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<document id="28E7BA45859A0D99BF736B47C029A447" ID-DOI="10.11646/zootaxa.3995.1.21" ID-GBIF-Dataset="810c1097-eff4-4262-acd6-1f76d05f1f83" ID-ISSN="1175-5326" ID-Zenodo-Dep="236857" ID-ZooBank="1D5CD052-9754-493C-836D-6A69756747D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461278201543" checkinUser="plazi" docAuthor="Oliver, P. Graham" docDate="2015" docId="3956C47FFFE2E320FF44FE48FF64FE07" docLanguage="en" docName="zt03995p263.pdf" docOrigin="Zootaxa 3995 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parathyasira Iredale 1930" docType="treatment" docVersion="8" lastPageNumber="255" masterDocId="C56FBC07FFE3E323FFD3FFE5FF80FFD7" masterDocTitle="Deep-water Thyasiridae (Mollusca: Bivalvia) from the Oman Margin, Arabian Sea, new species and examples of endemism and cosmopolitanism" masterLastPageNumber="263" masterPageNumber="252" pageNumber="253" updateTime="1698613566852" updateUser="plazi">
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<mods:title id="A3D13AAFD2ACEC5DF3D2647594A75F9C">Deep-water Thyasiridae (Mollusca: Bivalvia) from the Oman Margin, Arabian Sea, new species and examples of endemism and cosmopolitanism</mods:title>
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<mods:namePart id="32622CC68177958E726A3E4346EDF239">Oliver, P. Graham</mods:namePart>
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<treatment id="3956C47FFFE2E320FF44FE48FF64FE07" ID-DOI="http://doi.org/10.5281/zenodo.6098571" ID-GBIF-Taxon="119640208" ID-Zenodo-Dep="6098571" LSID="urn:lsid:plazi:treatment:3956C47FFFE2E320FF44FE48FF64FE07" httpUri="http://treatment.plazi.org/id/3956C47FFFE2E320FF44FE48FF64FE07" lastPageId="3" lastPageNumber="255" pageId="1" pageNumber="253">
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<paragraph id="B1407569FFE2E322FF44FE48FE5DFE10" blockId="1.[151,477,429,455]" box="[151,477,429,455]" pageId="1" pageNumber="253">
<heading id="EA08C205FFE2E322FF44FE48FE5DFE10" bold="true" box="[151,477,429,455]" fontSize="11" level="1" pageId="1" pageNumber="253" reason="1">
<taxonomicName id="76FF0EEAFFE2E322FF44FE48FE5DFE10" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" box="[151,477,429,455]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="genus">
<emphasis id="838BA97BFFE2E322FF44FE48FE5DFE10" bold="true" box="[151,477,429,455]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF44FE48FEB4FE10" bold="true" box="[151,308,429,455]" italics="true" pageId="1" pageNumber="253">Parathyasira</emphasis>
<bibRefCitation id="D56E0898FFE2E322FEE9FE48FE5DFE10" author="Iredale" box="[314,477,429,455]" pageId="1" pageNumber="253" refString="Iredale, T. (1930) More notes on the marine Mollusca of New South Wales. Records of the Australian Museum, 17, 384 - 407. http: // dx. doi. org / 10.3853 / J. 0067 - 1975.17.1930.773" type="journal article" year="1930">Iredale, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="B1407569FFE2E322FF44FE11FD4FFDDB" blockId="1.[151,1437,500,885]" box="[151,719,500,525]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF44FE11FEABFDDA" bold="true" box="[151,299,500,525]" pageId="1" pageNumber="253">
<typeStatus id="6E44CBCBFFE2E322FF44FE11FF52FDDA" box="[151,210,500,525]" pageId="1" pageNumber="253">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="76FF0EEAFFE2E322FEEAFE11FD4FFDDB" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" box="[313,719,500,525]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="resupina">
<emphasis id="838BA97BFFE2E322FEEAFE11FDB5FDDB" box="[313,565,500,525]" italics="true" pageId="1" pageNumber="253">Parathyasira resupina</emphasis>
<bibRefCitation id="D56E0898FFE2E322FDEFFE11FD4FFDDB" author="Iredale" box="[572,719,500,525]" pageId="1" pageNumber="253" refString="Iredale, T. (1930) More notes on the marine Mollusca of New South Wales. Records of the Australian Museum, 17, 384 - 407. http: // dx. doi. org / 10.3853 / J. 0067 - 1975.17.1930.773" type="journal article" year="1930">Iredale, 1930</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFE2E322FF14FDF2FA12FDAF" pageId="1" pageNumber="253" type="diagnosis">
<paragraph id="B1407569FFE2E322FF14FDF2FA12FDAF" blockId="1.[151,1437,500,885]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14FDF2FEBAFDE7" bold="true" box="[199,314,535,560]" pageId="1" pageNumber="253">Diagnosis</emphasis>
. Small to medium sized thyasirids with a weakly defined posterior area, a well-defined supramarginal sulcus without an auricle, a microsculpture of calcareous spines arranged in dense radial rows. Hinge lacking teeth, ligament weak, in a shallow resilifer. Gills of two demibranchs, delicate,
<typeStatus id="6E44CBCBFFE2E322FBBBFD85FB21FDAF" box="[1128,1185,608,632]" pageId="1" pageNumber="253">Type</typeStatus>
2aR of Oliver (2014).
