122 lines
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122 lines
16 KiB
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<document id="E4FD510003D974B8AAEF420816CE43C8" ID-CLB-Dataset="25728" ID-DOI="http://dx.doi.org/10.3897/zookeys.66.683" ID-GBIF-Dataset="a164e717-82d3-4b7b-abb9-2e202a7a1597" ID-PMC="PMC3088416" ID-Pensoft-Pub="1313-2970-66-29" ID-PubMed="21594030" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-66-29" ModsDocOrigin="ZooKeys 66" ModsDocTitle="Annotated checklist of the recent and extinct pythons (Serpentes, Pythonidae), with notes on nomenclature, taxonomy, and distribution" checkinTime="1451250909259" checkinUser="pensoft" docAuthor="Schleip, Wulf D. & O'Shea, Mark" docDate="2010" docId="382E501E628E152A69EC5BFFD22F0415" docLanguage="en" docName="ZooKeys 66: 29-79" docOrigin="ZooKeys 66" docSource="http://dx.doi.org/10.3897/zookeys.66.683" docTitle="Python molurus subsp. pimbura Deraniyagala 1945, [subspecies inquirenda, APP7]" docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="FFFFFFC1FFA4551A4D30CF25AE54FFB9" masterDocTitle="Annotated checklist of the recent and extinct pythons (Serpentes, Pythonidae), with notes on nomenclature, taxonomy, and distribution" masterLastPageNumber="80" masterPageNumber="29" pageNumber="52" updateTime="1691877344321" updateUser="ExternalLinkService">
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<mods:title id="E0CB25DCE85DF1FCF44EB7A45D05BC97">Annotated checklist of the recent and extinct pythons (Serpentes, Pythonidae), with notes on nomenclature, taxonomy, and distribution</mods:title>
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<mods:namePart id="372344CCE2BAB521BD312BEA9C075B21">Schleip, Wulf D.</mods:namePart>
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<mods:namePart id="3F17CBE9EFE9743EA420E7EC370A1A75">O'Shea, Mark</mods:namePart>
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<mods:date id="8CCBB5DE9C7913DAE91B18320177C0B1">2010</mods:date>
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<mods:number id="4553445B5F1663A4B6FE5F91BEE3D045">66</mods:number>
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<mods:start id="0494B83B1641D563CF3F3D8BF5FCC21F">29</mods:start>
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<mods:url id="FD5774166AA360D2A178DA9723618FA3">http://dx.doi.org/10.3897/zookeys.66.683</mods:url>
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<mods:classification id="B90867F64D3C6D24EDD8A701464F2DA8">journal article</mods:classification>
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<mods:identifier id="7B424BEFEDF03328CF498298EFFEF2A6" type="DOI">http://dx.doi.org/10.3897/zookeys.66.683</mods:identifier>
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<treatment id="382E501E628E152A69EC5BFFD22F0415" ID-GBIF-Taxon="159360591" LSID="urn:lsid:plazi:treatment:382E501E628E152A69EC5BFFD22F0415" httpUri="http://treatment.plazi.org/id/382E501E628E152A69EC5BFFD22F0415" lastPageNumber="52" pageId="23" pageNumber="52">
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<subSubSection id="2BD17769EEBDF6B6E5C6E6743B16D5EE" pageId="23" pageNumber="52" type="nomenclature">
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<paragraph id="73D6D6CEDEAB6A761FBD58ED0DD25405" pageId="23" pageNumber="52">
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<taxonomicName id="357598C3231C80C0A2E0ABEE37324058" ID-CoL="5KTG2" authority="Deraniyagala, 1945" authorityName="Deraniyagala" authorityYear="1945" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="pimbura">Python molurus pimbura (Deraniyagala, 1945)</taxonomicName>
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<taxonomicNameLabel id="8942D4B6C5A836769796E635221D9030" pageId="23" pageNumber="52">[subspecies inquirenda, APP7]</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="70807988C9DB904CFF71BCED5AFEEC03" pageId="23" pageNumber="52" type="synonyms">
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<paragraph id="3B269287F395F461A84D41DC86CE5619" pageId="23" pageNumber="52">Synonyms:</paragraph>
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<paragraph id="97B3F3ECFF5158B9AEFEBBD2823644E3" pageId="23" pageNumber="52">
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<taxonomicName id="7E06732A4D8132623E479EB77C1A0036" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="molurus">Python molurus molurus</taxonomicName>
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(Linnaeus) (part)
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</paragraph>
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<paragraph id="5D13E50B7089A1E71C685E15C7143ECD" pageId="23" pageNumber="52">
