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<document id="8BD4B309FC46C8F3EDBCACFB3F72B89A" ID-CLB-Dataset="4053" ID-DOI="10.11646/zootaxa.5048.3.6" ID-GBIF-Dataset="6456e942-b39b-4899-b2a9-f4deecde44d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556495" ID-ZooBank="AB9AF652-CA1B-4BB5-927F-BC3A12BA73D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633692557300" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2021" docId="3852D52EFFC63B22FF6EFF7FB250A13D" docLanguage="en" docName="zootaxa.5048.3.6.pdf" docOrigin="Zootaxa 5048 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cycloptiloides bimaculata Tan, Japir &amp; Chung 2021, new species" docType="treatment" docVersion="8" lastPageNumber="410" masterDocId="C46BAD56FFC53B21FFF9FFE8B158A723" masterDocTitle="New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo" masterLastPageNumber="421" masterPageNumber="407" pageNumber="410" updateTime="1699459133217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EE14F74391F34B6F6CEAC37F842293CF">New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo</mods:title>
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<mods:namePart id="7F68395FF80EEFC1705CA865CBA2BD25">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="2D9BFF316F5FB3A61311393A56FCD215">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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</mods:role>
<mods:namePart id="468DD300D88FDABE3140982B877D1DD3">Japir, Razy</mods:namePart>
<mods:affiliation id="60BBDA1AAEEC21B3229EA423D5E5CC46">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. &amp; Razy. Japir @ sabah. gov. my</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="28929F5D1FA6AD727360F2B22FBB00EA">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="D8DF612073D3CE4F0661FE1BF832C15F">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. &amp; Arthur. Chung @ sabah. gov. my; https: // orcid / org / 0000 - 0002 - 9529 - 4114</mods:affiliation>
<mods:nameIdentifier id="A2D6D28A078CDE56647FC9855D4123B1" type="email">hung@sabah.gov.my</mods:nameIdentifier>
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<mods:namePart id="40A7FB158725B52713132F403ADB5EF1">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="FCED3196ECD78DDBAE8DAFA10AAF1DDF">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam. rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
<mods:nameIdentifier id="1FEA4BBCB52EACACBDCCEA24F3FEE921" type="email">rodzay.wahab@ubd.edu.bn</mods:nameIdentifier>
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<mods:date id="1094AAF4376BE33185245D53ACABC287">2021</mods:date>
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<mods:number id="CCD9E3766E64F4C30021236B8341227E">2021-10-07</mods:number>
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<treatment id="3852D52EFFC63B22FF6EFF7FB250A13D" ID-DOI="http://doi.org/10.5281/zenodo.5569504" ID-GBIF-Taxon="188638707" ID-Zenodo-Dep="5569504" LSID="urn:lsid:plazi:treatment:3852D52EFFC63B22FF6EFF7FB250A13D" httpUri="http://treatment.plazi.org/id/3852D52EFFC63B22FF6EFF7FB250A13D" lastPageNumber="410" pageId="3" pageNumber="410">
<subSubSection id="F8E137B3FFC63B22FF6EFF7FB22AA792" box="[151,882,151,178]" pageId="3" pageNumber="410" type="nomenclature">
<paragraph id="B0446438FFC63B22FF6EFF7FB22AA792" blockId="3.[151,882,151,213]" box="[151,882,151,178]" pageId="3" pageNumber="410">
<heading id="EB0CD354FFC63B22FF6EFF7FB22AA792" bold="true" box="[151,882,151,178]" fontSize="11" level="1" pageId="3" pageNumber="410" reason="1">
<emphasis id="828FB82AFFC63B22FF6EFF7FB22AA792" bold="true" box="[151,882,151,178]" pageId="3" pageNumber="410">
<taxonomicName id="77FB1FBBFFC63B22FF6EFF7FB38CA792" ID-CoL="9GZR6" authority="Tan, Japir &amp; Chung" authorityName="Tan, Japir &amp; Chung" authorityYear="2021" box="[151,724,151,178]" class="Insecta" family="Mogoplistidae" genus="Cycloptiloides" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="828FB82AFFC63B22FF6EFF7FB097A791" bold="true" box="[151,463,151,178]" italics="true" pageId="3" pageNumber="410">Cycloptiloides bimaculata</emphasis>
Tan, Japir &amp; Chung
</taxonomicName>
,
<taxonomicNameLabel id="99BC0551FFC63B22FD1BFF7FB22AA792" box="[738,882,151,177]" pageId="3" pageNumber="410" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="B0446438FFC63B22FF6EFF53B04AA7F6" blockId="3.[151,882,151,213]" box="[151,274,187,213]" pageId="3" pageNumber="410">
(
<figureCitation id="28C078BDFFC63B22FF66FF53B052A7F6" box="[159,266,187,213]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1928,1952]" captionStartId-1="5.[151,250,1613,1637]" captionStartId-2="6.[151,250,1918,1942]" captionTargetBox-0="[151,1436,181,1904]" captionTargetBox-1="[170,1412,200,1569]" captionTargetBox-2="[154,1434,564,1884]" captionTargetId-0="figure-20@4.[151,1436,181,1904]" captionTargetId-1="figure-19@5.[151,1436,181,1589]" captionTargetId-2="figure-206@6.[151,1436,561,1894]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male (A) and female (B) adults on leaf litters in Rainforest Discovery Centre, Sandakan." captionText-1="FIGURE 3. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype habitus in ethanol in dorsal (A) and lateral (B) views. Scale bar: 1 mm." captionText-2="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.5556501" figureDoi-1="http://doi.org/10.5281/zenodo.5556503" figureDoi-2="http://doi.org/10.5281/zenodo.5556505" httpUri-0="https://zenodo.org/record/5556501/files/figure.png" httpUri-1="https://zenodo.org/record/5556503/files/figure.png" httpUri-2="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Figs. 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF6EFEE8B367A676" pageId="3" pageNumber="410" type="reference_group">
<paragraph id="B0446438FFC63B22FF6EFEE8B03DA615" blockId="3.[151,1436,256,341]" pageId="3" pageNumber="410">
<treatmentCitationGroup id="90EB4316FFC63B22FF6EFEE8B03DA615" pageId="3" pageNumber="410">
<taxonomicName id="77FB1FBBFFC63B22FF6EFEE8B346A63B" ID-CoL="435J5" authority="(Ingrisch, 1998)" baseAuthorityName="Ingrisch" baseAuthorityYear="1998" box="[151,542,256,280]" class="Insecta" family="Mogoplistidae" genus="Micrornebius" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="828FB82AFFC63B22FF6EFEE8B020A634" box="[151,376,256,280]" italics="true" pageId="3" pageNumber="410">Micrornebius incertus</emphasis>
(
<bibRefCitation id="D46A19C9FFC63B22FE7CFEE8B34FA63B" author="Ingrisch, S." box="[389,535,256,280]" pageId="3" pageNumber="410" pagination="225 - 234" refId="ref4894" refString="Ingrisch, S. (1998) New Mogoplistinae from the Kinabalu region in Sabah, North Borneo (Insecta: Ensifera: Grylloidea: Mogoplistidae). Senckenbergiana Biologica, 77, 225 - 234." type="journal article" year="1998">Ingrisch, 1998</bibRefCitation>
)
</taxonomicName>
—
<treatmentCitation id="315A4229FFC63B22FDC0FEE8B3A7A63B" author="Ingrisch, S." box="[569,767,256,280]" page="232" pageId="3" pageNumber="410" year="1998">
<bibRefCitation id="D46A19C9FFC63B22FDC0FEE8B3A7A63B" author="Ingrisch, S." box="[569,767,256,280]" pageId="3" pageNumber="410" pagination="225 - 234" refId="ref4894" refString="Ingrisch, S. (1998) New Mogoplistinae from the Kinabalu region in Sabah, North Borneo (Insecta: Ensifera: Grylloidea: Mogoplistidae). Senckenbergiana Biologica, 77, 225 - 234." type="journal article" year="1998">Ingrisch, 1998: 232</bibRefCitation>
</treatmentCitation>
[one of the females (not a
<typeStatus id="6F40DA9AFFC63B22FBF5FEE8B561A63B" box="[1036,1081,256,280]" pageId="3" pageNumber="410">type</typeStatus>
) erroneous described under
<taxonomicName id="77FB1FBBFFC63B22FAAEFEE8B005A615" ID-CoL="34W6D" authorityName="Ingrisch" authorityYear="1998" class="Insecta" family="Mogoplistidae" genus="Derectaotus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="828FB82AFFC63B22FAAEFEE8B005A615" italics="true" pageId="3" pageNumber="410">Derectaotus incertus</emphasis>
</taxonomicName>
]
</treatmentCitationGroup>
</paragraph>
<paragraph id="B0446438FFC63B22FF6EFED5B367A676" blockId="3.[151,1436,256,341]" box="[151,575,317,341]" pageId="3" pageNumber="410">
<treatmentCitationGroup id="90EB4316FFC63B22FF6EFED5B367A676" box="[151,575,317,341]" pageId="3" pageNumber="410">
<taxonomicName id="77FB1FBBFFC63B22FF6EFED5B003A676" box="[151,347,317,341]" class="Insecta" family="Mogoplistidae" genus="Cycloptiloides" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="828FB82AFFC63B22FF6EFED5B071A676" box="[151,297,317,341]" italics="true" pageId="3" pageNumber="410">Cycloptiloides</emphasis>
sp. 1
</taxonomicName>
—
<treatmentCitation id="315A4229FFC63B22FE8FFED5B367A676" author="Ingrisch, S." box="[374,575,317,341]" page="185" pageId="3" pageNumber="410" year="2006">
<bibRefCitation id="D46A19C9FFC63B22FE8FFED5B367A676" author="Ingrisch, S." box="[374,575,317,341]" pageId="3" pageNumber="410" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006: 185</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF6EFE6BB051A556" pageId="3" pageNumber="410" type="materials_examined">
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<emphasis id="828FB82AFFC63B22FF6EFE6BB0D8A6BE" bold="true" box="[151,384,387,413]" pageId="3" pageNumber="410">Material examined.</emphasis>
<materialsCitation id="00936E65FFC63B22FE7FFE6BB0BFA6C6" ID-GBIF-Occurrence="3383518306" collectingDate="2019-01-09" collectionCode="FRC" collectorName="M. K. Tan &amp; R. Japir &amp; M. Binti &amp; J. L. Yukang" country="Malaysia" county="Sandakan" elevation="5" latitude="5.8758" location="Rainforest Discovery Centre" longLatPrecision="1" longitude="117.94299" municipality="Sepilok" pageId="3" pageNumber="410" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="828FB82AFFC63B22FE7FFE6BB0ABA6BE" bold="true" box="[390,499,387,413]" pageId="3" pageNumber="410">
<typeStatus id="6F40DA9AFFC63B22FE7FFE6BB0ABA6BE" box="[390,499,387,413]" pageId="3" pageNumber="410" type="holotype">Holotype</typeStatus>
</emphasis>
, male (SDK.19.20), EAST
<collectingCountry id="C8EC24A8FFC63B22FCDCFE6BB2EEA6BE" box="[805,950,387,413]" name="Malaysia" pageId="3" pageNumber="410">MALAYSIA</collectingCountry>
,
<collectingRegion id="723FAADAFFC63B22FC3AFE6BB552A6BE" box="[963,1034,387,413]" country="Malaysia" name="Sabah" pageId="3" pageNumber="410">Sabah</collectingRegion>
,
<collectingCounty id="59251CB4FFC63B22FBECFE6BB5DDA6BE" box="[1045,1157,387,413]" pageId="3" pageNumber="410">Sandakan</collectingCounty>
,
<collectingMunicipality id="5020FE42FFC63B22FB76FE6BB5BFA6BE" box="[1167,1255,387,413]" pageId="3" pageNumber="410">Sepilok</collectingMunicipality>
,
<location id="B52432E3FFC63B22FB08FE6BB06FA6E2" LSID="urn:lsid:plazi:treatment:3852D52EFFC63B22FF6EFF7FB250A13D:B52432E3FFC63B22FB08FE6BB06FA6E2" country="Malaysia" county="Sandakan" latitude="5.8758" longLatPrecision="1" longitude="117.94299" municipality="Sepilok" name="Rainforest Discovery Centre" pageId="3" pageNumber="410" stateProvince="Sabah">Rainforest Discovery Centre</location>
,
<geoCoordinate id="D5CF02FFFFC63B22FEB9FE4FB0EBA6E1" box="[320,435,423,450]" degrees="5.87580" direction="north" orientation="latitude" pageId="3" pageNumber="410" precision="1" value="5.8758">N5.87580</geoCoordinate>
,
<geoCoordinate id="D5CF02FFFFC63B22FE45FE4FB31FA6E2" box="[444,583,423,449]" degrees="117.94299" direction="east" orientation="longitude" pageId="3" pageNumber="410" precision="1" value="117.94299">E117.94299</geoCoordinate>
, 39.1±
<quantity id="7703C9DDFFC63B22FD69FE4FB38EA6E2" box="[656,726,423,450]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="410" unit="m" value="5.4">
<elevation id="3BD6830BFFC63B22FD69FE4FB38EA6E2" box="[656,726,423,450]" metricMagnitude="0" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="410" unit="m" value="5.4">5.4 m</elevation>
</quantity>
.a.s.l., leaf litter,
<date id="C44542F8FFC63B22FC7EFE4FB56AA6E2" box="[903,1074,423,450]" pageId="3" pageNumber="410" value="2019-01-09">
<collectingDate id="D401BB10FFC63B22FC7EFE4FB56AA6E2" box="[903,1074,423,450]" pageId="3" pageNumber="410" value="2019-01-09">9 January 2019</collectingDate>
</date>
, 1911h, coll.
<collectorName id="1D0E01EEFFC63B22FB3DFE40B46EA6E1" box="[1220,1334,424,450]" pageId="3" pageNumber="410">M. K. Tan</collectorName>
, R.
<collectorName id="1D0E01EEFFC63B22FA99FE4FB1EBA6C5" pageId="3" pageNumber="410">Japir, M</collectorName>
.
<collectorName id="1D0E01EEFFC63B22FF47FE23B069A6C5" box="[190,305,459,486]" pageId="3" pageNumber="410">Binti, J. L</collectorName>
. Yukang (
<collectionCode id="D6EAFCFDFFC63B22FE59FE23B083A6C6" box="[416,475,459,485]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="3" pageNumber="410">FRC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B0446438FFC63B22FF3EFE07B051A556" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFE07B067A52A" bold="true" box="[199,319,495,521]" pageId="3" pageNumber="410">
<typeStatus id="6F40DA9AFFC63B22FF3EFE07B067A52A" box="[199,319,495,521]" pageId="3" pageNumber="410" type="paratype">Paratypes</typeStatus>
</emphasis>
,
<specimenCount id="A6FDAFB1FFC63B22FEB4FE07B0E6A52A" box="[333,446,495,521]" pageId="3" pageNumber="410" type="female">2 females</specimenCount>
:
<materialsCitation id="00936E65FFC63B22FE31FE07B2B6A50E" ID-GBIF-Occurrence="3383518304" collectingDate="2019-01-09" collectorName="M. K. Tan &amp; R. Japir &amp; M. Binti &amp; J. L. Yukang" elevation="5" latitude="5.87356" longLatPrecision="1" longitude="117.94172" pageId="3" pageNumber="410" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A6FDAFB1FFC63B22FE31FE07B371A52A" box="[456,553,495,521]" pageId="3" pageNumber="410" type="female">1 female</specimenCount>
(SDK.19.19), same locality,
<geoCoordinate id="D5CF02FFFFC63B22FC96FE07B2BAA529" box="[879,994,495,522]" degrees="5.87356" direction="north" orientation="latitude" pageId="3" pageNumber="410" precision="1" value="5.87356">N5.87356</geoCoordinate>
,
<geoCoordinate id="D5CF02FFFFC63B22FC15FE07B52EA52A" box="[1004,1142,495,521]" degrees="117.94172" direction="east" orientation="longitude" pageId="3" pageNumber="410" precision="1" value="117.94172">E117.94172</geoCoordinate>
, 49.1±
<quantity id="7703C9DDFFC63B22FB38FE07B451A52A" box="[1217,1289,495,522]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="410" unit="m" value="5.2">
<elevation id="3BD6830BFFC63B22FB38FE07B451A52A" box="[1217,1289,495,522]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="410" unit="m" value="5.2">5.2 m</elevation>
</quantity>
.a.s.l., leaf litter,
<date id="C44542F8FFC63B22FF3BFDFBB029A50E" box="[194,369,531,558]" pageId="3" pageNumber="410" value="2019-01-09">
<collectingDate id="D401BB10FFC63B22FF3BFDFBB029A50E" box="[194,369,531,558]" pageId="3" pageNumber="410" value="2019-01-09">9 January 2019</collectingDate>
</date>
, 1105h, coll.