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFE2E322FF14FD61FE90FC37" pageId="1" pageNumber="253" type="discussion">
<paragraph id="B1407569FFE2E322FF14FD61FBCDFCA2" blockId="1.[151,1437,500,885]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14FD61FEBBFD4A" bold="true" box="[199,315,644,669]" pageId="1" pageNumber="253">Remarks.</emphasis>
The diagnosis here is limited to those species with a spiny micro-sculpture arranged in radial rows. There are many other species with a similar overall shell form such as the North Atlantic
<taxonomicName id="76FF0EEAFFE2E322FBA4FD42FE99FD33" authority="Verrill &amp; Bush, 1898" authorityName="Verrill &amp; Bush" authorityYear="1898" class="Bivalvia" family="Thyasiridae" genus="Thyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="equalis">
<emphasis id="838BA97BFFE2E322FBA4FD42FAB2FD17" box="[1143,1330,679,704]" italics="true" pageId="1" pageNumber="253">Thyasira equalis</emphasis>
<bibRefCitation id="D56E0898FFE2E322FAEBFD42FE99FD33" author="Verrill" pageId="1" pageNumber="253" refString="Verrill, A. E. &amp; Bush, K. J. (1898) Revision of the deep-water Mollusca of the Atlantic coast of North America, with descriptions of new genera and species. Proceedings of the United States National Museum, 20, 775 - 901. http: // dx. doi. org / 10.5479 / si. 00963801.20 - 1139.775" type="journal article" year="1898">Verrill &amp; Bush, 1898</bibRefCitation>
</taxonomicName>
that have been placed in
<taxonomicName id="76FF0EEAFFE2E322FD9BFD29FD5CFD32" box="[584,732,716,741]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="genus">
<emphasis id="838BA97BFFE2E322FD9BFD29FD5CFD32" box="[584,732,716,741]" italics="true" pageId="1" pageNumber="253">Parathyasira</emphasis>
</taxonomicName>
by previous studies (
<bibRefCitation id="D56E0898FFE2E322FC05FD29FB49FD33" author="Payne" box="[982,1225,716,741]" pageId="1" pageNumber="253" refString="Payne, C. M. &amp; Allen, J. A. (1991) The morphology of deep-sea Thyasiridae (Mollusca: Bivalvia) from the Atlantic Ocean. Philosophical Transactions of the Royal Society of London Series B, 334, 481 - 566. http: // dx. doi. org / 10.1098 / rstb. 1991.0128" type="journal article" year="1991">Payne &amp; Allen 1991</bibRefCitation>
;
<bibRefCitation id="D56E0898FFE2E322FB0BFD29FF53FCDF" author="Oliver" pageId="1" pageNumber="253" refString="Oliver, P. G. &amp; Killeen, I. J. (2002) The Thyasiridae of the British Continental Shelf and North Sea Oilfields. Studies in Marine Biodiversity and Systematics from the National Museum of Wales. Biomor Reports, 3, 73 pp." type="book" year="2002">Oliver &amp; Killeen 2002</bibRefCitation>
;
<bibRefCitation id="D56E0898FFE2E322FF0EFD0AFEEEFCDF" author="Zelaya" box="[221,366,751,776]" pageId="1" pageNumber="253" refString="Zelaya, D. G. (2009) The genera Thyasira and Parathyasira in the Magellan Region and adjacent Antarctic waters (Bivalvia: Thyasiridae). Malacologia, 51 (2), 271 - 290. http: // dx. doi. org / 10.4002 / 040.051.0204" type="journal article" year="2009">Zelaya 2009</bibRefCitation>