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<taxonomicName id="699827EB76F0D910042C3A3539A1538F" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="molurus">Python molurus molurus</taxonomicName>
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-
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<bibRefCitation id="32B466FD66CAB6F98E5DD4D931FD87B6" author="Constable, JD" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, Harvard College" pageId="24" pageNumber="53" pagination="60 - 160" title="Reptiles from the Indian Peninsula in the Museum of Comparative Zooelogy." volume="103" year="1949">Constable 1949</bibRefCitation>
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</paragraph>
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<paragraph id="E18603AC9F4ED77FB0B99A7D92BB7EBC" pageId="23" pageNumber="52">
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<taxonomicName id="C8B0D20B15A992FD17534D374D0CC12C" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. pimbura" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="pimbura">Python molurus pimbura</taxonomicName>
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-
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<bibRefCitation id="28D7955CA1FBAB5CDD6A3E284DBCCB38" author="Deraniyagala, PEP" journalOrPublisher="Government Press, Colombo, Sri Lanka" pageId="24" pageNumber="53" title="Python molurus" volumeTitle="A Colored Atlas of some vertebrates from Ceylon, Vol. 3, Serpentoid Reptilia." year="1955">Deraniyagala 1955</bibRefCitation>
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</paragraph>
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<paragraph id="4F14E9A8D193A3826AB6027CE1E23C30" pageId="23" pageNumber="52">
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<taxonomicName id="72580771F356FE047A25B102E84203BF" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="molurus">Python molurus molurus</taxonomicName>
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-
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<bibRefCitation id="F4C80E6D535BE0398F10F55D646BAA0D" author="Stimson, AF" journalOrPublisher="Das Tierreich" pageId="25" pageNumber="54" pagination="1 - 49" title="Liste der rezenten Amphibien und Reptilien. Boidae (Boinae + Bolyeriinae + Loxoceminae + Pythoninae)." volume="89" year="1969">Stimson 1969</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EE08C463DAC4D69E5DE385BC487D0FA5" pageId="23" pageNumber="52" type="distribution">
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<paragraph id="237BF51188BFB78AED34C8368AAE4238" pageId="23" pageNumber="52">Distribution:</paragraph>
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<paragraph id="D37EE02879BFE1387459B4ABDB7A4332" pageId="23" pageNumber="52">
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First reported from Nunavil (Thenmarachi), Jaffna Peninsula, Sri Lanka by
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<bibRefCitation id="F88B626371F2502C3C664044918FDD24" author="Abyerami, S" journalOrPublisher="Pakistan Journal of Biological Sciences" pageId="24" pageNumber="53" pagination="1969 - 1978" title="Diversity of snakes from the Jaffna Peninsula, Sri Lanka." volume="11" year="2008">Abyerami and Sivashanthini (2008)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E959C6A91FDB4ADE08A5DF80A06AE4FE" pageId="23" pageNumber="52" type="remarks">
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<paragraph id="7AEE6C478C2C9FC7CB86EBDAAEA28421" pageId="23" pageNumber="52">Remarks:</paragraph>
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<paragraph id="09A4AF406BD1B2B6ACF6FA00A0F16845" pageId="23" pageNumber="52">
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<bibRefCitation id="0E21BFAC492CD685A73A6DA02EECF630" author="Hoser, RT" journalOrPublisher="Crocodilian - Journal of the Victorian Association of Amateur Herpetologists 4 (3): 31 - 37 and" pageId="24" pageNumber="53" pagination="21 - 40" title="A reclassification of the Pythoninae including the description of two new genera, two new species and nine new subspecies." volume="4" year="2004">Hoser (2004)</bibRefCitation>
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resurrected this taxon from the synonymy of
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<taxonomicName id="23455531B98BB0173F34D7C4566DDC46" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="molurus">Python molurus molurus</taxonomicName>
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without providing reasons for this action.
