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<document id="91573DF818C435718B5C7C27F5079D38" ID-CLB-Dataset="4053" ID-DOI="10.11646/zootaxa.5048.3.6" ID-GBIF-Dataset="6456e942-b39b-4899-b2a9-f4deecde44d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5556495" ID-ZooBank="AB9AF652-CA1B-4BB5-927F-BC3A12BA73D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1633692557300" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2021" docId="3852D52EFFC03B28FF6EF932B3F2A6F1" docLanguage="en" docName="zootaxa.5048.3.6.pdf" docOrigin="Zootaxa 5048 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ectatoderus nigrofasciatus Tan &amp; Wahab 2021, new species" docType="treatment" docVersion="7" lastPageNumber="416" masterDocId="C46BAD56FFC53B21FFF9FFE8B158A723" masterDocTitle="New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo" masterLastPageNumber="421" masterPageNumber="407" pageNumber="412" updateTime="1699459133217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CB862C761E737AC24358D50C5405C99F">New species and taxonomic notes of scaly crickets (Orthoptera: Mogoplistidae Mogoplistinae) from Borneo</mods:title>
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<mods:name id="6087222AF5431E914DB7338B5C520C22" type="personal">
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<mods:namePart id="FA2AB1949673674FFD33E8AAD51DC0BF">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="A7E661FFF5E1033E89C8BFA760CB5A60">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:namePart id="A58F8B3F018E31177946DAE2522BD802">Japir, Razy</mods:namePart>
<mods:affiliation id="CED885FDDBC8DF05B80B6C46D39A7FD0">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. &amp; Razy. Japir @ sabah. gov. my</mods:affiliation>
</mods:name>
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<mods:namePart id="BC82D7DD629247A87931E592EBF1400C">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="B19425C226C7D39622FF1B594A8BBED4">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. &amp; Arthur. Chung @ sabah. gov. my; https: // orcid / org / 0000 - 0002 - 9529 - 4114</mods:affiliation>
<mods:nameIdentifier id="8DED3B3A70A108341BC13F4C4455FC77" type="email">hung@sabah.gov.my</mods:nameIdentifier>
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<mods:role id="9140E590A49A46C884A9A77049E8E7C2">
<mods:roleTerm id="B5238A8B3CCFFB61EA4131E0FF055A9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="95E7EBA724A1D045E81EB7F5181B9811">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="EF3F5789282E16FDB1FDED3460833358">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam. rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
<mods:nameIdentifier id="41E831C5A142FE406C322B688DEABCDE" type="email">rodzay.wahab@ubd.edu.bn</mods:nameIdentifier>
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<mods:title id="5A4F2DC1C097C30CBE9BB4DB5507DC53">Zootaxa</mods:title>
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<mods:date id="59C9259DC8E8948F4EB818AE3AEE3D7B">2021</mods:date>
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<mods:number id="04CE37B6FBA0F433C56A9B01546755A8">2021-10-07</mods:number>
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<mods:number id="8481BE65A78FA9D147A5C8EA1165667A">5048</mods:number>
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<treatment id="3852D52EFFC03B28FF6EF932B3F2A6F1" ID-DOI="http://doi.org/10.5281/zenodo.5556493" ID-GBIF-Taxon="188638713" ID-Zenodo-Dep="5556493" LSID="urn:lsid:plazi:treatment:3852D52EFFC03B28FF6EF932B3F2A6F1" httpUri="http://treatment.plazi.org/id/3852D52EFFC03B28FF6EF932B3F2A6F1" lastPageId="9" lastPageNumber="416" pageId="5" pageNumber="412">