<collectorName id="1D0E01EEFFC63B22FDF0FDFCB326A50D" box="[521,638,532,558]" pageId="3" pageNumber="410">M. K. Tan</collectorName>
, R.
<collectorName id="1D0E01EEFFC63B22FD52FDFBB253A50D" box="[683,779,531,558]" pageId="3" pageNumber="410">Japir, M</collectorName>
.
<collectorName id="1D0E01EEFFC63B22FCEFFDFBB2D2A50D" box="[790,906,531,558]" pageId="3" pageNumber="410">Binti, J. L</collectorName>
. Yukang
</materialsCitation>
;
<materialsCitation id="00936E65FFC63B22FC03FDFBB05DA556" ID-GBIF-Occurrence="3383518305" collectingDate="2019-01-09" collectionCode="FRC" collectorName="M. K. Tan &amp; R. Japir &amp; M. Binti &amp; J. L. Yukang" elevation="5" latitude="5.87553" longLatPrecision="1" longitude="117.94116" pageId="3" pageNumber="410" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A6FDAFB1FFC63B22FC03FDFBB503A50E" box="[1018,1115,531,557]" pageId="3" pageNumber="410" type="female">1 female</specimenCount>
(SDK.19.35), same locality,
<geoCoordinate id="D5CF02FFFFC63B22FF6EFDDFB051A571" box="[151,265,567,594]" degrees="5.87553" direction="north" orientation="latitude" pageId="3" pageNumber="410" precision="1" value="5.87553">N5.87553</geoCoordinate>
,
<geoCoordinate id="D5CF02FFFFC63B22FEECFDDFB0C6A572" box="[277,414,567,593]" degrees="117.94116" direction="east" orientation="longitude" pageId="3" pageNumber="410" precision="1" value="117.94116">E117.94116</geoCoordinate>
, 33.4±
<quantity id="7703C9DDFFC63B22FE13FDDFB369A572" box="[490,561,567,593]" metricMagnitude="0" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="410" unit="m" value="4.9">
<elevation id="3BD6830BFFC63B22FE13FDDFB369A572" box="[490,561,567,593]" metricMagnitude="0" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="410" unit="m" value="4.9">4.9 m</elevation>
</quantity>
.a.s.l.,
<date id="C44542F8FFC63B22FD8AFDDFB27AA572" box="[627,802,567,594]" pageId="3" pageNumber="410" value="2019-01-09">
<collectingDate id="D401BB10FFC63B22FD8AFDDFB27AA572" box="[627,802,567,594]" pageId="3" pageNumber="410" value="2019-01-09">9 January 2019</collectingDate>
</date>
, 2040h, coll.
<collectorName id="1D0E01EEFFC63B22FC42FDD0B568A571" box="[955,1072,568,594]" pageId="3" pageNumber="410">M. K. Tan</collectorName>
, R.
<collectorName id="1D0E01EEFFC63B22FBA4FDDFB5E5A571" box="[1117,1213,567,594]" pageId="3" pageNumber="410">Japir, M</collectorName>
.
<collectorName id="1D0E01EEFFC63B22FB31FDDFB464A571" box="[1224,1340,567,594]" pageId="3" pageNumber="410">Binti, J. L</collectorName>
. Yukang (all
<collectionCode id="D6EAFCFDFFC63B22FF3DFDB3B1A6A556" box="[196,254,603,629]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="3" pageNumber="410">FRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF3EFD97B5D7A59E" pageId="3" pageNumber="410" type="discussion">
<paragraph id="B0446438FFC63B22FF3EFD97B5D7A59E" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFD97B063A5BA" bold="true" box="[199,315,639,665]" pageId="3" pageNumber="410">Remarks.</emphasis>
<bibRefCitation id="D46A19C9FFC63B22FEBBFD97B0B7A5B9" author="Ingrisch, S." box="[322,495,639,666]" pageId="3" pageNumber="410" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch (2006)</bibRefCitation>
described the female of this species as
<taxonomicName id="77FB1FBBFFC63B22FC5EFD97B5DCA5BA" box="[935,1156,639,666]" class="Insecta" family="Mogoplistidae" genus="Cycloptiloides" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="828FB82AFFC63B22FC5EFD97B511A5BA" box="[935,1097,639,665]" italics="true" pageId="3" pageNumber="410">Cycloptiloides</emphasis>
sp. 1
</taxonomicName>
without providing a species name. Here, we provide a formal name, assign a male
<typeStatus id="6F40DA9AFFC63B22FCD2FD4BB2D5A59E" box="[811,909,675,701]" pageId="3" pageNumber="410" type="holotype">holotype</typeStatus>
and describe the male.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF3EFD2FB00BA46D" pageId="3" pageNumber="410" type="diagnosis">
<paragraph id="B0446438FFC63B22FF3EFD2FB00BA46D" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFD2FB01AA5C2" bold="true" box="[199,322,711,737]" pageId="3" pageNumber="410">Diagnosis.</emphasis>
The male differs from congeners by the supra-anal plate trapezoidal in shape with apical margin broadly rounded, with dark scales and yellow brown setae laterally and shiny pale in the middle (forming two spots). The female differs from congeners by shorter maxillary palps and ovipositor and more transverse supra-anal plate (
<bibRefCitation id="D46A19C9FFC63B22FF66FCDBB01FA46D" author="Ingrisch, S." box="[159,327,819,846]" pageId="3" pageNumber="410" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF3EFCBFB089A4B6" pageId="3" pageNumber="410" type="etymology">
<paragraph id="B0446438FFC63B22FF3EFCBFB089A4B6" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFCBFB016A452" bold="true" box="[199,334,855,881]" pageId="3" pageNumber="410">Etymology.</emphasis>
The species name refers to the two spots in the middle of the supra-anal plate; bi = two in Latin, maculata = spotted in Latin.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF3EFC77B339A2DA" pageId="3" pageNumber="410" type="description">
<paragraph id="B0446438FFC63B22FF3EFC77B324A3B1" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFC77B001A49A" bold="true" box="[199,345,927,953]" pageId="3" pageNumber="410">Description.</emphasis>
Habitus of male as shown in
<figureCitation id="28C078BDFFC63B22FD53FC77B3A1A49A" box="[682,761,927,953]" captionStart="FIGURE 2" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[151,1436,181,1904]" captionTargetId="figure-20@4.[151,1436,181,1904]" captionTargetPageId="4" captionText="FIGURE 2. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male (A) and female (B) adults on leaf litters in Rainforest Discovery Centre, Sandakan." figureDoi="http://doi.org/10.5281/zenodo.5556501" httpUri="https://zenodo.org/record/5556501/files/figure.png" pageId="3" pageNumber="410">Figs. 2</figureCitation>
,
<figureCitation id="28C078BDFFC63B22FCFFFC77B24EA49A" box="[774,790,927,953]" captionStart="FIGURE 3" captionStartId="5.[151,250,1613,1637]" captionTargetBox="[170,1412,200,1569]" captionTargetId="figure-19@5.[151,1436,181,1589]" captionTargetPageId="5" captionText="FIGURE 3. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype habitus in ethanol in dorsal (A) and lateral (B) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556503" httpUri="https://zenodo.org/record/5556503/files/figure.png" pageId="3" pageNumber="410">3</figureCitation>
. Dorsum of head flattened (
<figureCitation id="28C078BDFFC63B22FBABFC77B5F5A49A" box="[1106,1197,927,953]" captionStart="FIGURE 3" captionStartId="5.[151,250,1613,1637]" captionTargetBox="[170,1412,200,1569]" captionTargetId="figure-19@5.[151,1436,181,1589]" captionTargetPageId="5" captionText="FIGURE 3. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype habitus in ethanol in dorsal (A) and lateral (B) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556503" httpUri="https://zenodo.org/record/5556503/files/figure.png" pageId="3" pageNumber="410">Fig. 3A</figureCitation>
). Frontal rostrum 1.2 times wider than scapus. Maxillary palps with apical (= fifth) a little longer than subapical (= fourth) and third segments, subapical and third segments subequal lengths; apical segment little widened (
<figureCitation id="28C078BDFFC63B22FBABFC0FB5F4A322" box="[1106,1196,999,1025]" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Fig. 4A</figureCitation>
). Pronotum about 1.5 times longer than wide with anterior dorsal margin distinctly concave; lateral margin subparallel in anterior half and faintly widening posteriorly; covering tegmen entirely; posterior margin convex. Fore tibia with internal tympanum large and elongate oval (
<figureCitation id="28C078BDFFC63B22FE48FBBBB352A34E" box="[433,522,1107,1133]" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Fig. 4B</figureCitation>
); without external tympanum. Hind femur 1.4 times longer than hind tibia; hind tibia 2.1 times longer than hind metatarsus.