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<bibRefCitation id="A20C6B8913CA9FB21D1BECCBC2B5A1E5" author="Deraniyagala, PEP" journalOrPublisher="Spolia Zeylanica," pageId="24" pageNumber="53" pagination="103 - 113" title="Some new races of the Python, Chrysopelea, Binocellate Cobra and Tith-Polonga inhabiting Ceylon and India." volume="24" year="1945">Deraniyagala (1945)</bibRefCitation>
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separated the subspecies from
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<taxonomicName id="386375235D2F6DA78AAF1EEE7C40109D" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus subsp. molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="subSpecies" species="molurus" subSpecies="molurus">Python molurus molurus</taxonomicName>
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based on lower subcaudal scale counts and the irregular shape of the lateral markings. Dorsal midbody scale rows and ventral scale counts overlap those of the nominate subspecies.
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<bibRefCitation id="1FF0B0D56179B7541BD92A59764D9439" author="Constable, JD" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, Harvard College" pageId="24" pageNumber="53" pagination="60 - 160" title="Reptiles from the Indian Peninsula in the Museum of Comparative Zooelogy." volume="103" year="1949">Constable (1949: 124)</bibRefCitation>
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did not follow this placement and synonymized this taxon with the nominate subspecies, which was followed by
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<bibRefCitation id="D9D28F9D2B23D97306C41FE2CF1BEE9A" author="Stimson, AF" journalOrPublisher="Das Tierreich" pageId="25" pageNumber="54" pagination="1 - 49" title="Liste der rezenten Amphibien und Reptilien. Boidae (Boinae + Bolyeriinae + Loxoceminae + Pythoninae)." volume="89" year="1969">Stimson (1969)</bibRefCitation>
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. A second paper by
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<bibRefCitation id="944F578150D854C4F069BB47D7D8DC69" author="Deraniyagala, PEP" journalOrPublisher="Government Press, Colombo, Sri Lanka" pageId="24" pageNumber="53" title="Python molurus" volumeTitle="A Colored Atlas of some vertebrates from Ceylon, Vol. 3, Serpentoid Reptilia." year="1955">Deraniyagala (1955: 6)</bibRefCitation>
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provided a more detailed description of the subspecies. Therein, he stated that this taxon is also separated from the nominate form "in generally possessing three preoculars instead of two" or four as stated by
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<bibRefCitation id="9F7D707BCD1FEA08328B12073946879C" author="Wall, F" journalOrPublisher="HR Cottle, Colombo" pageId="26" pageNumber="55" title="Ophidia Taprobanica or the Snakes of Ceylon." year="1921">Wall (1921: 47)</bibRefCitation>
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for some Indian populations of the nominate form. There appears to be a range in preocular scale counts across India, from three in the northeast, to four in the north-center, and two in northwest (
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<normalizedToken id="24FECB734E0DCD8A3EA8113A2C1E01D2" originalValue="O’Shea">O'Shea</normalizedToken>
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pers. obs.) but this data, from only a few specimens, requires further verification. Contrary to his findings in 1945,
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<bibRefCitation id="95D808765537EA4BE02EE7CECA16E4CF" author="Deraniyagala, PEP" journalOrPublisher="Government Press, Colombo, Sri Lanka" pageId="24" pageNumber="53" title="Python molurus" volumeTitle="A Colored Atlas of some vertebrates from Ceylon, Vol. 3, Serpentoid Reptilia." year="1955">Deraniyagala (1955)</bibRefCitation>
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reports this taxon to have "more subcaudals" than the nominate form, obviously a typographic error according to the scale count data provided therein. It seems likely that subsequent workers overlooked this latter work, since neither