<subSubSection id="F8E137B3FFC03B24FF6EF932B26AA1D6" box="[151,818,1754,1781]" pageId="5" pageNumber="412" type="nomenclature">
<paragraph id="B0446438FFC03B24FF6EF932B26AA1D6" blockId="5.[151,818,1754,1818]" box="[151,818,1754,1781]" pageId="5" pageNumber="412">
<heading id="EB0CD354FFC03B24FF6EF932B26AA1D6" bold="true" box="[151,818,1754,1781]" fontSize="11" level="1" pageId="5" pageNumber="412" reason="1">
<emphasis id="828FB82AFFC03B24FF6EF932B26AA1D6" bold="true" box="[151,818,1754,1781]" pageId="5" pageNumber="412">
<taxonomicName id="77FB1FBBFFC03B24FF6EF932B3CDA1D6" ID-CoL="9GZR8" authority="Tan &amp; Wahab" authorityName="Tan &amp; Wahab" authorityYear="2021" box="[151,661,1754,1781]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="412" phylum="Arthropoda" rank="species" species="nigrofasciatus" status="sp. nov.">
<emphasis id="828FB82AFFC03B24FF6EF932B085A1D7" bold="true" box="[151,477,1754,1781]" italics="true" pageId="5" pageNumber="412">Ectatoderus nigrofasciatus</emphasis>
Tan &amp; Wahab
</taxonomicName>
,
<taxonomicNameLabel id="99BC0551FFC03B24FD5BF933B26AA1D6" box="[674,818,1755,1781]" pageId="5" pageNumber="412" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC03B24FF6EF917B04AA039" box="[151,274,1791,1818]" pageId="5" pageNumber="412" type="description">
<paragraph id="B0446438FFC03B24FF6EF917B04AA039" blockId="5.[151,818,1754,1818]" box="[151,274,1791,1818]" pageId="5" pageNumber="412">
(
<figureCitation id="28C078BDFFC03B24FF66F917B052A039" box="[159,266,1791,1818]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,250,1061,1085]" captionStartId-1="8.[151,250,1943,1967]" captionStartId-2="10.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1037]" captionTargetBox-1="[184,1414,201,1920]" captionTargetBox-2="[179,1408,181,1920]" captionTargetId-0="figure-19@7.[151,1436,181,1037]" captionTargetId-1="figure-20@8.[171,1417,181,1920]" captionTargetId-2="figure-20@10.[179,1408,181,1920]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIGURE 5. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male holotype on a palm/ rattan leaf in Belait." captionText-1="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." captionText-2="FIGURE 7. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male phallic complex in dorsal (A), ventral (B), lateral (C) and latero-ventral (D) views; and apex of medial valve in ventral view (E). Scale bars: 1 mm (A—D), 0.5 mm (E)." figureDoi-0="http://doi.org/10.5281/zenodo.5556507" figureDoi-1="http://doi.org/10.5281/zenodo.5556509" figureDoi-2="http://doi.org/10.5281/zenodo.5556511" httpUri-0="https://zenodo.org/record/5556507/files/figure.png" httpUri-1="https://zenodo.org/record/5556509/files/figure.png" httpUri-2="https://zenodo.org/record/5556511/files/figure.png" pageId="5" pageNumber="412">Figs. 5–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC03B24FF6EF8AFB000A08A" pageId="5" pageNumber="412" type="materials_examined">
<paragraph id="B0446438FFC03B24FF6EF8AFB000A08A" blockId="5.[151,1437,1863,2034]" pageId="5" pageNumber="412">
<emphasis id="828FB82AFFC03B24FF6EF8AFB0DCA042" bold="true" box="[151,388,1863,1889]" pageId="5" pageNumber="412">Material examined.</emphasis>
<materialsCitation id="00936E65FFC03B24FE76F8AFB000A08A" ID-GBIF-Occurrence="3383518303" collectingDate="2019-07-07" collectionCode="BRU, UBDM" collectorName="M. K. Tan" country="Brunei" latitude="4.63354" location="Andulau Forest Reserve" longLatPrecision="1" longitude="114.51047" municipality="Jalan Labi" pageId="5" pageNumber="412" specimenCount="1" stateProvince="Belait" typeStatus="holotype">