). There remains many uncertainties concerning the relationships of these species in conjunction with many others currently placed in
<taxonomicName id="76FF0EEAFFE2E322FD93FCF1FD24FCFA" box="[576,676,788,813]" class="Bivalvia" family="Thyasiridae" genus="Thyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="genus">
<emphasis id="838BA97BFFE2E322FD93FCF1FD24FCFA" box="[576,676,788,813]" italics="true" pageId="1" pageNumber="253">Thyasira</emphasis>
</taxonomicName>
but not agreeing in shell form or anatomy with the
<typeStatus id="6E44CBCBFFE2E322FB20FCF0FAA3FCFA" box="[1267,1315,789,813]" pageId="1" pageNumber="253">type</typeStatus>
species of
<taxonomicName id="76FF0EEAFFE2E322FF44FCD2FF7BFC87" box="[151,251,823,848]" class="Bivalvia" family="Thyasiridae" genus="Thyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="genus">
<emphasis id="838BA97BFFE2E322FF44FCD2FF7BFC87" box="[151,251,823,848]" italics="true" pageId="1" pageNumber="253">Thyasira</emphasis>
</taxonomicName>
,
<taxonomicName id="76FF0EEAFFE2E322FEDDFCDCFDE1FC87" authority="Montagu, 1803" authorityName="Montagu" authorityYear="1803" box="[270,609,823,848]" class="Bivalvia" family="Thyasiridae" genus="Thyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="flexuosa">
<emphasis id="838BA97BFFE2E322FEDDFCDCFE0DFC87" box="[270,397,823,848]" italics="true" pageId="1" pageNumber="253">T. flexuosa</emphasis>
(
<bibRefCitation id="D56E0898FFE2E322FE71FCDDFDD9FC87" author="Montagu" box="[418,601,824,848]" pageId="1" pageNumber="253" refString="Montagu, G. (1803) Testacea Britannica, or natural history of British shells, marine, land and the fresh-water, including the most minute: systematically arranged and embellished with figures, Romsey, London, 606 pp., 16 pls." type="book" year="1803">Montagu, 1803</bibRefCitation>
)
</taxonomicName>
. Further research is required to resolve how many genera should be recognized, while those with a distinct radial micro-sculpture form a coherent group.
</paragraph>
<paragraph id="B1407569FFE2E322FF44FC41FE90FC37" blockId="1.[151,492,932,992]" pageId="1" pageNumber="253">
<taxonomicName id="76FF0EEAFFE2E322FF44FC41FE0FFC6A" box="[151,399,932,957]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="bamberi" status="sp. nov.">
<emphasis id="838BA97BFFE2E322FF44FC41FE0FFC6A" box="[151,399,932,957]" italics="true" pageId="1" pageNumber="253">Parathyasira bamberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98B81400FFE2E322FE45FC40FE6CFC6B" box="[406,492,933,956]" pageId="1" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
<figureCitation id="29C469ECFFE2E322FF44FC22FE90FC37" box="[151,272,967,992]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="1" pageNumber="253">Figs 1AE</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="B1407569FFE2E322FF44FBEAFB51FBA7" blockId="1.[151,1437,1039,1967]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF44FBEAFE3CFBFF" bold="true" box="[151,444,1039,1064]" pageId="1" pageNumber="253">
<typeStatus id="6E44CBCBFFE2E322FF44FBEAFF52FBFF" box="[151,210,1039,1064]" pageId="1" pageNumber="253">Type</typeStatus>
material.
