192 lines
21 KiB
XML
192 lines
21 KiB
XML
<document id="5444A49FA05F6181DA8280EA45097F27" ID-CLB-Dataset="53872" ID-DOI="10.11646/phytotaxa.542.3.4" ID-GBIF-Dataset="79173f68-095d-4473-8104-91fd01bde9f5" ID-ISSN="1179-3163" ID-Zenodo-Dep="6416852" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649228408651" checkinUser="plazi" docAuthor="Ballard, Harvey E." docDate="2022" docId="391B87F24C058763BFFAFA82FA3EFE6E" docLanguage="en" docName="phytotaxa.542.3.4.pdf" docOrigin="Phytotaxa 542 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.542.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Paypayrola Aublet 1775" docType="key" docVersion="5" lastPageNumber="291" masterDocId="C522FF8A4C02876BBF72FFC0FF94FF9F" masterDocTitle="A new Paypayrola (Violaceae) from Panama, with notes about the species complexes for the genus" masterLastPageNumber="292" masterPageNumber="283" pageNumber="290" updateTime="1699319390668" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="04A59B22711E715BD6ADD91232F5D5CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="845AC8ABCD69665DEFCAEBDF3F93D2CA">
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<mods:title id="82CD1CC9BB6BF2288C5F2BBD7E4DCA41">A new Paypayrola (Violaceae) from Panama, with notes about the species complexes for the genus</mods:title>
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<mods:name id="C9DB5382615030CB46EC22C3DDC14214" type="personal">
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<mods:namePart id="C96F6578590FB16D1CE94898C645B3E7">Ballard, Harvey E.</mods:namePart>
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<mods:nameIdentifier id="132BB1F3B499A089D7BA42FA2E9FF9CA" type="ORCID">0000-0003-2396-784X</mods:nameIdentifier>
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<mods:affiliation id="EE427798F8BFFEFD6A129E74C5AE2D25">ballard @ ohio. edu; https: // orcid. org / 0000 - 0003 - 2396 - 784 X</mods:affiliation>
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<mods:nameIdentifier id="908DE75CD456B2F751D64EA1067E83BA" type="email">ballard@ohio.edu</mods:nameIdentifier>
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<mods:title id="1B807E931F61920E8E1155264F8C5B4D">Phytotaxa</mods:title>
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<mods:part id="ED252033544E01927009CC357043864A">
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<mods:date id="5483B8B46ED771D968A8FA51E8E740B7">2022</mods:date>
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<mods:number id="F749DBFF774AA6EB2F410D34F787BA32">2022-04-06</mods:number>
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<mods:number id="A55AF1490AC768AA0C988A66A5221497">542</mods:number>
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<mods:url id="C56E541B87D1A066AFA267D2F3A84FD1">http://dx.doi.org/10.11646/phytotaxa.542.3.4</mods:url>
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<mods:identifier id="FC9BC9B88B7CE58B89D1208A6D815719" type="DOI">10.11646/phytotaxa.542.3.4</mods:identifier>
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<treatment id="391B87F24C058763BFFAFA82FA3EFE6E" ID-DOI="http://doi.org/10.5281/zenodo.6421441" ID-GBIF-Taxon="194471927" ID-Zenodo-Dep="6421441" LSID="urn:lsid:plazi:treatment:391B87F24C058763BFFAFA82FA3EFE6E" httpUri="http://treatment.plazi.org/id/391B87F24C058763BFFAFA82FA3EFE6E" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290">
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<subSubSection id="F9A8656F4C05876CBFFAFA82FDC3FAC3" box="[136,599,1346,1373]" pageId="7" pageNumber="290" type="nomenclature">
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<paragraph id="B10D36E44C05876CBFFAFA82FDC3FAC3" blockId="7.[136,1452,1346,1444]" box="[136,599,1346,1373]" pageId="7" pageNumber="290">
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<heading id="EA4581884C05876CBFFAFA82FDC3FAC3" bold="true" box="[136,599,1346,1373]" fontSize="11" level="1" pageId="7" pageNumber="290" reason="1">
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<emphasis id="83C6EAF64C05876CBFFAFA82FDC3FAC3" bold="true" box="[136,599,1346,1373]" pageId="7" pageNumber="290">
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<emphasis id="83C6EAF64C05876CBFFAFA82FE5CFAC2" bold="true" box="[136,456,1346,1373]" italics="true" pageId="7" pageNumber="290">Taxonomic key to flowering</emphasis>
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<taxonomicName id="76B24D674C05876CBEBDFA82FDC3FAC3" ID-CoL="6JTB" authorityName="Aublet" authorityYear="1775" box="[463,599,1346,1372]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="genus">Paypayrola</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9A8656F4C05876CBFFAFAA7FE2DFA3B" pageId="7" pageNumber="290" type="discussion">
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<paragraph id="B10D36E44C05876CBFFAFAA7FE2DFA3B" blockId="7.[136,1452,1346,1444]" pageId="7" pageNumber="290">Note: all traits refer to those observed on dried herbarium specimens unless otherwise noted; corolla color was interpreted from label data.</paragraph>
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</subSubSection>
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<subSubSection id="F9A8656F4C058763BFFAFA0FFA3EFE6E" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290" type="key">
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<key id="C52361D54C058763BFFAFA0FFA3EFE6E" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290">
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<keyStep id="0A462E414C05876CBFFAFA0FFA3FF9C8" pageId="7" pageNumber="290">
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<paragraph id="B10D36E44C05876CBFFAFA0FFA3FF982" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAFA0FFA3FF982" pageId="7" pageNumber="290">
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1. Inflorescence long-pedunculate, rachis somewhat short to elongate (commonly much longer than
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), laxly flowered (sometimes moderately densely flowered but with rachis visible in
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<taxonomicName id="76B24D674C05876CBD95FA2CFCF2F99E" ID-CoL="6TW2J" authorityName="Aublet" authorityYear="1775" box="[743,870,1516,1537]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="guianensis">
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<emphasis id="83C6EAF64C05876CBD95FA2CFCF2F99E" box="[743,870,1516,1537]" italics="true" pageId="7" pageNumber="290">P. guianensis</emphasis>
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</taxonomicName>
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); sepals as broad as long or broader than long (much longer than broad in
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<taxonomicName id="76B24D674C05876CBE25F9C9FE58F982" ID-CoL="75W2J" authorityName="Tulasne" authorityYear="1847" box="[343,460,1544,1566]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="longifolia">
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<emphasis id="83C6EAF64C05876CBE25F9C9FE58F982" box="[343,460,1544,1566]" italics="true" pageId="7" pageNumber="290">P. longifolia</emphasis>
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</taxonomicName>
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); corolla deep yellow or yellow-orange to orange in life (Guianensis complex) ....................................2
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</keyLead>
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</paragraph>
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<keyLead id="0A4395D14C05876CBFFAF9E6FA3FF9C8" pageId="7" pageNumber="290">
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-. Inflorescence sessile or subsessile to short-pedunculate, rachis relatively short (usually to
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long), densely flowered with rachis not visible; sepals much longer than broad; corolla white often tinged with pink or red, or bright yellow in life .......................... 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="0A462E414C05876CBFFAF99FFA3FF955" pageId="7" pageNumber="290">
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<paragraph id="B10D36E44C05876CBFFAF99FFA3EF90F" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF99FFA3EF90F" pageId="7" pageNumber="290">
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2. Leaves long-petiolate, blades oblanceolate to linear-oblanceolate, base and apex attenuate; sepals distinctly longer than broad, elliptical ..........................................................................................................................................................................
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<taxonomicName id="76B24D674C05876CBA44F9BCFA3EF90F" ID-CoL="75W2J" authorityName="Tulasne" authorityYear="1847" box="[1334,1450,1659,1681]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="longifolia">
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<emphasis id="83C6EAF64C05876CBA44F9BCFA3EF90F" box="[1334,1450,1659,1681]" italics="true" pageId="7" pageNumber="290">P. longifolia</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B10D36E44C05876CBFFAF959FA3FF955" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF959FA3FF955" pageId="7" pageNumber="290">-. Leaves relatively short-petiolate, blades broadly oblanceolate to obovate, base cuneate to narrowly rounded, apex abruptly attenuate or subattenuate to narrowly rounded; sepals about as broad as long or broader than long, broadly ovate to deltate ....... 3</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="0A462E414C05876CBFFAF912FA3FF8E8" pageId="7" pageNumber="290">
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<paragraph id="B10D36E44C05876CBFFAF912FA3FF8BF" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF912FA3FF8BF" pageId="7" pageNumber="290">
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||
3. Leaf blades broadly lance-obovate to obovate, apex tapering or convexly rounded to a narrowly rounded tip, full-length lateral nerves per side 9–12; inflorescence elongate, laxly flowered; mature flower buds and unopened corollas long-conic ..................... ......................................................................................................................................................................................
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<taxonomicName id="76B24D674C05876CBA56F8CCFA3FF8BF" ID-CoL="6TVQJ" authorityName="Tulasne" authorityYear="1847" box="[1316,1451,1803,1825]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="grandiflora">
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<emphasis id="83C6EAF64C05876CBA56F8CCFA3FF8BF" box="[1316,1451,1803,1825]" italics="true" pageId="7" pageNumber="290">P. grandiflora</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B10D36E44C05876CBFFAF8E9FA3FF8E8" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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||
<keyLead id="0A4395D14C05876CBFFAF8E9FA3FF8E8" pageId="7" pageNumber="290">
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-. Leaf blades obovate, apex abruptly short- to long-attenuate to a sharply narrowly acute tip, full-length lateral nerves per side 5–8(–9); inflorescence shorter to elongate, laxly to moderately densely flowered; mature flower buds and unopened corollas cylindrical ......................................................................................................................................................................
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<taxonomicName id="76B24D674C05876CBA5EF8A3FA3FF8E8" ID-CoL="6TW2J" authorityName="Aublet" authorityYear="1775" box="[1324,1451,1890,1912]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="guianensis">
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<emphasis id="83C6EAF64C05876CBA5EF8A3FA3FF8E8" box="[1324,1451,1890,1912]" italics="true" pageId="7" pageNumber="290">P. guianensis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="0A462E414C05876CBFFAF8BFFA3FF82E" pageId="7" pageNumber="290">
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<paragraph id="B10D36E44C05876CBFFAF8BFFA3FF80B" blockId="7.[136,1452,1487,2084]" box="[136,1451,1918,1941]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF8BFFA3FF80B" box="[136,1451,1918,1941]" pageId="7" pageNumber="290">4. Sepals puberulent abaxially; corolla bright yellow in life (Hulkiana complex) ............................................................................... 5</keyLead>
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</paragraph>
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<paragraph id="B10D36E44C05876CBFFAF85CFA3FF82E" blockId="7.[136,1452,1487,2084]" box="[136,1451,1947,1969]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF85CFA3FF82E" box="[136,1451,1947,1969]" pageId="7" pageNumber="290">-. Sepals glabrous abaxially; corolla white often tinged with pink or red in life (Blanchetiana complex) ......................................... 6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="0A462E414C058763BFFAF878FA3EFE95" lastPageId="8" lastPageNumber="291" pageId="7" pageNumber="290">
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<paragraph id="B10D36E44C05876CBFFAF878FA3FF7BB" blockId="7.[136,1452,1487,2084]" pageId="7" pageNumber="290">
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<keyLead id="0A4395D14C05876CBFFAF878FA3FF7BB" pageId="7" pageNumber="290">
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5. Leaf blades 11.7–18.0 ×
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, apex abruptly subattenuate to long-attenuate; complete lateral nerves per side (5)6–10; inflorescence laxly to moderately densely flowered with up to 7 mature flowers at anthesis; sepals 1.8–2.4 × as long as broad, narrowly linear-oblong to narrowly oblong-triangular, apex narrowly rounded, the 1 medial nerve often raised and prominent ..... ...........................................................................................................................................................................................
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<taxonomicName id="76B24D674C05876CBA4DF7CFFA3FF7BB" ID-CoL="75VQJ" authorityName="Pulle" authorityYear="1912" box="[1343,1451,2063,2084]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="290" phylum="Tracheophyta" rank="species" species="hulkiana">
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<emphasis id="83C6EAF64C05876CBA4DF7CFFA3FF7BB" box="[1343,1451,2063,2084]" italics="true" pageId="7" pageNumber="290">P. hulkiana</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B10D36E44C0A8763BFFAFF5FFA3EFE95" blockId="8.[136,1452,159,498]" pageId="8" pageNumber="291">
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<keyLead id="0A4395D14C0A8763BFFAFF5FFA3EFE95" pageId="8" pageNumber="291">
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5. Leaf blades 18.4–29.3 × (4.1)5.9–9.8, cm, apex weakly subattenuate to narrowly rounded, rarely convexly rounded to an obtuse or broadly rounded tip; complete lateral nerves per side 8–12; inflorescence densely flowered with 6–15 mature flowers at anthesis; sepals 1.0–1.7(2.0) × as long as broad, narrowly to broadly oblong-elliptical or uncommonly suborbicular, apex broadly rounded to truncate or retuse, the (1)3 or 5 nerves frequently raised and prominent ...............................................................