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<bibRefCitation id="1DD0FB6C642F788E0D7E3577EBEE586E" author="Stimson, AF" journalOrPublisher="Das Tierreich" pageId="25" pageNumber="54" pagination="1 - 49" title="Liste der rezenten Amphibien und Reptilien. Boidae (Boinae + Bolyeriinae + Loxoceminae + Pythoninae)." volume="89" year="1969">Stimson (1969)</bibRefCitation>
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nor
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<bibRefCitation id="694D2C896F24B9B92026AF9D842F40BA" author="McDiarmid, RW" journalOrPublisher="The Herpetologist's League, Washington" pageId="25" pageNumber="54" title="Snake species of the world: a taxonomic and geographic reference. Volume 1" year="1999">McDiarmid et al. (1999)</bibRefCitation>
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or
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<bibRefCitation id="D821BD6CF7C42FE52D2559C3E569419C" author="Henderson, RW" editor="Henderson, RW" journalOrPublisher="Eagle Mountain Publishing, Utah, USA" pageId="24" pageNumber="53" pagination="3 - 21" title="The biology of boas and pythons: a retrospective look to the future." volumeTitle="Biology of the Boas and Pythons." year="2007">Henderson and Powell (2007)</bibRefCitation>
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cited it. Several subsequent workers accepted the placement to the synonymy of the nominate form, but no further studies have been conducted on the molurus-complex. However, besides the lower subcaudal scale counts and the higher number of preoculars, the pink surface of the head may also constitite a morphological difference. (
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<bibRefCitation id="6273D07971F4BDFFA90D73113E7F6F4C" author="Boulenger, GA" journalOrPublisher="Taylor and Francis, London, U. K." pageId="24" pageNumber="53" title="Reptilia and Batrachia: The Fauna of British India, including Ceylon and Burma." year="1890">Boulenger (1890</bibRefCitation>
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,
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<bibRefCitation id="B01BC33C31B910B8EFBE49A56487B4D1" author="Boulenger, GA" journalOrPublisher="British Museum (Natural History), London, U. K." pageId="24" pageNumber="53" title="Catalogue of the snakes in the British Museum (Natural History). Vol. I" year="1893">1893</bibRefCitation>
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) and MA
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<bibRefCitation id="01F60DE6D64B69CC688EC1730C5A134F" author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="25" pageNumber="54" title="The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptilia and Amphibia. Vol. III Serpentes." year="1943">Smith (1943)</bibRefCitation>
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recorded two preoculars for
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<taxonomicName id="522834DBE7610A9D4BEA9BC0C8D6972F" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python molurus" pageId="23" pageNumber="52" phylum="Chordata" rank="species" species="molurus">Python molurus</taxonomicName>
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, while
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<bibRefCitation id="5058794F99ABBB244F68CC2C62DA2014" author="Wall, F" journalOrPublisher="HR Cottle, Colombo" pageId="26" pageNumber="55" title="Ophidia Taprobanica or the Snakes of Ceylon." year="1921">Wall (1921)</bibRefCitation>
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records three preoculars for specimens from Ceylon. Since Sri Lanka is a known biodiversity hot spot with a high level of endemism, this allopatric population may represent a cryptic species. Because of the evidence provided by
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<bibRefCitation id="1EDB3604EF8D2A2EBD5368545FF058AA" author="Deraniyagala, PEP" journalOrPublisher="Government Press, Colombo, Sri Lanka" pageId="24" pageNumber="53" title="Python molurus" volumeTitle="A Colored Atlas of some vertebrates from Ceylon, Vol. 3, Serpentoid Reptilia." year="1955">Deraniyagala (1955)</bibRefCitation>
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, these authors tentatively list this taxon as a valid subspecies and call for further research regarding its true status (APP7).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |