separate group, more related with 
<taxonomicName id="75A84D50FFFD6273C8A152C1FC67F9F3" authorityName="Morgan" authorityYear="1990" box="[724,945,1519,1545]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="boriaustraliensis">
<emphasis id="80DCEAC1FFFD6273C8A152C1FD33F9F2" box="[724,741,1519,1544]" italics="true" pageId="6" pageNumber="432">P</emphasis>
. 
<emphasis id="80DCEAC1FFFD6273C88652C1FC67F9F3" box="[755,945,1519,1545]" italics="true" pageId="6" pageNumber="432">boriaustraliensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFFD6273C99D52C1FB54F9F3" authorityName="McLaughlin" authorityYear="2002" box="[1000,1154,1519,1545]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="pitagsaleei">
<emphasis id="80DCEAC1FFFD6273C99D52C1FC2FF9F2" box="[1000,1017,1519,1544]" italics="true" pageId="6" pageNumber="432">P</emphasis>
. 
<emphasis id="80DCEAC1FFFD6273CE7252C1FB54F9F3" box="[1031,1154,1519,1545]" italics="true" pageId="6" pageNumber="432">pitagsaleei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B21736D3FFFD6273CAB2513DFDA2F8D3" blockId="6.[151,1437,151,1905]" pageId="6" pageNumber="432">
Notwithstanding, the new species shows enough morphological differences to avoid any confusion with the species of the group or any other congeners known so far. The oblique calcified scale present in the base of the peduncles, is not known to occur in any other species of 
<taxonomicName id="75A84D50FFFD6273C95B5175FC5BF98E" authorityName="Fabricius" authorityYear="1775" box="[814,909,1627,1652]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="80DCEAC1FFFD6273C95B5175FC5BF98E" box="[814,909,1627,1652]" italics="true" pageId="6" pageNumber="432">Pagurus</emphasis>
</taxonomicName>
. The shape and armature of the chelipeds are diagnostic, with a subquadrate shape in dorsal view, clearly elongate. The palm surface is flat and smooth, but with numerous distinctive clumps of stout short setae forming circular tufts arranged in longitudinal rows. The masticatory border of the left cheliped dactylus is also very conspicuous, lacking teeth, but bearing instead a row of closely-spaced stout setae, forming a well-developed brush-like structure, which has not been described for any other species of 
<taxonomicName id="75A84D50FFFD6273CB3B5021FE7BF8D2" authorityName="Fabricius" authorityYear="1775" box="[334,429,1807,1832]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="80DCEAC1FFFD6273CB3B5021FE7BF8D2" box="[334,429,1807,1832]" italics="true" pageId="6" pageNumber="432">Pagurus</emphasis>
</taxonomicName>
in the study area.
</paragraph>
<paragraph id="B21736D3FFFD627FCAB2501DFA83FD9B" blockId="6.[151,1437,151,1905]" lastBlockId="10.[151,1437,151,1365]" lastPageId="10" lastPageNumber="436" pageId="6" pageNumber="432">
Although those characters are sufficiently distinctive to avoid any confusions, the new species shows a closer morphological resemblance with some species of the 
<taxonomicName id="75A84D50FFFD6273C96D5079FC77F88B" authorityName="Risso" authorityYear="1827" box="[792,929,1879,1905]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="80DCEAC1FFFD6273C96D5079FC77F88B" box="[792,929,1879,1905]" italics="true" pageId="6" pageNumber="432">anachoretus</emphasis>
</taxonomicName>
group, specifically with 
<taxonomicName id="75A84D50FFFD6273CEA45079FA40F88B" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[1233,1430,1879,1905]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="432" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFFD6273CEA45079FA40F88B" box="[1233,1430,1879,1905]" italics="true" pageId="6" pageNumber="432">Pagurus liochele</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFF1627FCAE257B9FEE7FF4A" authorityName="McLaughlin &amp; Forest" authorityYear="1999" box="[151,305,151,176]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="emmersoni">
<emphasis id="80DCEAC1FFF1627FCAE257B9FEE7FF4A" box="[151,305,151,176]" italics="true" pageId="10" pageNumber="436">P. emmersoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFF1627FCB3757B9FE31FF4B" baseAuthorityName="Chevreux &amp; Bouvier" baseAuthorityYear="1982" box="[322,487,151,177]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="80DCEAC1FFF1627FCB3757B9FE31FF4B" box="[322,487,151,177]" italics="true" pageId="10" pageNumber="436">P. triangularis</emphasis>
</taxonomicName>
, 
<taxonomicName id="75A84D50FFF1627FCB8C57B9FD5DFF4B" authorityName="Forest" authorityYear="1966" box="[505,651,151,177]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFF1627FCB8C57B9FD5DFF4B" box="[505,651,151,177]" italics="true" pageId="10" pageNumber="436">P. fimbriatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFF1627FC8BD57B9FCB8FF4B" baseAuthorityName="Balss" baseAuthorityYear="1921" box="[712,878,151,177]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="similimanus">
<emphasis id="80DCEAC1FFF1627FC8BD57B9FCB8FF4B" box="[712,878,151,177]" italics="true" pageId="10" pageNumber="436">P. similimanus</emphasis>