<emphasis id="828FB82AFFC03B24FE76F8AFB0A4A042" bold="true" box="[399,508,1863,1889]" pageId="5" pageNumber="412">
<typeStatus id="6F40DA9AFFC03B24FE76F8AFB0A4A042" box="[399,508,1863,1889]" pageId="5" pageNumber="412" type="holotype">Holotype</typeStatus>
</emphasis>
, male (
<collectionCode id="D6EAFCFDFFC03B24FDAFF8AFB3CDA042" box="[598,661,1863,1889]" country="USA" lsid="urn:lsid:biocol.org:col:15210" name="Brown University" pageId="5" pageNumber="412" type="Herbarium">BRU</collectionCode>
.19.70),
<collectingCountry id="C8EC24A8FFC03B24FD0BF8A0B571A042" box="[754,1065,1863,1890]" name="Brunei" pageId="5" pageNumber="412">BRUNEI DARUSSALAM</collectingCountry>
,
<collectingRegion id="723FAADAFFC03B24FBCEF8AFB584A042" box="[1079,1244,1863,1889]" country="Brunei" name="Belait" pageId="5" pageNumber="412">Belait District</collectingRegion>
,
<collectingMunicipality id="5020FE42FFC03B24FB13F8AFB439A042" box="[1258,1377,1863,1889]" pageId="5" pageNumber="412">Jalan Labi</collectingMunicipality>
near
<location id="B52432E3FFC03B24FF6EF883B0F2A0A5" LSID="urn:lsid:plazi:treatment:3852D52EFFC03B28FF6EF932B3F2A6F1:B52432E3FFC03B24FF6EF883B0F2A0A5" box="[151,426,1899,1926]" country="Brunei" latitude="4.63354" longLatPrecision="1" longitude="114.51047" municipality="Jalan Labi" name="Andulau Forest Reserve" pageId="5" pageNumber="412" stateProvince="Belait">Andulau Forest Reserve</location>
,
<geoCoordinate id="D5CF02FFFFC03B24FE4FF883B370A0A5" box="[438,552,1899,1926]" degrees="4.63354" direction="north" orientation="latitude" pageId="5" pageNumber="412" precision="1" value="4.63354">N4.63354</geoCoordinate>
,
<geoCoordinate id="D5CF02FFFFC03B24FDCDF883B3E6A0A5" box="[564,702,1899,1926]" degrees="114.51047" direction="east" orientation="longitude" pageId="5" pageNumber="412" precision="1" value="114.51047">E114.51047</geoCoordinate>
, 76.6±5.0 m.a.s.l., on rattan foliage,
<date id="C44542F8FFC03B24FBA5F884B5BBA0A6" box="[1116,1251,1899,1926]" pageId="5" pageNumber="412" value="2019-07-07">
<collectingDate id="D401BB10FFC03B24FBA5F884B5BBA0A6" box="[1116,1251,1899,1926]" pageId="5" pageNumber="412" value="2019-07-07">7 July 2019</collectingDate>
</date>
, 2042h, coll.
<collectorName id="1D0E01EEFFC03B24FA85F884B1BBA089" pageId="5" pageNumber="412">M. K. Tan</collectorName>
(
<collectionCode id="D6EAFCFDFFC03B24FF08F867B013A08A" box="[241,331,1935,1961]" pageId="5" pageNumber="412">UBDM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC03B26FF3EF85BB433A3EE" lastPageId="7" lastPageNumber="414" pageId="5" pageNumber="412" type="diagnosis">
<paragraph id="B0446438FFC03B27FF3EF85BB097A7DA" blockId="5.[151,1437,1863,2034]" lastBlockId="6.[151,1437,151,538]" lastPageId="6" lastPageNumber="413" pageId="5" pageNumber="412">
<emphasis id="828FB82AFFC03B24FF3EF85BB01AA0EE" bold="true" box="[199,322,1971,1997]" pageId="5" pageNumber="412">Diagnosis.</emphasis>
The new species is characterised by the scapus being dorsally black but ventrally and laterally brown, and the antennae being wholly black; the head has a distinct black band behind eyes; and the male phallic complex has the medial valve elongated, not curved to a spiral, sinuous both dorso-ventrally and laterally and with the ventro-external sclerite of the lateral valve being elongated, in basal third broader, and tapering into a narrow lamella with subacute apex.
</paragraph>
<paragraph id="B0446438FFC33B27FF3EFEEBB421A6AA" blockId="6.[151,1437,151,538]" pageId="6" pageNumber="413">
The new species is closest in distribution to
<taxonomicName id="77FB1FBBFFC33B27FD48FEEBB553A63E" authority="Chopard, 1969" authorityName="Chopard" authorityYear="1969" box="[689,1035,259,285]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="828FB82AFFC33B27FD48FEEBB206A63E" box="[689,862,259,285]" italics="true" pageId="6" pageNumber="413">E. angusticollis</emphasis>
<bibRefCitation id="D46A19C9FFC33B27FC9CFEEBB553A63E" author="Chopard, L." box="[869,1035,259,285]" pageId="6" pageNumber="413" refId="ref4785" refString="Chopard, L. (1969) Family Gryllidae: Subfamilies Mogoplistinae, Myrecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Family Oecanthidae, Gryllotalpidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Vol. 12. Uitgeverij Dr. W. Junk N. V. ' s, Gravenhage, pp. 215 - 500." type="book" year="1969">Chopard, 1969</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="C8EC24A8FFC33B27FBB7FEEBB59BA63E" box="[1102,1219,259,285]" name="Singapore" pageId="6" pageNumber="413">Singapore</collectingCountry>
. In both species the pronotum is more strongly widening apicad, and fore and middle tibiae are provided with black dorsal stripes. The new species differs by the scapus and antennae being black instead of testaceous, by the head being provided with a distinct black band behind eyes and by the male phallic complex having the medial valve not curved to a spiral.
</paragraph>
<paragraph id="B0446438FFC33B27FF3EFE7BB5D0A53A" blockId="6.[151,1437,151,538]" pageId="6" pageNumber="413">
The new species is most similar to
<taxonomicName id="77FB1FBBFFC33B27FDA1FE7BB2C3A68E" authority="Ingrisch, 2006" authorityName="Ingrisch" authorityYear="2006" box="[600,923,403,429]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="argentatus">
<emphasis id="828FB82AFFC33B27FDA1FE7BB3A8A68F" box="[600,752,403,428]" italics="true" pageId="6" pageNumber="413">E. argentatus</emphasis>
<bibRefCitation id="D46A19C9FFC33B27FD01FE7BB2C3A68E" author="Ingrisch, S." box="[760,923,403,429]" pageId="6" pageNumber="413" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="C8EC24A8FFC33B27FC18FE7BB51DA68E" box="[993,1093,403,429]" name="Thailand" pageId="6" pageNumber="413">Thailand</collectingCountry>
having the pronotum strongly widening apicad and the frons dark; it differs by the darked scapus and basal antennal segments instead of being yellow brown, by the male phallic complex which is more elongated and having the medial valve sinuous rather than curving upward (basally) and by the shape of the apical sclerotization of the medial valve.
</paragraph>
<caption id="E48434B0FFC33B27FF6EF896B0F2A0FD" ID-DOI="http://doi.org/10.5281/zenodo.5556505" ID-Zenodo-Dep="5556505" httpUri="https://zenodo.org/record/5556505/files/figure.png" pageId="6" pageNumber="413" startId="6.[151,250,1918,1942]" targetBox="[154,1434,564,1884]" targetPageId="6">
<paragraph id="B0446438FFC33B27FF6EF896B0F2A0FD" blockId="6.[151,1436,1918,2014]" pageId="6" pageNumber="413">
<emphasis id="828FB82AFFC33B27FF6EF896B04AA0B5" bold="true" box="[151,274,1918,1942]" pageId="6" pageNumber="413">FIGURE 4.</emphasis>
<taxonomicName id="77FB1FBBFFC33B27FEE1F896B3B6A0B5" authority="Tan, Japir &amp; Chung" authorityName="Tan, Japir &amp; Chung" authorityYear="2021" box="[280,750,1918,1942]" class="Insecta" family="Mogoplistidae" genus="Cycloptiloides" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="413" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="828FB82AFFC33B27FEE1F896B379A0B5" box="[280,545,1918,1942]" italics="true" pageId="6" pageNumber="413">Cycloptiloides bimaculata</emphasis>
Tan, Japir &amp; Chung
</taxonomicName>
,
<emphasis id="828FB82AFFC33B27FD01F896B21FA0B5" bold="true" box="[760,839,1918,1942]" pageId="6" pageNumber="413">
<taxonomicNameLabel id="99BC0551FFC33B27FD01F896B21FA0B5" box="[760,839,1918,1942]" pageId="6" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype: head (A) and inner tympanum (B) in lateral view; male abdominal apex in dorsal (C in ethanol, D from live photograph), lateral (E) and ventral (F) views. Scale bars: 1 mm (all except B), 0.5 mm (B).
</paragraph>
</caption>
<caption id="E48434B0FFC23B26FF6EFBCDB5A9A31E" ID-DOI="http://doi.org/10.5281/zenodo.5556507" ID-Zenodo-Dep="5556507" box="[151,1265,1061,1085]" httpUri="https://zenodo.org/record/5556507/files/figure.png" pageId="7" pageNumber="414" startId="7.[151,250,1061,1085]" targetBox="[151,1436,181,1037]" targetPageId="7">
<paragraph id="B0446438FFC23B26FF6EFBCDB5A9A31E" blockId="7.[151,1265,1061,1085]" box="[151,1265,1061,1085]" pageId="7" pageNumber="414">
<emphasis id="828FB82AFFC23B26FF6EFBCDB04BA31E" bold="true" box="[151,275,1061,1085]" pageId="7" pageNumber="414">FIGURE 5.</emphasis>
<taxonomicName id="77FB1FBBFFC23B26FEE0FBCDB3E6A31E" authority="Tan &amp; Wahab" authorityName="Tan &amp; Wahab" authorityYear="2021" box="[281,702,1061,1085]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="414" phylum="Arthropoda" rank="species" species="nigrofasciatus" status="sp. nov.">
<emphasis id="828FB82AFFC23B26FEE0FBCDB372A31E" box="[281,554,1061,1085]" italics="true" pageId="7" pageNumber="414">Ectatoderus nigrofasciatus</emphasis>
Tan &amp; Wahab
</taxonomicName>
,
<emphasis id="828FB82AFFC23B26FD31FBCDB240A31E" bold="true" box="[712,792,1061,1085]" pageId="7" pageNumber="414">
<taxonomicNameLabel id="99BC0551FFC23B26FD31FBCDB240A31E" box="[712,792,1061,1085]" pageId="7" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype on a palm/ rattan leaf in Belait.
</paragraph>
</caption>
<paragraph id="B0446438FFC23B26FF3EFB83B433A3EE" blockId="7.[151,1437,1131,1986]" pageId="7" pageNumber="414">
This new species also differs from
<taxonomicName id="77FB1FBBFFC23B26FDB3FB84B20AA3A6" authority="Ingrisch, 2006" authorityName="Ingrisch" authorityYear="2006" box="[586,850,1131,1157]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="414" phylum="Arthropoda" rank="species" species="samui">
<emphasis id="828FB82AFFC23B26FDB3FB84B3F3A3A6" box="[586,683,1132,1157]" italics="true" pageId="7" pageNumber="414">E. samui</emphasis>
<bibRefCitation id="D46A19C9FFC23B26FD48FB83B20AA3A6" author="Ingrisch, S." box="[689,850,1131,1157]" pageId="7" pageNumber="414" pagination="133 - 227" refId="ref4931" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="C8EC24A8FFC23B26FC6CFB83B2A1A3A6" box="[917,1017,1131,1157]" name="Thailand" pageId="7" pageNumber="414">Thailand</collectingCountry>
by the pronotum being more strongly widening apicad and by the male phallic complex having the medial valve not curved to a spiral; from
<taxonomicName id="77FB1FBBFFC23B26FAF9FB78B03DA3EE" authority="Bey-Bienko, 1966" authorityName="Bey-Bienko" authorityYear="1966" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="414" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="828FB82AFFC23B26FAF9FB78B4C5A38A" box="[1280,1437,1168,1193]" italics="true" pageId="7" pageNumber="414">E. marginatus</emphasis>
Bey-Bienko, 1966
</taxonomicName>
from Lesser Sunda Islands it differs by the pronotum being more strongly widening apicad.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC23B26FF3EFB30B26FA236" pageId="7" pageNumber="414" type="etymology">
<paragraph id="B0446438FFC23B26FF3EFB30B26FA236" blockId="7.[151,1437,1131,1986]" pageId="7" pageNumber="414">
<emphasis id="828FB82AFFC23B26FF3EFB30B016A3D1" bold="true" box="[199,334,1240,1266]" pageId="7" pageNumber="414">Etymology.</emphasis>
The species name refers to the black band posterior of eyes and the black scapus (dorsally only) and antennae; nigro = black in Latin, fascia = band in Latin.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFC23B28FF3EFAF7B3BCA68E" lastPageId="9" lastPageNumber="416" pageId="7" pageNumber="414" type="description">
<paragraph id="B0446438FFC23B26FF3EFAF7B23DA131" blockId="7.[151,1437,1131,1986]" pageId="7" pageNumber="414">
<emphasis id="828FB82AFFC23B26FF3EFAF7B001A21A" bold="true" box="[199,345,1311,1337]" pageId="7" pageNumber="414">Description.</emphasis>
Habitus of male as shown in
<figureCitation id="28C078BDFFC23B26FD5AFAF7B3BEA219" box="[675,742,1311,1338]" captionStart="FIGURE 5" captionStartId="7.[151,250,1061,1085]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure-19@7.[151,1436,181,1037]" captionTargetPageId="7" captionText="FIGURE 5. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male holotype on a palm/ rattan leaf in Belait." figureDoi="http://doi.org/10.5281/zenodo.5556507" httpUri="https://zenodo.org/record/5556507/files/figure.png" pageId="7" pageNumber="414">Fig. 5</figureCitation>
. Dorsum of head flattened (
<figureCitation id="28C078BDFFC23B26FBE4FAF7B5E8A219" box="[1053,1200,1311,1338]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Figs. 6A, 6B</figureCitation>
). Frontal rostrum 1.3 times wider than scapus (
<figureCitation id="28C078BDFFC23B26FE4FFAABB349A27D" box="[438,529,1347,1374]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Fig. 6A</figureCitation>
). Maxillary palps with apical (= fifth) and subapical (= fourth) segments of subequal length, third segment longer than apical and subapical segments; apical segment triangular and widened apically (
<figureCitation id="28C078BDFFC23B26FF24FA63B06EA285" box="[221,310,1419,1446]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Fig. 6B</figureCitation>
). Pronotum about 1.7 times longer than wide with anterior dorsal margin narrow and straight; lateral margin widening posteriorly; strongly produced posteriorly and covering near-entirely tegmen; posterior margin convex. Fore tibia with internal tympanum small and oval; without external tympanum. Hind femur 1.1 times longer than hind tibia; hind tibia 4.4 times longer than hind metatarsus.
</paragraph>
<paragraph id="B0446438FFC23B26FF3EF9F3B2F5A1C9" blockId="7.[151,1437,1131,1986]" pageId="7" pageNumber="414">
Male. Supra-anal plate with last abdominal tergite and epiproct distinctly separated by a transverse suture; last abdominal tergite transverse, with hind margin emarginate; supra-anal plate less transverse than last abdominal tergite, flattened, with fewer setae, apex rounded (
<figureCitation id="28C078BDFFC23B26FD37F98BB27EA15D" box="[718,806,1635,1662]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Fig. 6E</figureCitation>
). Genitalia as in
<figureCitation id="28C078BDFFC23B26FC1FF98BB573A15D" box="[998,1067,1635,1662]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@10.[179,1408,181,1920]" captionTargetPageId="10" captionText="FIGURE 7. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male phallic complex in dorsal (A), ventral (B), lateral (C) and latero-ventral (D) views; and apex of medial valve in ventral view (E). Scale bars: 1 mm (A—D), 0.5 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.5556511" httpUri="https://zenodo.org/record/5556511/files/figure.png" pageId="7" pageNumber="414">Fig. 7</figureCitation>
. Epiphallus elongated and membranous. Medial valve elongated, not curved to a spiral, sinuous both dorso-ventrally and laterally. Lateral valves weakly sclerotised, forming stout and truncated lobes at apex. Ventro-external sclerite of lateral valve elongate, basal third broader, tapering into a narrow lamella with subacute apex.
</paragraph>
<paragraph id="B0446438FFC23B26FF3EF91BB0C6A02E" blockId="7.[151,1437,1131,1986]" box="[199,414,1779,1805]" pageId="7" pageNumber="414">Female. Unknown.</paragraph>
<paragraph id="B0446438FFC23B28FF3EF8FFB5A2A646" blockId="7.[151,1437,1131,1986]" lastBlockId="9.[151,1437,151,466]" lastPageId="9" lastPageNumber="416" pageId="7" pageNumber="414">
Colouration. Head dorsum light brown, with white stripe running along lateral margin from posterior end, along inner margin of eyes to frontal rostrum (
<figureCitation id="28C078BDFFC23B26FDA8F8D3B3BBA075" box="[593,739,1851,1878]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Figs. 6B, 6D</figureCitation>
). Scapus dorsally black, otherwise brown (
<figureCitation id="28C078BDFFC23B26FB45F8D3B417A075" box="[1212,1359,1851,1878]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Figs. 6A, 6D</figureCitation>
); basal antennal segments black, thereafter brown with some segments black (
<figureCitation id="28C078BDFFC23B26FC45F8B7B50DA059" box="[956,1109,1887,1914]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Figs. 6A, 6D</figureCitation>
). Frons and mouthparts dark coloured (
<figureCitation id="28C078BDFFC23B26FEF1F86BB039A0BD" box="[264,353,1923,1950]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Fig. 6C</figureCitation>
). Maxillary and labial palps yellow brown (
<figureCitation id="28C078BDFFC23B26FCB3F86BB283A0BD" box="[842,987,1923,1950]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="7" pageNumber="414">Figs. 6B, 6C</figureCitation>
). Lateral parts of head, including genae, light brown; with a black horizontal band behind eyes. Pronotal dorsal disc brown, apical margin with white scales. Pronotal lateral lobes also with white scales and with brown margins (
<figureCitation id="28C078BDFFCC3B28FC63FF7FB2ABA791" box="[922,1011,151,178]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="9" pageNumber="416">Fig. 6D</figureCitation>
). Tegmen mostly infumated white, but infumated grey at apical area of lateral field (
<figureCitation id="28C078BDFFCC3B28FD71FF53B3B9A7F5" box="[648,737,187,214]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="9" pageNumber="416">Fig. 6D</figureCitation>
). Femur generally pale brown to brown. Fore and middle tibiae dorsally black, otherwise brown. Tarsi with basal segment brown, with middle and apical segments black. Hind femur generally brown with knee dark. Hind tibia brown, dorsally pale yellow. Abdomen with tergites black covered with greyish brown scales; sternites black with whitish scales especially on those nearer to thorax. Abdominal apex black; last abdominal tergite back with brown setae along posterior margin (
<figureCitation id="28C078BDFFCC3B28FC15FEA3B51BA645" box="[1004,1091,331,358]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[184,1414,201,1920]" captionTargetId="figure-20@8.[171,1417,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Ectatoderus nigrofasciatus Tan &amp; Wahab, sp. nov. male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="9" pageNumber="416">Fig. 6E</figureCitation>
). Cercus brown.
</paragraph>
<caption id="E48434B0FFCD3B29FF6EF87FB53FA0F0" ID-DOI="http://doi.org/10.5281/zenodo.5556509" ID-Zenodo-Dep="5556509" httpUri="https://zenodo.org/record/5556509/files/figure.png" pageId="8" pageNumber="415" startId="8.[151,250,1943,1967]" targetBox="[184,1414,201,1920]" targetPageId="8">
<paragraph id="B0446438FFCD3B29FF6EF87FB53FA0F0" blockId="8.[151,1436,1943,2004]" pageId="8" pageNumber="415">
<emphasis id="828FB82AFFCD3B29FF6EF87FB04CA08C" bold="true" box="[151,276,1943,1967]" pageId="8" pageNumber="415">FIGURE 6.</emphasis>
<taxonomicName id="77FB1FBBFFCD3B29FEE2F870B39BA093" authority="Tan &amp; Wahab" authorityName="Tan &amp; Wahab" authorityYear="2021" box="[283,707,1944,1968]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="415" phylum="Arthropoda" rank="species" species="nigrofasciatus" status="sp. nov.">
<emphasis id="828FB82AFFCD3B29FEE2F870B375A093" box="[283,557,1944,1968]" italics="true" pageId="8" pageNumber="415">Ectatoderus nigrofasciatus</emphasis>
Tan &amp; Wahab
</taxonomicName>
,
<emphasis id="828FB82AFFCD3B29FD37F870B278A093" bold="true" box="[718,800,1944,1968]" pageId="8" pageNumber="415">
<taxonomicNameLabel id="99BC0551FFCD3B29FD37F870B278A093" box="[718,800,1944,1968]" pageId="8" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male: head and pronotum in dorsal (A) and lateral (D) views, head in lateral (B) and anterior (C) views; abdominal apex in dorsal view (E). Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="B0446438FFCC3B28FF3EFE87B3BCA68E" blockId="9.[151,1437,151,466]" pageId="9" pageNumber="416">
<emphasis id="828FB82AFFCC3B28FF3EFE87B0BDA6AB" bold="true" box="[199,485,366,393]" pageId="9" pageNumber="416">Measurements (in mm).</emphasis>
<typeStatus id="6F40DA9AFFCC3B28FE15FE87B30CA6AA" box="[492,596,367,393]" pageId="9" pageNumber="416" type="holotype">Holotype</typeStatus>
BL = 7.6; FRW = 0.28; SW = 0.22; EW = 0.26; PL = 3.9; PW = 2.2; TL = 2.3; HFL = 4.3; HFW = 1.3; HTL = 4.0; HML = 0.9.
</paragraph>
</subSubSection>
<subSubSection id="F8E137B3FFCC3B28FF3EFE5EB3F2A6F1" box="[199,682,438,466]" pageId="9" pageNumber="416" type="distribution">
<paragraph id="B0446438FFCC3B28FF3EFE5EB3F2A6F1" blockId="9.[151,1437,151,466]" box="[199,682,438,466]" pageId="9" pageNumber="416">
<emphasis id="828FB82AFFCC3B28FF3EFE5EB039A6F3" bold="true" box="[199,353,438,464]" pageId="9" pageNumber="416">Distribution.</emphasis>
Borneo (
<collectingCountry id="C8EC24A8FFCC3B28FE31FE5FB3FBA6F1" box="[456,675,439,466]" name="Brunei" pageId="9" pageNumber="416">Brunei Darussalam</collectingCountry>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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