<typeStatus id="6E44CBCBFFE2E322FE9CFBEAFE3CFBFF" box="[335,444,1039,1064]" pageId="1" pageNumber="253" type="holotype">Holotype</typeStatus>
</emphasis>
- single live collected specimen, Off Ras Madrakah, southern
<collectingCountry id="C9E835F9FFE2E322FB5AFBF5FB50FBFF" box="[1161,1232,1040,1064]" name="Oman" pageId="1" pageNumber="253">Oman</collectingCountry>
, Discovery cruise 211 st. 12687#2,
<geoCoordinate id="D4CB13AEFFE2E322FE89FBD1FE65FB9B" box="[346,485,1076,1101]" direction="north" orientation="latitude" pageId="1" pageNumber="253" precision="9" value="18.930834">18°55.85´N</geoCoordinate>
<geoCoordinate id="D4CB13AEFFE2E322FE3EFBD1FDF5FB9B" box="[493,629,1076,1101]" direction="east" orientation="longitude" pageId="1" pageNumber="253" precision="9" value="58.8695">58°52.17´E</geoCoordinate>
to
<geoCoordinate id="D4CB13AEFFE2E322FD48FBD1FCA6FB9B" box="[667,806,1076,1101]" direction="north" orientation="latitude" pageId="1" pageNumber="253" precision="9" value="18.869333">18°52.16´N</geoCoordinate>
<geoCoordinate id="D4CB13AEFFE2E322FCFDFBD1FC39FB9B" box="[814,953,1076,1101]" direction="east" orientation="longitude" pageId="1" pageNumber="253" precision="9" value="59.120667">59°07.24´E</geoCoordinate>
,
<quantity id="7607D88CFFE2E322FC16FBD1FB9DFB9B" box="[965,1053,1076,1101]" metricMagnitude="3" metricUnit="m" metricValue="3.356" pageId="1" pageNumber="253" unit="m" value="3356.0">3356 m</quantity>
, collected P.G. Oliver,
<date id="C54153A9FFE2E322FACDFBD1FF53FBA7" pageId="1" pageNumber="253" value="1994-10-19">19 October 1994</date>
.
<collectionCode id="D7EEEDACFFE2E322FF0EFBBDFEA9FBB8" box="[221,297,1112,1135]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="253">NMW</collectionCode>
.Z. 1995.009.13.
<typeStatus id="6E44CBCBFFE2E322FE31FBBDFDC7FBA7" box="[482,583,1112,1136]" pageId="1" pageNumber="253" type="paratype">Paratype</typeStatus>
, one fragmented shell as
<typeStatus id="6E44CBCBFFE2E322FCB1FBB2FC47FBA7" box="[866,967,1111,1136]" pageId="1" pageNumber="253" type="holotype">holotype</typeStatus>
,
<collectionCode id="D7EEEDACFFE2E322FC01FBBDFB9EFBB8" box="[978,1054,1112,1135]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="253">NMW</collectionCode>
.Z. 1995.009.14.
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFE2E322FF14FB99FC52F8B0" pageId="1" pageNumber="253" type="description">
<paragraph id="B1407569FFE2E322FF14FB99FDF1FA47" blockId="1.[151,1437,1039,1967]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14FB99FED9FB42" bold="true" box="[199,345,1148,1173]" pageId="1" pageNumber="253">Description.</emphasis>
<emphasis id="838BA97BFFE2E322FEB1FB99FE1AFB42" box="[354,410,1148,1173]" italics="true" pageId="1" pageNumber="253">Shell</emphasis>
(
<figureCitation id="29C469ECFFE2E322FE78FB99FDB0FB43" box="[427,560,1148,1173]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="1" pageNumber="253">Figs 1AD</figureCitation>
).
<typeStatus id="6E44CBCBFFE2E322FD96FB99FD2DFB42" box="[581,685,1148,1173]" pageId="1" pageNumber="253" type="holotype">Holotype</typeStatus>
<quantity id="7607D88CFFE2E322FD64FB99FC9EFB43" box="[695,798,1148,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="1" pageNumber="253" unit="mm" value="18.7">18.7 mm</quantity>
in length,
<quantity id="7607D88CFFE2E322FC4DFB99FB84FB43" box="[926,1028,1148,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="1" pageNumber="253" unit="mm" value="18.4">18.4 mm</quantity>
in height. Thin, fragile. Equivalve. Compressed. ~Equilateral, beaks prosogyrous, just anterior of midline. Outline subcircular, slightly longer than high. Posterior dorsal margin long, almost straight, sloping steeply; anterior dorsal margin short, slightly concave; anterior and ventral margins a continuous curve. Posterior area flattened not sulcate, demarcated by a weak radial ridge. Sub-marginal sulcus (sms = escutcheon) long, narrow, sunken, edges vertical. Ligament slender not sunken but not set on an auricle, ~half the length of sms. Lunule a flattened area, not depressed. Hinge edentulous. Sculpture of commarginal growth lines and weak ridges with a micro-sculpture of dense radial rows of blunt calcareous spines. Muscle scars indistinct.