</taxonomicName>
. The first two are considered endemic to South African waters. The new species differs from the two South African congeners in having broader carapace and rostrum, longer ocular peduncles, and more ovate ocular acicles. The antennules are shorter in 
<taxonomicName id="75A84D50FFF1627FCE8F57F1FA43FF02" authorityName="McLaughlin &amp; Forest" authorityYear="1999" box="[1274,1429,223,248]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="emmersoni">
<emphasis id="80DCEAC1FFF1627FCE8F57F1FA43FF02" box="[1274,1429,223,248]" italics="true" pageId="10" pageNumber="436">P. emmersoni</emphasis>
</taxonomicName>
, overreaching distal margin of corneas by approximately 0.2 length of antennular segment 3, while they are similar in 
<taxonomicName id="75A84D50FFF1627FCAC05609FEAAFEBB" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[181,380,295,321]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF1627FCAC05609FEAAFEBB" box="[181,380,295,321]" italics="true" pageId="10" pageNumber="436">P. pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF1627FCBF15609FE15FEBB" bold="true" box="[388,451,295,321]" pageId="10" pageNumber="436">
<taxonomicNameLabel id="9BEF57BAFFF1627FCBF15609FE15FEBB" box="[388,451,295,321]" pageId="10" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="75A84D50FFF1627FCB885609FDA4FEBB" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[509,626,295,321]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFF1627FCB885609FDA4FEBB" box="[509,626,295,321]" italics="true" pageId="10" pageNumber="436">P. liochele</emphasis>
</taxonomicName>
, overreaching the ocular peduncles by approximately 0.3–0.4 length of antennular segment 3. Another distinctive character can be found in the antennal segment 2 dorsolateral distal angle, which is strongly produced in both 
<taxonomicName id="75A84D50FFF1627FC8695641FD59FE73" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[540,655,367,393]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFF1627FC8695641FD59FE73" box="[540,655,367,393]" italics="true" pageId="10" pageNumber="436">P. liochele</emphasis>
</taxonomicName>
and 
<taxonomicName id="75A84D50FFF1627FC8B75641FC51FE73" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[706,903,367,393]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF1627FC8B75641FC51FE73" box="[706,903,367,393]" italics="true" pageId="10" pageNumber="436">P. pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF1627FC9F95641FC1FFE73" bold="true" box="[908,969,367,393]" pageId="10" pageNumber="436">
<taxonomicNameLabel id="9BEF57BAFFF1627FC9F95641FC1FFE73" box="[908,969,367,393]" pageId="10" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, with mesial margin more densely dentate in the new species with at least 10 small spines, while in 
<taxonomicName id="75A84D50FFF1627FC96356BDFC5CFE57" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[790,906,403,429]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFF1627FC96356BDFC5CFE57" box="[790,906,403,429]" italics="true" pageId="10" pageNumber="436">P. liochele</emphasis>
</taxonomicName>
there are 1–5. Moreover, the new species has a longer palm, with the characteristic clumps of setae arranged in circles on dorsal surface instead of any tubercles or spines, character that has not been described in any of the other species. The pereiopods are stout but more elongated in the new species, with the dactylus longer than propodus, bearing 5 (second) to 7 (third) strong corneous spines on ventral margin. The sternite of third pereiopods anterior lobe is roundly rectangular in 
<taxonomicName id="75A84D50FFF1627FCE14550DFAF1FDC7" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[1121,1319,547,573]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF1627FCE14550DFAF1FDC7" box="[1121,1319,547,573]" italics="true" pageId="10" pageNumber="436">P. pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF1627FCF58550DFABAFDC7" bold="true" box="[1325,1388,547,573]" pageId="10" pageNumber="436">
<taxonomicNameLabel id="9BEF57BAFFF1627FCF58550DFABAFDC7" box="[1325,1388,547,573]" pageId="10" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="75A84D50FFF1627FCAE25569FEF8FD9A" authorityName="McLaughlin &amp; Forest" authorityYear="1999" box="[151,302,583,608]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="emmersoni">
<emphasis id="80DCEAC1FFF1627FCAE25569FEF8FD9A" box="[151,302,583,608]" italics="true" pageId="10" pageNumber="436">P. emmersoni</emphasis>
</taxonomicName>
, although with differences in setation, while in 
<taxonomicName id="75A84D50FFF1627FC9315569FC6FFD9B" baseAuthorityName="Barnard" baseAuthorityYear="1947" box="[836,953,583,609]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="liochele">
<emphasis id="80DCEAC1FFF1627FC9315569FC6FFD9B" box="[836,953,583,609]" italics="true" pageId="10" pageNumber="436">P. liochele</emphasis>
</taxonomicName>
the shape of the sternite is subovate.
</paragraph>
<caption id="E6D7665BFFFC6272CAE2507CFA4BF82D" ID-DOI="http://doi.org/10.5281/zenodo.7333832" ID-Zenodo-Dep="7333832" httpUri="https://zenodo.org/record/7333832/files/figure.png" pageId="7" pageNumber="433" startId="7.[151,250,1874,1898]" targetBox="[193,1395,193,1836]" targetPageId="7">
<paragraph id="B21736D3FFFC6272CAE2507CFA4BF82D" blockId="7.[151,1437,1874,2007]" pageId="7" pageNumber="433">
<emphasis id="80DCEAC1FFFC6272CAE2507CFEDBF890" bold="true" box="[151,269,1874,1898]" pageId="7" pageNumber="433">FIGURE 2</emphasis>
. 
<taxonomicName id="75A84D50FFFC6272CB6F507DFDD9F890" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[282,527,1874,1898]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="433" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFFC6272CB6F507DFDD9F890" box="[282,527,1874,1898]" italics="true" pageId="7" pageNumber="433">Pagurus pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFFC6272C863507DFD86F891" bold="true" box="[534,592,1875,1899]" pageId="7" pageNumber="433">
<taxonomicNameLabel id="9BEF57BAFFFC6272C863507DFD86F891" box="[534,592,1875,1899]" pageId="7" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Anterior part of carapace and cephalic appendages, dorsal view (arrow highlighting the position of the distinctive oblique calcified scales); (B) branchial number and disposition on body appendages; (C) right antennule, lateral view; (D) carapace, dorsal setae and blister-like tubercles general disposition; (E) left maxilliped 3; (F) sternite of somite XII (third pereiopods), ventral view. (Scales: A–F = 1 mm).
</paragraph>
</caption>
<caption id="E6D7665BFFF3627DCAE25042FB69F80A" ID-DOI="http://doi.org/10.5281/zenodo.7333834" ID-Zenodo-Dep="7333834" httpUri="https://zenodo.org/record/7333834/files/figure.png" pageId="8" pageNumber="434" startId="8.[151,250,1900,1924]" targetBox="[205,1391,191,1870]" targetPageId="8">
<paragraph id="B21736D3FFF3627DCAE25042FB69F80A" blockId="8.[151,1436,1900,2032]" pageId="8" pageNumber="434">
<emphasis id="80DCEAC1FFF3627DCAE25042FED9F87E" bold="true" box="[151,271,1900,1924]" pageId="8" pageNumber="434">FIGURE 3</emphasis>
. 
<taxonomicName id="75A84D50FFF3627DCB6B5043FDC3F87E" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[286,533,1900,1924]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="434" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF3627DCB6B5043FDC3F87E" box="[286,533,1900,1924]" italics="true" pageId="8" pageNumber="434">Pagurus pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF3627DC86B5042FD8FF87E" bold="true" box="[542,601,1900,1924]" pageId="8" pageNumber="434">
<taxonomicNameLabel id="9BEF57BAFFF3627DC86B5042FD8FF87E" box="[542,601,1900,1924]" pageId="8" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) Right cheliped, dorsal view; (B) right cheliped, mesial view; (C) anterior part of the right cheliped, lateral view; (D) left cheliped, dorsal view; anterior part of left cheliped, lateral view (inset: detail of the dactylus masticatory border); (E) anterior part of the left cheliped, lateral view (F) detail of circular clumps of setae. (Scales: A, B = 3 mm; C = 1 mm; D, E = 2 mm; F = 0.5 mm).
</paragraph>
</caption>
<caption id="E6D7665BFFF2627CCAE25064FDA2F808" ID-DOI="http://doi.org/10.5281/zenodo.7333836" ID-Zenodo-Dep="7333836" httpUri="https://zenodo.org/record/7333836/files/figure.png" pageId="9" pageNumber="435" startId="9.[151,250,1866,1890]" targetBox="[222,1380,187,1836]" targetPageId="9">
<paragraph id="B21736D3FFF2627CCAE25064FDA2F808" blockId="9.[151,1437,1866,2034]" pageId="9" pageNumber="435">
<emphasis id="80DCEAC1FFF2627CCAE25064FEDBF898" bold="true" box="[151,269,1866,1890]" pageId="9" pageNumber="435">FIGURE 4</emphasis>
. 