</paragraph>
<paragraph id="B0446438FFC63B22FF3EFB73B06DA3DE" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
Male. Supra-anal plate trapezoidal with apical margin broadly rounded, with setae at lateral parts (
<figureCitation id="28C078BDFFC63B22FACAFB73B1E6A3FA" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Figs. 4C, 4D</figureCitation>
). Subgenital plate trapezoidal, posterior margin slightly convex (
<figureCitation id="28C078BDFFC63B22FC6FFB57B2B4A3FA" box="[918,1004,1215,1241]" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Fig. 4F</figureCitation>
). Epiphallus and phallic complex hyaline (
<figureCitation id="28C078BDFFC63B22FF28FB0BB070A3DE" box="[209,296,1251,1277]" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="B0446438FFC63B22FF3EFAEFB043A266" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
Female. Already sufficiently described in
<bibRefCitation id="D46A19C9FFC63B22FD61FAEFB210A201" author="Ingrisch, S." box="[664,840,1287,1314]" pageId="3" pageNumber="410" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch (2006)</bibRefCitation>
. Our female specimens from Sandakan agree with the description.
</paragraph>
<paragraph id="B0446438FFC63B22FF3EFAA7B0E3A292" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
Colouration. General colouration similar to female described in
<bibRefCitation id="D46A19C9FFC63B22FC50FAA7B504A249" author="Ingrisch, S." box="[937,1116,1359,1386]" pageId="3" pageNumber="410" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch (2006)</bibRefCitation>
. Male supra-anal plate with dark scales laterally, middle shining pale coloured (forming two spots) (
<figureCitation id="28C078BDFFC63B22FC30FA9BB538A2AE" box="[969,1120,1395,1421]" captionStart="FIGURE 4" captionStartId="6.[151,250,1918,1942]" captionTargetBox="[154,1434,564,1884]" captionTargetId="figure-206@6.[151,1436,561,1894]" captionTargetPageId="6" captionText="FIGURE 4. Cycloptiloides bimaculata Tan, Japir &amp; Chung, sp. nov. male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="3" pageNumber="410">Figs. 4C, 4D</figureCitation>
). Cercus (both male and females) with brown scales.
</paragraph>
<paragraph id="B0446438FFC63B22FF3EFA53B339A2DA" blockId="3.[151,1437,387,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EFA53B0BBA2F6" bold="true" box="[199,483,1467,1493]" pageId="3" pageNumber="410">Measurements (in mm).</emphasis>
<typeStatus id="6F40DA9AFFC63B22FE13FA53B30AA2F6" box="[490,594,1467,1493]" pageId="3" pageNumber="410" type="holotype">Holotype</typeStatus>
BL = 3.9; FRW = 0.27; SW = 0.14; EW = 0.14; PL = 1.9; PW = 1.3; HFL = 2.5; HFW = 0.9; HTL = 1.8; HML = 0.8.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC63B22FF3EF9EBB250A13D" box="[199,776,1539,1566]" pageId="3" pageNumber="410" type="distribution">
<paragraph id="B0446438FFC63B22FF3EF9EBB250A13D" blockId="3.[151,1437,387,1566]" box="[199,776,1539,1566]" pageId="3" pageNumber="410">
<emphasis id="828FB82AFFC63B22FF3EF9EBB039A13E" bold="true" box="[199,353,1539,1565]" pageId="3" pageNumber="410">Distribution.</emphasis>
Borneo (
<collectingRegion id="723FAADAFFC63B22FE30F9EBB34BA13E" box="[457,531,1539,1565]" country="Malaysia" name="Sabah" pageId="3" pageNumber="410">Sabah</collectingRegion>
:
<taxonomicName id="77FB1FBBFFC63B22FDE4F9EBB259A13D" authority=", Sandakan" authorityName="Sandakan" box="[541,769,1539,1566]" class="Insecta" family="Mogoplistidae" genus="Apterornebius" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="kinabalu">Kinabalu, Sandakan</taxonomicName>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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