</paragraph>
<paragraph id="B1407569FFE2E322FF14FA79FEEBF967" blockId="1.[151,1437,1039,1967]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14FA79FEABFA64" box="[199,299,1436,1459]" italics="true" pageId="1" pageNumber="253">Anatomy</emphasis>
(
<figureCitation id="29C469ECFFE2E322FEE9FA79FE0FFA63" box="[314,399,1436,1461]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="1" pageNumber="253">Fig. 1E</figureCitation>
). Mantle thin; edges un-fused except for the formation of a small exhalant aperture Anterior adductor scar slender, elongate and approximately two times longer than posterior adductor scar; free from mantle edge for about one-half of its ventral edge; posterior adductor muscle oval. Foot vermiform very long, tip noticeably expanded, heel obsolete, protractor muscles very slender. Labial palps small, with distinctly grooved lamellae leading to a short tubular portion leading to the mouth. Lateral body pouches large, thin walled, branches few, terminations simple, rounded. Kidney large with golden coloured granules. Ctenidium large, both demibranchs with fully reflected filaments; outer demibranch about half the depth of the inner demibranch. Filaments delicate.
</paragraph>
<paragraph id="B1407569FFE2E322FF14F959FC52F8B0" blockId="1.[151,1437,1039,1967]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14F959FE3CF902" bold="true" box="[199,444,1724,1749]" pageId="1" pageNumber="253">Habitat and ecology.</emphasis>
Taken from abyssal depths in ooze along with a high diversity of protobranchs and large numbers of suspension feeding
<taxonomicName id="76FF0EEAFFE2E322FDD7F904FCA1F92F" authority="Jeffreys" authorityName="Jeffreys" box="[516,801,1759,1784]" class="Bivalvia" family="Limopsidae" genus="Limopsis" kingdom="Animalia" order="Arcoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="tenella">
<emphasis id="838BA97BFFE2E322FDD7F904FD3FF92F" box="[516,703,1759,1784]" italics="true" pageId="1" pageNumber="253">Limopsis tenella</emphasis>
Jeffreys
</taxonomicName>
and
<taxonomicName id="76FF0EEAFFE2E322FC8FF93AFB0BF92F" authority="Dall" authorityName="Dall" box="[860,1163,1759,1784]" class="Bivalvia" family="Arcidae" genus="Bentharca" kingdom="Animalia" order="Arcoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="asperula">
<emphasis id="838BA97BFFE2E322FC8FF93AFBBCF92F" box="[860,1084,1759,1784]" italics="true" pageId="1" pageNumber="253">Bentharca asperula</emphasis>
(Dall)
</taxonomicName>
. The water temperature was 1.7°C and the oxygen concentration was 2.99 ml l -1. The delicate gill structure indicates that the abfrontal extension is poor and that any bacterial symbiosis is limited and this would be in keeping with the observations made on
<taxonomicName id="76FF0EEAFFE2E322FF2EF8ABFD7EF8B0" authority="Monterosato, 1874" authorityName="Monterosato" authorityYear="1874" box="[253,766,1870,1895]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="1" pageNumber="253" phylum="Mollusca" rank="species" species="granulosa">
<emphasis id="838BA97BFFE2E322FF2EF8ABFD89F8B0" box="[253,521,1870,1895]" italics="true" pageId="1" pageNumber="253">Parathyasira granulosa</emphasis>
, (
<bibRefCitation id="D56E0898FFE2E322FDCCF8AAFD76F8B0" author="Monterosato" box="[543,758,1871,1895]" pageId="1" pageNumber="253" refString="Monterosato, T. A. (1874) Recherches conchyliologiques, effectuees au Cap Santo Vito, en Sicile. (Traduz. dall'italiano di H. Crosse). Journal de Conchyliologie, 22 (3), 243 - 282." type="journal article" year="1874">Monterosato, 1874</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="D56E0898FFE2E322FCFBF8ABFC4DF8B0" author="Dufour" box="[808,973,1870,1895]" pageId="1" pageNumber="253" refString="Dufour, S. C. (2005) Gill anatomy and evolution of symbiosis in the bivalve family Thyasiridae. Biological Bulletin, 208, 200 - 212. http: // dx. doi. org / 10.2307 / 3593152" type="journal article" year="2005">Dufour (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F9E526E2FFE2E320FF14F897FF64FE07" lastPageId="3" lastPageNumber="255" pageId="1" pageNumber="253" type="etymology">
<paragraph id="B1407569FFE2E322FF14F897FCC6F878" blockId="1.[151,1437,1039,1967]" pageId="1" pageNumber="253">
<emphasis id="838BA97BFFE2E322FF14F897FECEF85C" bold="true" box="[199,334,1906,1931]" pageId="1" pageNumber="253">Etymology.</emphasis>
Named for Roger Bamber, fellow student from our days at the Dove Marine Laboratory, Cullercoats, Northumberland, sadly departed February, 2015.
</paragraph>
<caption id="E58025E1FFE1E321FF44F8F9FF5DF826" httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="2" pageNumber="254" targetBox="[188,1409,181,1796]" targetPageId="2">
<paragraph id="B1407569FFE1E321FF44F8F9FF5DF826" blockId="2.[151,1436,1820,2033]" pageId="2" pageNumber="254">
<emphasis id="838BA97BFFE1E321FF44F8F9FE96F8E6" bold="true" box="[151,278,1820,1842]" pageId="2" pageNumber="254">FIGURE 1.</emphasis>
AE,
<taxonomicName id="76FF0EEAFFE1E321FE8DF8F9FDC1F8E5" box="[350,577,1820,1842]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="2" pageNumber="254" phylum="Mollusca" rank="species" species="bamberi" status="sp. nov.">
<emphasis id="838BA97BFFE1E321FE8DF8F9FDC1F8E5" box="[350,577,1820,1842]" italics="true" pageId="2" pageNumber="254">Parathyasira bamberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98B81400FFE1E321FD9FF8F9FD1FF8E6" box="[588,671,1820,1841]" pageId="2" pageNumber="254" rank="species">
<emphasis id="838BA97BFFE1E321FD9FF8F9FD1FF8E6" bold="true" box="[588,671,1820,1841]" pageId="2" pageNumber="254">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype: AB, external and internal of the right valve; CD, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F,
<taxonomicName id="76FF0EEAFFE1E321FF44F8BCFE49F8B8" authority="Iredale" authorityName="Iredale" box="[151,457,1881,1903]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="2" pageNumber="254" phylum="Mollusca" rank="species" species="resupina">
<emphasis id="838BA97BFFE1E321FF44F8BCFEFBF8B8" box="[151,379,1881,1903]" italics="true" pageId="2" pageNumber="254">Parathyasira resupina</emphasis>
Iredale
</taxonomicName>
, holotype, Australian Museum, C57834; G,
<taxonomicName id="76FF0EEAFFE1E321FC55F8BCFAB9F8B8" authority="Monterosato, Gulf" authorityName="Monterosato, Gulf" box="[902,1337,1881,1903]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="2" pageNumber="254" phylum="Mollusca" rank="species" species="granulosa">
<emphasis id="838BA97BFFE1E321FC55F8BCFBF9F8B8" box="[902,1145,1881,1903]" italics="true" pageId="2" pageNumber="254">Parathyasira granulosa</emphasis>
Monterosato, Gulf
</taxonomicName>
of Cadiz, National Museum Wales; H,
<taxonomicName id="76FF0EEAFFE1E321FE6EF89DFBD5F859" authority="Payne &amp; Allen, Rockall Trough, 1595" authorityName="Payne &amp; Allen, Rockall Trough" authorityYear="1595" box="[445,1109,1912,1934]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="2" pageNumber="254" phylum="Mollusca" rank="species" species="subcircularis">
<emphasis id="838BA97BFFE1E321FE6EF89DFD4EF859" box="[445,718,1912,1934]" italics="true" pageId="2" pageNumber="254">Parathyasira subcircularis</emphasis>
Payne &amp; Allen, Rockall Trough, 1595
</taxonomicName>
m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus.
</paragraph>
</caption>
<paragraph id="B1407569FFE0E320FF14FF72FF64FE07" blockId="3.[151,1437,151,464]" pageId="3" pageNumber="255">
<emphasis id="838BA97BFFE0E320FF14FF72FEECFF67" bold="true" box="[199,364,151,176]" pageId="3" pageNumber="255">Comparisons.</emphasis>
The number of species of
<taxonomicName id="76FF0EEAFFE0E320FD49FF72FC9EFF67" box="[666,798,151,176]" class="Bivalvia" family="Thyasiridae" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="family">Thyasiridae</taxonomicName>
with a radial microsculpture of calcareous spines is few. The
<typeStatus id="6E44CBCBFFE0E320FF19FF58FF7AFF02" box="[202,250,189,213]" pageId="3" pageNumber="255">type</typeStatus>
species
<taxonomicName id="76FF0EEAFFE0E320FE89FF58FE57FF03" box="[346,471,189,212]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="resupina">
<emphasis id="838BA97BFFE0E320FE89FF58FE57FF03" box="[346,471,189,212]" italics="true" pageId="3" pageNumber="255">P. resupina</emphasis>
</taxonomicName>
(
<figureCitation id="29C469ECFFE0E320FE34FF59FDA6FF03" box="[487,550,188,213]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="3" pageNumber="255">Fig. 1</figureCitation>
F) and the closely related
<taxonomicName id="76FF0EEAFFE0E320FC9CFF58FB0DFF03" authority="Iredale, 1930" authorityName="Iredale" authorityYear="1930" box="[847,1165,188,213]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="neozelanica">
<emphasis id="838BA97BFFE0E320FC9CFF58FC72FF02" box="[847,1010,188,213]" italics="true" pageId="3" pageNumber="255">P. neozelanica</emphasis>
<bibRefCitation id="D56E0898FFE0E320FC2AFF59FB0DFF03" author="Iredale" box="[1017,1165,188,213]" pageId="3" pageNumber="255" refString="Iredale, T. (1930) More notes on the marine Mollusca of New South Wales. Records of the Australian Museum, 17, 384 - 407. http: // dx. doi. org / 10.3853 / J. 0067 - 1975.17.1930.773" type="journal article" year="1930">Iredale, 1930</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="76FF0EEAFFE0E320FB16FF58FEE7FF2F" authority="Cotton &amp; Godfrey, 1938" authorityName="Cotton &amp; Godfrey" authorityYear="1938" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="verconis">
<emphasis id="838BA97BFFE0E320FB16FF58FABFFF03" box="[1221,1343,189,212]" italics="true" pageId="3" pageNumber="255">P. verconis</emphasis>
(
<bibRefCitation id="D56E0898FFE0E320FA9DFF59FEDFFF2F" author="Cotton" pageId="3" pageNumber="255" refString="Cotton, B. C. &amp; Godfrey, F. K. (1938) The Molluscs of South Australia. Trigg, Adelaide, 314 pp." type="book" year="1938">Cotton &amp; Godfrey, 1938</bibRefCitation>
)
</taxonomicName>
inhabit shelf depths off southern
<collectingCountry id="C9E835F9FFE0E320FD28FF3AFCE2FF2F" box="[763,866,223,248]" name="Australia" pageId="3" pageNumber="255">Australia</collectingCountry>
and
<collectingCountry id="C9E835F9FFE0E320FC72FF05FBC0FF2F" box="[929,1088,223,248]" name="New Zealand" pageId="3" pageNumber="255">New Zealand</collectingCountry>
. These are much smaller, not exceeding
<quantity id="7607D88CFFE0E320FEDCFEE1FECBFECB" box="[271,331,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="255" unit="mm" value="8.0">8mm</quantity>
, are higher than long in outline with a much stronger posterior sinus. The other known species are from the North Atlantic,
<taxonomicName id="76FF0EEAFFE0E320FE78FECCFCA4FE97" authority="Monterosato, 1874" authorityName="Monterosato" authorityYear="1874" box="[427,804,295,320]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="granulosa">
<emphasis id="838BA97BFFE0E320FE78FECCFDB7FE97" box="[427,567,295,320]" italics="true" pageId="3" pageNumber="255">P. granulosa</emphasis>
(
<bibRefCitation id="D56E0898FFE0E320FD95FECDFC9CFE97" author="Monterosato" box="[582,796,296,320]" pageId="3" pageNumber="255" refString="Monterosato, T. A. (1874) Recherches conchyliologiques, effectuees au Cap Santo Vito, en Sicile. (Traduz. dall'italiano di H. Crosse). Journal de Conchyliologie, 22 (3), 243 - 282." type="journal article" year="1874">Monterosato, 1874</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="29C469ECFFE0E320FCE1FEC2FCF0FE97" box="[818,880,295,320]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="3" pageNumber="255">Fig. 1</figureCitation>
G) is known from the lower shelf and upper slope while
<taxonomicName id="76FF0EEAFFE0E320FF37FEA8FD3EFEB3" authority="Payne &amp; Allen, 1991" authorityName="Payne &amp; Allen" authorityYear="1991" box="[228,702,332,357]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="subcircularis">
<emphasis id="838BA97BFFE0E320FF37FEA8FE1AFEB2" box="[228,410,332,357]" italics="true" pageId="3" pageNumber="255">P. subcircularis</emphasis>
(
<bibRefCitation id="D56E0898FFE0E320FE63FEA9FD36FEB3" author="Payne" box="[432,694,332,357]" pageId="3" pageNumber="255" refString="Payne, C. M. &amp; Allen, J. A. (1991) The morphology of deep-sea Thyasiridae (Mollusca: Bivalvia) from the Atlantic Ocean. Philosophical Transactions of the Royal Society of London Series B, 334, 481 - 566. http: // dx. doi. org / 10.1098 / rstb. 1991.0128" type="journal article" year="1991">Payne &amp; Allen, 1991</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="29C469ECFFE0E320FD06FEA9FC9BFEB3" box="[725,795,332,357]" captionStart="FIGURE 1. A E" captionStartId="2.[151,250,1820,1842]" captionTargetBox="[188,1409,181,1796]" captionTargetId="figure@2.[188,1409,181,1798]" captionTargetPageId="2" captionText="FIGURE 1. A E, Parathyasira bamberi sp. nov. Holotype: A B, external and internal of the right valve; C D, scanning electron micrographs of the microsculpture; E, gross anatomy viewed from the right side and right mantle removed; F, Parathyasira resupina Iredale, holotype, Australian Museum, C 57834; G, Parathyasira granulosa Monterosato, Gulf of Cadiz, National Museum Wales; H, Parathyasira subcircularis Payne &amp; Allen, Rockall Trough, 1595 m, National Museum Wales. aa, anterior adductor muscle; f, foot; hg, hind gut; id, inner demibranch; k, kidney; lbp, lateral body pouch; lig, ligament; lp, labial palps; lun, lunule; me, mantle edge; od, outer demibranch; pa, posterior adductor muscle; r, rectum; sms, submarginal sulcus." httpUri="https://zenodo.org/record/236858/files/figure.png" pageId="3" pageNumber="255">Fig. 1</figureCitation>
H) is known from the lower slope and abyssal rise.
<taxonomicName id="76FF0EEAFFE0E320FF44FE8AFE25FE5F" box="[151,421,367,392]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="granulosa">
<emphasis id="838BA97BFFE0E320FF44FE8AFE25FE5F" box="[151,421,367,392]" italics="true" pageId="3" pageNumber="255">Parathyasira granulosa</emphasis>
</taxonomicName>
is very similar to the Australian species while
<taxonomicName id="76FF0EEAFFE0E320FC6FFE94FBEBFE5F" box="[956,1131,367,392]" class="Bivalvia" family="Thyasiridae" genus="Parathyasira" kingdom="Animalia" order="Lucinoida" pageId="3" pageNumber="255" phylum="Mollusca" rank="species" species="subcircularis">
<emphasis id="838BA97BFFE0E320FC6FFE94FBEBFE5F" box="[956,1131,367,392]" italics="true" pageId="3" pageNumber="255">P. subcircularis</emphasis>
</taxonomicName>
has a more poorly defined posterior area. The latter is however higher than long in outline and has a less defined lunule and sub-marginal sulcus.
</paragraph>
</subSubSection>
</treatment>
</document>