<taxonomicName id="75A84D50FFF2627CCB6F5065FDC6F898" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[282,528,1866,1890]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="435" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF2627CCB6F5065FDC6F898" box="[282,528,1866,1890]" italics="true" pageId="9" pageNumber="435">Pagurus pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF2627CC8625064FD86F898" bold="true" box="[535,592,1866,1890]" pageId="9" pageNumber="435">
<taxonomicNameLabel id="9BEF57BAFFF2627CC8625064FD86F898" box="[535,592,1866,1890]" pageId="9" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Mauritania, ♂ sl. 3.2 mm, holotype, (IEOCD/2979): (A) right pereiopod 2, lateral view (inset- dactylus mesial view); (B) right pereiopod 3, lateral view (inset- dactylus mesial view); (C) Sternite of somite XIV and coxa of fifth male pereiopods, ventral view. 
<taxonomicName id="75A84D50FFF2627CC8A350BDFC19F850" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[726,975,1938,1962]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="435" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF2627CC8A350BDFC19F850" box="[726,975,1938,1962]" italics="true" pageId="9" pageNumber="435">Pagurus pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF2627CC9AC50BCFBC0F850" bold="true" box="[985,1046,1938,1962]" pageId="9" pageNumber="435">
<taxonomicNameLabel id="9BEF57BAFFF2627CC9AC50BCFBC0F850" box="[985,1046,1938,1962]" pageId="9" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Mauritania, ♀ sl. 3.9 mm, paratype, (IEOCD/2978): (D) Sternite of somite XII and coxa of third female pereiopods, ventral view; (E) left pereiopod 4, lateral view; (F) 
<taxonomicName id="75A84D50FFF2627CCAC950F4FED4F808" authorityName="Pearse" authorityYear="1952" box="[188,258,2010,2034]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="432" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
, dorsal view. (Scales: A–F = 1 mm).
</paragraph>
</caption>
<paragraph id="B21736D3FFF1627FCAB25545FD74FAAF" blockId="10.[151,1437,151,1365]" pageId="10" pageNumber="436">
The other three species share also some morphological characters, the last two even with a distribution relatively close to that of the new species (see introduction). 
<taxonomicName id="75A84D50FFF1627FC8A555A1FC88FD53" authorityName="Forest" authorityYear="1966" box="[720,862,655,681]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFF1627FC8A555A1FC88FD53" box="[720,862,655,681]" italics="true" pageId="10" pageNumber="436">P. fimbriatus</emphasis>
</taxonomicName>
differs in the rostrum, which is obtuse, not pointed and not thickened. The shield is as long as broad while in the new species is broader than long. The antennal scales are shorter in 
<taxonomicName id="75A84D50FFF1627FCBF455F9FDC7FD0B" authorityName="Forest" authorityYear="1966" box="[385,529,727,753]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFF1627FCBF455F9FDC7FD0B" box="[385,529,727,753]" italics="true" pageId="10" pageNumber="436">P. fimbriatus</emphasis>
</taxonomicName>
, reaching only the base of the cornea, while in the new species reach distal end of corneas. Another important character can be found at dorsolateral distal angle of the second antennal segment, which is strongly produced in 
<taxonomicName id="75A84D50FFF1627FCB855431FD60FCC3" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[496,694,799,825]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF1627FCB855431FD60FCC3" box="[496,694,799,825]" italics="true" pageId="10" pageNumber="436">P. pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF1627FC8CB5431FD28FCC3" bold="true" box="[702,766,799,825]" pageId="10" pageNumber="436">
<taxonomicNameLabel id="9BEF57BAFFF1627FC8CB5431FD28FCC3" box="[702,766,799,825]" pageId="10" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, reaching to 3/4 of fourth peduncular segment, terminating in strong spine and with mesial margin bearing at least 10 additional small spines; based on the descriptions and figures provided by 
<bibRefCitation id="D6394B22FFF1627FCB095449FDC1FC7B" author="Forest, J." box="[380,535,871,897]" pageId="10" pageNumber="436" pagination="125 - 172" refId="ref8336" refString="Forest, J., 1966. Crustaces Decapodes: Pagurides. In: Campagne de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome ' et Annobon (1956). Annales de l'Institut Oceanographique de Monaco, 44, pp. 125 - 172." type="journal article" year="1966">Forest (1966)</bibRefCitation>
, this character is not present in 
<taxonomicName id="75A84D50FFF1627FC9F55449FBD9FC7B" authorityName="Forest" authorityYear="1966" box="[896,1039,871,897]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFF1627FC9F55449FBD9FC7B" box="[896,1039,871,897]" italics="true" pageId="10" pageNumber="436">P. fimbriatus</emphasis>
</taxonomicName>
. 
<taxonomicName id="75A84D50FFF1627FCE6B5449FADDFC7B" baseAuthorityName="Chevreux &amp; Bouvier" baseAuthorityYear="1982" box="[1054,1291,871,897]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis id="80DCEAC1FFF1627FCE6B5449FADDFC7B" box="[1054,1291,871,897]" italics="true" pageId="10" pageNumber="436">Pagurus triangularis</emphasis>
</taxonomicName>
and the new species share a pointed rostrum, but the former (like 
<taxonomicName id="75A84D50FFF1627FC97254A5FC4FFC5F" authorityName="Forest" authorityYear="1966" box="[775,921,907,933]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="80DCEAC1FFF1627FC97254A5FC4FFC5F" box="[775,921,907,933]" italics="true" pageId="10" pageNumber="436">P. fimbriatus</emphasis>
</taxonomicName>
) has a narrower shield and shorter antennal acicles, which does not reach the distal margin of the cornea (at most it reaches its base), and also has a shorter dorsolateral distal process at the second antennal segment. 
<taxonomicName id="75A84D50FFF1627FC95F54FDFC1AFC17" baseAuthorityName="Balss" baseAuthorityYear="1921" box="[810,972,979,1005]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="similimanus">
<emphasis id="80DCEAC1FFF1627FC95F54FDFC1AFC17" box="[810,972,979,1005]" italics="true" pageId="10" pageNumber="436">P. similimanus</emphasis>
</taxonomicName>
also has a narrower shield, and a rostrum that is less projected than in the new species. The ocular peduncles are also shorter in 
<taxonomicName id="75A84D50FFF1627FCE1E54D9FAD8FBEB" baseAuthorityName="Balss" baseAuthorityYear="1921" box="[1131,1294,1015,1041]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="similimanus">
<emphasis id="80DCEAC1FFF1627FCE1E54D9FAD8FBEB" box="[1131,1294,1015,1041]" italics="true" pageId="10" pageNumber="436">P. similimanus</emphasis>
</taxonomicName>
, reaching to midlength of antennular segment 3, and proximal 1/3 of antennal segment 5, while in the new species reach the distal 1/3 of the antennular segment 3 and antennal segment 5. The antennal acicle is much longer, clearly exceeding the cornea in 
<taxonomicName id="75A84D50FFF1627FCB4D534DFE0BFB87" baseAuthorityName="Balss" baseAuthorityYear="1921" box="[312,477,1123,1149]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="similimanus">
<emphasis id="80DCEAC1FFF1627FCB4D534DFE0BFB87" box="[312,477,1123,1149]" italics="true" pageId="10" pageNumber="436">P. similimanus</emphasis>
</taxonomicName>
while in the new species reaches the distal margin of the cornea. The dorsolateral distal process of the second antennal segment has 6–8 little teeth on mesial margin, while in the new species the number is higher (up to 10). The shape of the chelipeds is also different in all these species, with a variable degree of elongation that never reaches that of the new species. The dorsal surface of the chelipeds is flat and unarmed in 
<taxonomicName id="75A84D50FFF1627FCAE253DDFE8AFAF7" authority="Almón &amp; Cuesta &amp; García-Raso, 2022" authorityName="Almón &amp; Cuesta &amp; García-Raso" authorityYear="2022" box="[151,348,1267,1293]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="436" phylum="Arthropoda" rank="species" species="pectinidactylus" status="sp. nov.">
<emphasis id="80DCEAC1FFF1627FCAE253DDFE8AFAF7" box="[151,348,1267,1293]" italics="true" pageId="10" pageNumber="436">P. pectinidactylus</emphasis>
</taxonomicName>
<emphasis id="80DCEAC1FFF1627FCB1653DDFE77FAF7" bold="true" box="[355,417,1267,1293]" pageId="10" pageNumber="436">
<taxonomicNameLabel id="9BEF57BAFFF1627FCB1653DDFE77FAF7" box="[355,417,1267,1293]" pageId="10" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and the characteristic pattern formed by the clumps of setae is not mentioned in any of the other Eastern Atlantic species, neither the diagnostic masticatory border of the dactyl of the left cheliped formed by strong, densely arranged setae instead of teeth.
</paragraph>
</subSubSection>
</treatment>
</document>