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<taxonomicName id="76B24D674C0A8763BA6CFF36FA3EFE95" ID-CoL="98XWG" authorityName="H. E. Ballard" authorityYear="2022" box="[1310,1450,245,267]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="291" phylum="Tracheophyta" rank="species" species="panamensis">
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<emphasis id="83C6EAF64C0A8763BA6CFF36FA3EFE95" bold="true" box="[1310,1450,245,267]" italics="true" pageId="8" pageNumber="291">P. panamensis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="0A462E414C0A8763BFFAFED2FA3FFEDA" pageId="8" pageNumber="291">
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<paragraph id="B10D36E44C0A8763BFFAFED2FA3EFEB7" blockId="8.[136,1452,159,498]" box="[136,1450,274,296]" pageId="8" pageNumber="291">
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<keyLead id="0A4395D14C0A8763BFFAFED2FA3EFEB7" box="[136,1450,274,296]" pageId="8" pageNumber="291">
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6. Short shrub to
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; petioles <
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<quantity id="764A9B014C0A8763BE84FED2FDB9FEB7" box="[502,557,274,296]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="291" unit="mm" value="2.0">2 mm</quantity>
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long; sepals acute at apex ......................................................................................
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<taxonomicName id="76B24D674C0A8763BA49FED3FA3EFEB7" ID-CoL="6TW2H" box="[1339,1450,275,296]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="291" phylum="Tracheophyta" rank="species" species="arenacea">
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<emphasis id="83C6EAF64C0A8763BA49FED3FA3EFEB7" box="[1339,1450,275,296]" italics="true" pageId="8" pageNumber="291">P. arenacea</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B10D36E44C0A8763BFFAFEEFFA3FFEDA" blockId="8.[136,1452,159,498]" box="[136,1451,303,325]" pageId="8" pageNumber="291">
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<keyLead id="0A4395D14C0A8763BFFAFEEFFA3FFEDA" box="[136,1451,303,325]" pageId="8" pageNumber="291">
|
||
6. Shrub or treelet to
|
||
<quantity id="764A9B014C0A8763BEF5FEEFFE2CFEDA" box="[391,440,303,325]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="291" unit="m" value="12.0">12 m</quantity>
|
||
(rarely as short as
|
||
<quantity id="764A9B014C0A8763BD1FFEEFFD39FEDA" box="[621,685,303,325]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="8" pageNumber="291" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
||
); petioles>
|
||
<quantity id="764A9B014C0A8763BC51FEEFFCCEFEDA" box="[803,858,303,325]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="291" unit="mm" value="4.0">4 mm</quantity>
|
||
long; sepals broadly rounded to truncate at apex ..................7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="0A462E414C0A8763BFFAFE8CFA3EFE6E" pageId="8" pageNumber="291">
|
||
<paragraph id="B10D36E44C0A8763BFFAFE8CFA3EFE05" blockId="8.[136,1452,159,498]" pageId="8" pageNumber="291">
|
||
<keyLead id="0A4395D14C0A8763BFFAFE8CFA3EFE05" pageId="8" pageNumber="291">
|
||
7. Leaf blades linear-oblanceolate to oblong-oblanceolate, base truncate to convexly rounded (rarely subcuneate), apex weakly subattenuate, tapering or convexly rounded to a narrowly rounded or sharply acute tip; full-length lateral nerves per side 10–12; sepals 1.1–1.4 × as long as broad, broadly elliptical or oblong-ovate to suborbicular .............................................
|
||
<taxonomicName id="76B24D674C0A8763BA64FE46FA3EFE05" ID-CoL="75VQH" authorityName="Tulasne" authorityYear="1847" box="[1302,1450,389,411]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="291" phylum="Tracheophyta" rank="species" species="blanchetiana">
|
||
<emphasis id="83C6EAF64C0A8763BA64FE46FA3EFE05" box="[1302,1450,389,411]" italics="true" pageId="8" pageNumber="291">P. blanchetiana</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="B10D36E44C0A8763BFFAFE63FA3EFE6E" blockId="8.[136,1452,159,498]" pageId="8" pageNumber="291">
|
||
<keyLead id="0A4395D14C0A8763BFFAFE63FA3EFE6E" pageId="8" pageNumber="291">
|
||
7. Leaf blades oblanceolate to narrowly obovate, base cuneate, apex abruptly short- to long-attenuate to a sharply narrowly acute tip; full-length lateral nerves per side 7–8(11); sepals> 1.5 × as long as broad, narrowly elliptical or ovate-lanceolate to narrowly oblong-ovate ...............................................................................................................................................................
|
||
<taxonomicName id="76B24D674C0A8763BA68FE1DFA3EFE6E" ID-CoL="767QH" authorityName="Tulasne" authorityYear="1847" box="[1306,1450,476,498]" class="Magnoliopsida" family="Violaceae" genus="Paypayrola" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="291" phylum="Tracheophyta" rank="species" species="confertiflora">
|
||
<emphasis id="83C6EAF64C0A8763BA68FE1DFA3EFE6E" box="[1306,1450,476,498]" italics="true" pageId="8" pageNumber="291">P. confertiflora</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |