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709 lines
90 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.881.36744" ID-GBIF-Dataset="3cef5ca2-433f-45f7-a32b-087b5f1946a4" ID-PMC="PMC6813179" ID-Pensoft-Pub="1313-2970-881-135" ID-Pensoft-UUID="16F69FEA95835AE5BF6B9C34F83E49D7" ID-PubMed="31662614" ID-ZooBank="B058CA156A3D41A19ADFD00384234D8E" ModsDocID="1313-2970-881-135" checkinTime="1571384733377" checkinUser="pensoft" docAuthor="Mvogo Ndongo, Pierre A., von Rintelen, Thomas & Cumberlidge, Neil" docDate="2019" docId="393AB7F6A16A55149098E1A1C9709C6C" docLanguage="en" docName="ZooKeys 881: 135-164" docOrigin="ZooKeys 881" docSource="http://dx.doi.org/10.3897/zookeys.881.36744" docTitle="Louisea nkongsamba Ndongo, Rintelen & Cumberlidge, 2019, sp. nov." docType="treatment" docUuid="141A1FD3-DF3B-4E84-9296-5AA5A26A3B68" docUuidSource="ZooBank" docVersion="6" id="16F69FEA95835AE5BF6B9C34F83E49D7" lastPageNumber="135" masterDocId="16F69FEA95835AE5BF6B9C34F83E49D7" masterDocTitle="Taxonomic revision of the endemic Cameroonian freshwater crab genus Louisea Cumberlidge, 1994 (Crustacea, Decapoda, Brachyura, Potamonautidae), with descriptions of two new species from Nkongsamba and Yabassi" masterLastPageNumber="164" masterPageNumber="135" pageNumber="135" updateTime="1668167896540" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the endemic Cameroonian freshwater crab genus Louisea Cumberlidge, 1994 (Crustacea, Decapoda, Brachyura, Potamonautidae), with descriptions of two new species from Nkongsamba and Yabassi</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mvogo Ndongo, Pierre A.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>von Rintelen, Thomas</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cumberlidge, Neil</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>881</mods:number>
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<mods:start>135</mods:start>
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<mods:end>164</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.881.36744</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.881.36744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-881-135</mods:identifier>
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<treatment ID-GBIF-Taxon="159754741" LSID="urn:lsid:zoobank.org:act:141A1FD3-DF3B-4E84-9296-5AA5A26A3B68" httpUri="http://treatment.plazi.org/id/393AB7F6A16A55149098E1A1C9709C6C" lastPageNumber="135" pageId="0" pageNumber="135">
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<taxonomicName LSID="393AB7F6-A16A-5514-9098-E1A1C9709C6C" class="Malacostraca" family="Potamonautidae" genus="Louisea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Louisea nkongsamba" order="Decapoda" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="nkongsamba">Louisea nkongsamba</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Largest adult male (CW 17.5 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 26930) (missing right cheliped) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 1 ', www. mapress. com / j / zt) c largest adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 5, and right cheliped and p 2, p 3) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 8 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure2" httpUri="https://binary.pensoft.net/fig/347890" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure2">Figs 2d</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) (missing left p 4) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 4 and right p 2) c subadult male, paratype (CW 13.8 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 4, and right cheliped and p 2 - p 5) d second largest adult male (CW 18.38 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 10 mm (b), 8.42 mm (c), 11.63 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure3" httpUri="https://binary.pensoft.net/fig/347891" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure3">3d</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">4a</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">5d</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">6d</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">8g, h</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Four species of Louisea endemic to southwestern Cameroon, right cheliped merus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 d', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure9" httpUri="https://binary.pensoft.net/fig/347897" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure9">9d</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">10d</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">11d</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12d</figureCitation>
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">13d</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">14d</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">15d</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="materials_examined">
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<paragraph pageId="0" pageNumber="135">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="135">
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CAMEROON. Holotype: adult male (CW 20 mm, CL 14.85 mm, CH 8.4 mm, FW 6.6 mm), Littoral Region, Mount Nlonako Ecological Reserve (locality 1) (4.91046N, 9.976332E), 1,237 m asl, 23 May 2018, coll. P.A. Mvogo Ndongo (ZMB Crust. 31618). Paratype: adult male (CW 18.38 mm, CL 13.32 mm, CH 8.13 mm, FW 6.34 mm), Littoral Region, Mount Nlonako Ecological Reserve (locality 2) (4.91343N, 9.98500E), 1,176 m asl, 23 May 2018, coll. P.A. Mvogo Ndongo (ZMB Crust. 31620). Other material examined is listed in Table
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<tableCitation captionStartId="T4" httpUri="http://table.plazi.org/id/7D42479E00FA589E99E3E82D767D8174" pageId="0" pageNumber="135" tableUuid="7D42479E00FA589E99E3E82D767D8174">4</tableCitation>
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.
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</paragraph>
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<caption ID-Table-UUID="7D42479E00FA589E99E3E82D767D8174" httpUri="http://table.plazi.org/id/7D42479E00FA589E99E3E82D767D8174" pageId="0" pageNumber="135" start="Table 4" startId="T4">
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<paragraph pageId="0" pageNumber="135">
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<emphasis bold="true" pageId="0" pageNumber="135">Table 4.</emphasis>
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Morphometric analysis and collection data of specimens (
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<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
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= 27) of
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<taxonomicName class="Malacostraca" family="Potamonautidae" genus="Louisea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Louisea nkongsamba" order="Decapoda" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="nkongsamba">
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<emphasis italics="true" pageId="0" pageNumber="135">Louisea nkongsamba</emphasis>
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</taxonomicName>
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sp. nov. from Cameroon. All measurements are given in mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="135">
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<table pageId="0" pageNumber="135">
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<tr pageId="0" pageNumber="135">
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">Specimens</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CW</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CL</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CH</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">FW</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CW/FW</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CL/FW</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">CH/FW</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">FW/CL</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">Coll. Date</emphasis>
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</td>
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<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
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<emphasis bold="true" pageId="0" pageNumber="135">Museum</emphasis>
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</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">20</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">14.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2 ad ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">18.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31620</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">18.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4 ad ♂3</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.53</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 26.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31619</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5 ad ♂3</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 26.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31619</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6 ad ♂4</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">18.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.52</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 27.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31621</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7 ad ♂4</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 27.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31621</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8 ad ♂4</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 27.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31621</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">9 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">18.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.84</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16.75</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.77</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11.68</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16.90</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13 ad ♂1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">15.24</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.81</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31618</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">14 ad ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">15.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.93</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.46</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31620</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">15 ad ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16.50</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">ZMB Crust. 31620</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16 ad ♀2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">19.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">14.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">9.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.29</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17 ad ♀2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">17.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">8.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.96</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">18 ad ♀1</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">16.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11.66</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">7.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.26</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.48</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 23.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">19 ad ♀2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">15.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">11.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.22</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">20 ad ♀2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">14.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.80</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">21 ad ♀2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">14.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.82</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-005</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">22 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.49</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">23 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.76</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.54</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.16</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">24 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.30</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">6.10</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.25</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">25 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.87</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.86</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">3.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.40</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">26 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">13.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">10.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.60</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.28</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">27 sd ♂2</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">12.91</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">9.94</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">5.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">4.70</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.74</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">2.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">1.20</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">0.47</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">P.A.M.N 25.05.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">IFAS-006</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="135">
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">Mean</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">16.15</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">12.00</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">7.20</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">5.46</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">2.95</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">2.19</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">1.32</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">0.45</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="135" rowspan="1">-</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="135">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Carapace smooth, urogastric groove faint; postfrontal crest faint, complete, meeting anterolateral margin behind intermediate tooth (
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">Fig. 7d</figureCitation>
|
||
); exorbital, intermediate teeth large, triangular; epibranchial tooth undetectable (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Figs 4d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
|
||
). Vertical sulcus on carapace branchiostegal wall meeting anterolateral margin at intermediate tooth (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6d</figureCitation>
|
||
). Mandibular palp bi-segmented; terminal segment (TS) bilobed, with large distinct anterior lobe 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
terminal segment length (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">Fig. 15d</figureCitation>
|
||
). Third maxilliped exopod completely lacking flagellum; ischium with distinct vertical groove (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">Fig. 14d</figureCitation>
|
||
). Episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 complete (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5d</figureCitation>
|
||
). Major cheliped dactylus relatively stout, straight, not arched, enclosing long thin interspace when closed, with small distal tooth (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8g</figureCitation>
|
||
); propodus of major cheliped with three large teeth (proximal, medial, distal) (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8g</figureCitation>
|
||
); cheliped carpus inner margin with large, broad pointed distal tooth, robust subequal proximal tooth (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">Fig. 10d</figureCitation>
|
||
); cheliped merus medial inferior margin with large jagged distal tooth followed by several distinct smaller teeth (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Four species of Louisea endemic to southwestern Cameroon, right cheliped merus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 d', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure9" httpUri="https://binary.pensoft.net/fig/347897" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure9">Fig. 9d</figureCitation>
|
||
). G1TA short (TA/SS 0.22), directed outwards at 45° angle to longitudinal axis of G1SS, with distinct longitudinal groove on ventral face, proximally distinctly broad, abruptly narrow, slim and tube-like at distal two-thirds (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Figs 11d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12d</figureCitation>
|
||
). G1SS tapering slightly from broad basal margin to relatively wide distal margin (0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS basal margin); dorsal face with broad dorsal membrane (maximum width 0.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS length) at TA/SS junction (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Fig. 11d</figureCitation>
|
||
). G2TA long (TA/SS 0.44), flagellum-like, almost as long as G2SS (
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">Fig. 13d</figureCitation>
|
||
). Mature at CW 20 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="description">
|
||
<paragraph pageId="0" pageNumber="135">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Carapace ovoid, moderately high (CH/FW 1.32,
|
||
<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
|
||
= 27), wide (CW/FW 2.95,
|
||
<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
|
||
= 27), texture smooth, urogastric groove distinct. Front wide (FW/CW 0.34,
|
||
<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
|
||
= 27), deflexed, anterior margin straight (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Largest adult male (CW 17.5 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 26930) (missing right cheliped) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 1 ', www. mapress. com / j / zt) c largest adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 5, and right cheliped and p 2, p 3) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 8 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure2" httpUri="https://binary.pensoft.net/fig/347890" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure2">Figs 2d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) (missing left p 4) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 4 and right p 2) c subadult male, paratype (CW 13.8 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 4, and right cheliped and p 2 - p 5) d second largest adult male (CW 18.38 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 10 mm (b), 8.42 mm (c), 11.63 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure3" httpUri="https://binary.pensoft.net/fig/347891" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure3">3d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">4a</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
|
||
). Postfrontal crest faint but complete, ends meeting anterolateral margins at epibranchial teeth (
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">Fig. 7d</figureCitation>
|
||
); mid-groove faint, shallow; epigastric crests poorly defined (
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">Fig. 7d</figureCitation>
|
||
). Exorbital, intermediate teeth large, triangular, epibranchial tooth small but detectable (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Figs 4a</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
|
||
). Anterolateral margin of carapace lined by small granules (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Figs 4a</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
|
||
); posterolateral margin curving inward, continuous with anterolateral margin (
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">Fig. 7d</figureCitation>
|
||
); posterior carapace margin about 2/3 CW. Carapace branchiostegal wall with longitudinal, vertical sutures dividing sidewall into three parts (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6d</figureCitation>
|
||
). Longitudinal sulcus beginning at respiratory opening, curving backward across sidewall, dividing suborbital- and subhepatic regions from pterygostomial region (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6d</figureCitation>
|
||
); vertical sulcus on sidewall marked by row of granules, meeting anterolateral margin at intermediate tooth (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6d</figureCitation>
|
||
), dividing suborbital- from subhepatic regions (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Figs 6d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7d</figureCitation>
|
||
). Sternal sulcus s2/s3, deep, ends not meeting side margins of sternum (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5d</figureCitation>
|
||
); s3 with distinct central depression; s3/s4 reduced to two short lateral notches (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5d</figureCitation>
|
||
). Episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 complete (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5d</figureCitation>
|
||
). Mandibular palp bi-segmented; terminal segment (TS) bilobed, with large distinct anterior lobe 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
terminal segment length (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">Fig. 15d</figureCitation>
|
||
). Third maxilliped (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6d</figureCitation>
|
||
) filling entire buccal cavern, except for transversely oval efferent respiratory openings in superior lateral corners; exopod completely lacking flagellum; ischium with distinct vertical groove (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">Fig. 14d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Major cheliped dactylus relatively stout, straight, not arched enclosing long, thin interspace when closed, with small proximal tooth (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8g</figureCitation>
|
||
); propodus of major cheliped with three large teeth (proximal, medial, distal) (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8g</figureCitation>
|
||
); cheliped carpus inner margin with large, broad, pointed distal tooth, robust, subequal proximal tooth (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">Fig. 10d</figureCitation>
|
||
). Walking legs (p2-p5 slender, p4 longest, p5 shortest; dactyli (p2-p5) tapering to point, each bearing rows of downward-pointing sharp bristles, p5 dactylus shortest (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Largest adult male (CW 17.5 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 26930) (missing right cheliped) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 1 ', www. mapress. com / j / zt) c largest adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 5, and right cheliped and p 2, p 3) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 8 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure2" httpUri="https://binary.pensoft.net/fig/347890" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure2">Figs 2d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) (missing left p 4) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 4 and right p 2) c subadult male, paratype (CW 13.8 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 4, and right cheliped and p 2 - p 5) d second largest adult male (CW 18.38 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 10 mm (b), 8.42 mm (c), 11.63 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure3" httpUri="https://binary.pensoft.net/fig/347891" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure3">3d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Male pleon triangular, telson (a7) with rounded distal margin (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5d</figureCitation>
|
||
). G1TA short (TA/SS 0.22), directed outwards at 45° angle to longitudinal axis of G1SS, with distinct longitudinal groove on ventral face, proximally distinctly broad, abruptly narrow, slim and tube-like at distal two-thirds (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Figs 11d</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12d</figureCitation>
|
||
). G1SS tapering slightly from broad basal margin to relatively wide distal margin (0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS basal margin); dorsal face with broad dorsal membrane (maximum width 0.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS length) at TA/SS junction (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Fig. 11d</figureCitation>
|
||
). G2TA long (TA/SS 0.44), flagellum-like, almost as long as G2SS (
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">Fig. 13d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="color in life">
|
||
<paragraph pageId="0" pageNumber="135">Color in life.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Specimens of
|
||
<taxonomicName lsidName="L. nkongsamba" pageId="0" pageNumber="135" rank="species" species="nkongsamba">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. nkongsamba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. have a dark brown or green carapace and walking legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="type locality">
|
||
<paragraph pageId="0" pageNumber="135">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">Nlonako Wildlife Reserve, Nkongsamba, littoral region of Cameroon.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="etymology">
|
||
<paragraph pageId="0" pageNumber="135">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">The new species is named for Nkongsamba, the closest town to the type locality. The species epithet is a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="habitat">
|
||
<paragraph pageId="0" pageNumber="135">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
<taxonomicName lsidName="L. nkongsamba" pageId="0" pageNumber="135" rank="species" species="nkongsamba">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. nkongsamba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is known only from Nlonako Wildlife Reserve, one of the threatened tropical rainforest habitats in the littoral region of Cameroon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="remarks">
|
||
<paragraph pageId="0" pageNumber="135">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
<taxonomicName lsidName="L. nkongsamba" pageId="0" pageNumber="135" rank="species" species="nkongsamba">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. nkongsamba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. possesses numerous characters that link it to
|
||
<taxonomicName lsidName="L. edeaensis" pageId="0" pageNumber="135" rank="species" species="edeaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. edeaensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. balssi" pageId="0" pageNumber="135" rank="species" species="balssi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. balssi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="L. yabassi" pageId="0" pageNumber="135" rank="species" species="yabassi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. yabassi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Differences between these species are discussed below under general remarks.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.881.36744.figure16" httpUri="https://binary.pensoft.net/fig/347904" pageId="0" pageNumber="135" start="Figure 16" startId="F16">
|
||
<paragraph pageId="0" pageNumber="135">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">Figure 16.</emphasis>
|
||
ML tree topology for the freshwater crab taxa from Cameroon included in this study derived from mtDNA sequences corresponding to three loci (partial 16S rRNA, COI and 12S rRNA genes). BI and ML statistical values (%) on the nodes indicate posterior probabilities and bootstrap support, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |