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<document ID-DOI="10.25221/fee.365.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164252" approvalRequired="244" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="237" approvalRequired_for_treatments="1" checkinTime="1665241599349" checkinUser="felipe" docAuthor="Grichanov, I. Ya." docDate="2018" docId="382287BE1A506C0DFF7A53D4FDD56943" docLanguage="en" docName="FarEastEntomol.365.1-25.pdf" docOrigin="Far Eastern Entomologist 365" docSource="http://dx.doi.org/10.25221/fee.365.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Tenuopus lomholdti Grichanov 2018, sp. n." docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="C41BFFC61A5E6C1DFF99575FFFE0683F" masterDocTitle="A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="15" updateTime="1665268663009" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new subfamily of Dolichopodidae (Diptera) for Tenuopus Curran, 1924 with description of new species from Tropical Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grichanov, I. Ya.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-08-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>365</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.365.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.365.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7164252</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:382287BE1A506C0DFF7A53D4FDD56943" httpUri="http://treatment.plazi.org/id/382287BE1A506C0DFF7A53D4FDD56943" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection box="[227,650,1162,1187]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph blockId="14.[227,650,1162,1216]" box="[227,650,1162,1187]" pageId="14" pageNumber="15">
<heading box="[227,650,1162,1187]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis bold="true" box="[227,650,1162,1187]" pageId="14" pageNumber="15">
<taxonomicName authority="Grichanov" authorityName="Grichanov" authorityYear="2018" box="[227,578,1162,1187]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lomholdti" status="sp. n.">
<emphasis bold="true" box="[227,449,1162,1186]" italics="true" pageId="14" pageNumber="15">Tenuopus lomholdti</emphasis>
Grichanov
</taxonomicName>
,
<taxonomicNameLabel box="[590,650,1163,1187]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,373,1192,1216]" pageId="14" pageNumber="15" type="description">
<paragraph blockId="14.[227,650,1162,1216]" box="[227,373,1192,1216]" pageId="14" pageNumber="15">
<figureCitation box="[227,295,1192,1216]" captionStart="Figs 1–6" captionStartId="6.[265,306,1584,1606]" captionTargetBox="[229,1124,257,1519]" captionTargetId="figure-0@6.[229,1124,257,1520]" captionTargetPageId="6" captionText="Figs 1–6. Tenuopus spp., head: 1 – T. bururiensis sp. n.; 2 – T. gorongosaensis sp. n.; 3 – T. kirkspriggsi sp. n.; 4 – T. kylei sp. n.; 5 – T. lomholdti sp. n.; 6 – T. unicolor (Becker)." figureDoi="http://doi.org/10.5281/zenodo.7166072" httpUri="https://zenodo.org/record/7166072/files/figure.png" pageId="14" pageNumber="15">Figs 5</figureCitation>
,
<figureCitation box="[306,335,1192,1216]" captionStart="Figs 7–14" captionStartId="10.[265,306,1503,1525]" captionTargetBox="[229,1124,257,1481]" captionTargetId="figure-0@10.[229,1124,257,1482]" captionTargetPageId="10" captionText="Figs 7–14. Tenuopus spp. 7 – T. shcherbakovi Grichanov, head in ethanol with everted" figureDoi="http://doi.org/10.5281/zenodo.7166074" httpUri="https://zenodo.org/record/7166074/files/figure.png" pageId="14" pageNumber="15">12</figureCitation>
,
<figureCitation box="[346,373,1192,1216]" captionStart="Figs 19–22" captionStartId="17.[265,306,1423,1445]" captionTargetBox="[229,1124,717,1388]" captionTargetId="figure-199@17.[229,1124,717,1389]" captionTargetPageId="17" captionText="Figs 19–22. Tenuopus spp., epandrial lobes and surstylus, right lateral view: 19 – T. kylei" figureDoi="http://doi.org/10.5281/zenodo.7164266" httpUri="https://zenodo.org/record/7164266/files/figure.png" pageId="14" pageNumber="15">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="materials_examined">
<paragraph blockId="14.[227,1127,1237,1610]" box="[265,1126,1237,1261]" pageId="14" pageNumber="15">
<materialsCitation box="[265,1126,1237,1261]" collectionCode="TYPE, MATERIAL" country="Tanzania" elevation="1000" location="East Uzambara" municipality="Amani" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode box="[265,480,1237,1261]" pageId="14" pageNumber="15">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[491,590,1237,1261]" pageId="14" pageNumber="15">Holotype</typeStatus>
–
<specimenCount box="[619,641,1237,1260]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
,
<emphasis bold="true" box="[653,760,1237,1261]" pageId="14" pageNumber="15">
<collectingCountry box="[653,760,1237,1261]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:382287BE1A506C0DFF7A53D4FDD56943:B55460731A506C13FC9F538AFC4C6CD2" box="[774,940,1237,1261]" country="Tanzania" municipality="Amani" name="East Uzambara" pageId="14" pageNumber="15">East Uzambara</location>
,
<collectingMunicipality box="[952,1027,1237,1261]" pageId="14" pageNumber="15">Amani</collectingMunicipality>
,
<quantity box="[1038,1122,1237,1261]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="1000.0">
<elevation box="[1038,1122,1237,1261]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
</materialsCitation>
</paragraph>
<paragraph blockId="14.[227,1127,1237,1610]" pageId="14" pageNumber="15">
<materialsCitation box="[227,1042,1266,1290]" collectingDate="1977-01-26" collectionCode="ZMUC" collectorName="H. Enghoff &amp; O. Lomholdt &amp; O. Martin" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
<date box="[227,335,1266,1290]" pageId="14" pageNumber="15" value="1977-01-26">
<collectingDate box="[227,335,1266,1290]" pageId="14" pageNumber="15" value="1977-01-26">26.I 1977</collectingDate>
</date>
,
<collectorName box="[348,473,1266,1290]" pageId="14" pageNumber="15">H. Enghoff</collectorName>
,
<collectorName box="[487,627,1266,1290]" pageId="14" pageNumber="15">O. Lomholdt</collectorName>
&amp;
<collectorName box="[667,775,1266,1290]" pageId="14" pageNumber="15">O. Martin</collectorName>
leg. [
<collectionCode box="[839,924,1266,1290]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="15">ZMUC</collectionCode>
].
<typeStatus box="[944,1042,1266,1290]" pageId="14" pageNumber="15">Paratype</typeStatus>
</materialsCitation>
:
<materialsCitation collectorName="Udzungwa Mt. N. P." country="Tanzania" latitude="-7.8375278" location="Mito Mitatu" municipality="Udzungwa Mt." pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro" typeStatus="paratype">
<specimenCount box="[1054,1088,1266,1290]" pageId="14" pageNumber="15" type="male">1♂</specimenCount>
(in ethanol),
<collectingCountry box="[327,427,1295,1319]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
:
<collectingRegion box="[438,544,1296,1320]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="15">Morogoro</collectingRegion>
Reg.,
<collectorName box="[616,840,1295,1319]" pageId="14" pageNumber="15">
<collectingMunicipality box="[616,776,1295,1319]" pageId="14" pageNumber="15">Udzungwa Mt.</collectingMunicipality>
N. P.
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:382287BE1A506C0DFF7A53D4FDD56943:B55460731A506C13FCCC5250FC386D18" box="[853,984,1295,1319]" country="Tanzania" latitude="-7.8375278" municipality="Udzungwa Mt." name="Mito Mitatu" pageId="14" pageNumber="15" stateProvince="Morogoro">Mito Mitatu</location>
,
<geoCoordinate box="[997,1122,1295,1319]" degrees="7" direction="south" minutes="50" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="15.1" value="-7.8375278">7°50'15.1''S</geoCoordinate>
</materialsCitation>
,
</paragraph>
<paragraph blockId="14.[227,1127,1237,1610]" box="[227,1109,1324,1348]" pageId="14" pageNumber="15">
<materialsCitation box="[227,1105,1324,1348]" collectingDate="2013-05-17" collectionCode="T, ZMUC" collectorName="Pape &amp; N. Scharff" elevation="1198" longitude="36.847195" pageId="14" pageNumber="15" specimenCount="1">
<geoCoordinate box="[227,367,1324,1348]" degrees="36" direction="east" minutes="50" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="49.9" value="36.847195">36°50'49.9''E</geoCoordinate>
,
<quantity box="[377,459,1324,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.198" pageId="14" pageNumber="15" unit="m" value="1198.0">
<elevation box="[377,459,1324,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.198" pageId="14" pageNumber="15" unit="m" value="1198.0">1198 m</elevation>
</quantity>
,
<collectingMethod box="[469,601,1324,1348]" pageId="14" pageNumber="15">Malaise trap</collectingMethod>
,
<date box="[611,724,1324,1348]" pageId="14" pageNumber="15" value="2013-05-17">
<collectingDate box="[611,724,1324,1348]" pageId="14" pageNumber="15" value="2013-05-17">17.V 2013</collectingDate>
</date>
,
<collectionCode box="[734,753,1325,1348]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="15">T</collectionCode>
.
<collectorName box="[763,814,1325,1348]" pageId="14" pageNumber="15">Pape</collectorName>
&amp;
<collectorName box="[847,956,1324,1348]" pageId="14" pageNumber="15">N. Scharff</collectorName>
leg. [
<collectionCode box="[1013,1099,1324,1348]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="15">ZMUC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph blockId="14.[227,1127,1237,1610]" pageId="14" pageNumber="15">
DESCRIPTION. Male.
<emphasis box="[526,584,1353,1377]" italics="true" pageId="14" pageNumber="15">Head</emphasis>
: frons black, whitish pollinose; one pair of short postvertical setae, as long as uppermost postocular seta; upper postocular setae black, increasing in length upward; lateral and lower postoculars white; ventral postcranium covered with irregular white hairs; face silvery-white, 7 times as high as wide in middle, nearly as wide as postpedicel; clypeus slightly bulging; antennae as long as height of head, with yellow scape and pedicel and yellow-orange postpedicel; pedicel projected distally on inner side, with a crown of short black setae,
</paragraph>
<paragraph blockId="14.[227,1127,1237,1610]" lastBlockId="15.[227,1127,258,1666]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
one of dorsal setae as long as pedicel; postpedicel rounded, slightly longer than high at base (1.5/1.3); arista-like stylus dorsal, with short hairs; length ratio of scape to pedicel to postpedicel to stylus (1
<superScript attach="left" box="[598,610,258,272]" fontSize="6" pageId="15" pageNumber="16">st</superScript>
and 2
<superScript attach="left" box="[679,696,258,272]" fontSize="6" pageId="15" pageNumber="16">nd</superScript>
segments), 15/17/20/12/92; palpus and proboscis short, yellow, covered with white hairs, proboscis also with a pair of black lateral setae.
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">
<emphasis box="[265,342,354,378]" italics="true" pageId="15" pageNumber="16">Thorax</emphasis>
: pleura dirty yellow; mesonotum brownish orange, with narrow black stripe along acrostichals, wider posteriorly and black on scutellum dorsally; 6
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">
dorsocentral setae with posterior pair shifted laterally; acrostichals irregularly biseriate, short, nearly reaching 5
<superScript attach="left" box="[565,578,441,455]" fontSize="6" pageId="15" pageNumber="16">th</superScript>
dorsocentrals; scutellum with 2 strong bristles and
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,821,477,501]" pageId="15" pageNumber="16">2 short lateral hairs; proepisternum with 2 yellow setae.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">
<emphasis box="[265,316,508,531]" italics="true" pageId="15" pageNumber="16">Legs</emphasis>
yellow; last segments of tarsi brown; fore and mid coxae with black hairs anteriorly and 5-7 black apical bristles of various length; hind coxa with one long black outer bristle above middle; fore femur with dark fine erect ventral hairs, at most half as long as diameter of femur, and 1-2 black posteroventral cilia; fore tibia simple, with 1 anterodorsal and 1 posterodorsal at base, 1-2 dorsals in middle, 1-2
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">apical setae; tarsomeres simple, with rather short ventral semi-erect setulae on all segments, with 1-2 elongate dorsal setae on distal apex of segments 1-4; claws simple; mid femur with very small anterior preapical seta, glabrous ventrally; mid tibia with 3 anterior, 3 posterodorsal, 3 ventral, 3-4 apical seta; basitarsus with few short ventral setae; hind femur without long hairs, with 1 strong anterior preapical seta; hind tibia with 3 anterior, 2 dorsal, 3 posterodorsal, 2-3 apical setae; basitarsus with few short posterior setae. Femur, tibia and tarsomere (from first to fifth) length ratio: fore leg: 44/45/35/19/19/21/12, mid leg: 31/50/35/15/11/6/4, hind leg:</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,439,906,930]" pageId="15" pageNumber="16">28/51/17/15/10/6/3.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[265,1127,937,961]" pageId="15" pageNumber="16">
<emphasis box="[265,321,938,961]" italics="true" pageId="15" pageNumber="16">Wing</emphasis>
greyish, veins brown; subcosta very thin; ratio of part of costa between
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,1127,968,993]" pageId="15" pageNumber="16">
R
<subScript attach="left" box="[245,272,979,993]" fontSize="6" pageId="15" pageNumber="16">2+3</subScript>
and R
<subScript attach="left" box="[354,381,979,993]" fontSize="6" pageId="15" pageNumber="16">4+5</subScript>
to that between R
<subScript attach="left" box="[601,628,979,993]" fontSize="6" pageId="15" pageNumber="16">4+5</subScript>
and M
<subScript attach="left" box="[716,725,979,993]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
, 16/2; M
<subScript attach="left" box="[836,845,979,993]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
with gentle arc to apex,
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,1127,998,1023]" pageId="15" pageNumber="16">
reaching costa right before wing apex; M
<subScript attach="left" box="[676,685,1009,1023]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
present as fold on membrane; crossvein
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">
<emphasis box="[227,288,1029,1053]" italics="true" pageId="15" pageNumber="16">dm-m</emphasis>
straight; ratio of crossvein
<emphasis box="[583,644,1029,1053]" italics="true" pageId="15" pageNumber="16">dm-m</emphasis>
to apical part of M
<subScript attach="left" box="[851,879,1040,1054]" fontSize="6" pageId="15" pageNumber="16">1+2</subScript>
(fork-handle) to apical part of M
<subScript attach="left" box="[330,339,1071,1085]" fontSize="6" pageId="15" pageNumber="16">4</subScript>
, 9/31/23; anal vein foldlike, not reaching wing margin; anal angle obtuse;
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">lower calypter yellow, with black apex and pale setae; halter yellow with orange knob, halter stem thin and long, with dorsal and ventral groups of short hairs distally.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">
<emphasis box="[265,366,1182,1206]" italics="true" pageId="15" pageNumber="16">Abdomen</emphasis>
mostly yellow-orange, black setose; 2
<superScript attach="right" box="[778,795,1177,1191]" fontSize="6" pageId="15" pageNumber="16">nd</superScript>
–6
<superScript attach="left" box="[822,835,1177,1191]" fontSize="6" pageId="15" pageNumber="16">th</superScript>
yellow, with narrow brown or black posterior edging; 8
<superScript attach="left" box="[531,544,1208,1222]" fontSize="6" pageId="15" pageNumber="16">th</superScript>
segment and epandrium entirely yellow; 8
<superScript attach="left" box="[1015,1028,1208,1222]" fontSize="6" pageId="15" pageNumber="16">th</superScript>
segment with dark hairs; cerci and surstyli yellow; cercus covered with dark-yellow hairs and setae; surstylus dark at apex; hypandrium small, simple; phallus thin and simple; epandrial lobe prominent, finger-like, pointed at apex, with 4 ventral setae;
</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">5 epandrial setae of different length between hypandrium and base of epandrial lobe, including 1 thick pedunculate seta; surstylus about as large as cercus,</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,1127,1397,1421]" pageId="15" pageNumber="16">irregularly leaf-like, with small dorsal process at apex, with 2 inner setae at middle;</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" pageId="15" pageNumber="16">cercus about half as long as epandrium, subtriangular, pointed at apex, covered with short hairs dorsally, double row of long setae distally.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[265,1127,1489,1513]" pageId="15" pageNumber="16">MEASUREMENTS (in mm). Body length 5.8; antenna length 1.3; wing length</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,442,1519,1544]" pageId="15" pageNumber="16">4.9; wing width 1.4.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[265,470,1550,1574]" pageId="15" pageNumber="16">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="etymology">
<paragraph blockId="15.[227,1127,258,1666]" box="[265,1127,1581,1605]" pageId="15" pageNumber="16">ETYMOLOGY. The species is named after the Danish entomologist, Dr. O.</paragraph>
<paragraph blockId="15.[227,1127,258,1666]" box="[227,910,1611,1636]" pageId="15" pageNumber="16">
Lomholdt (ZMUC), one of the collectors of the
<typeStatus box="[739,785,1612,1636]" pageId="15" pageNumber="16">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="diagnosis">
<paragraph blockId="16.[227,1127,264,381]" pageId="16" pageNumber="17">
DIAGNOSIS.
<taxonomicName authority="Grichanov, 2018" authorityName="Grichanov" authorityYear="2018" box="[425,559,264,288]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lomholdti" status="sp. n.">
<emphasis box="[425,559,264,288]" italics="true" pageId="16" pageNumber="17">T. lomholdti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[570,631,265,288]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
belongs to a group of species with biseriate acrostichals, being the closest to
<taxonomicName authorityName="Grichanov" authorityYear="2000" box="[602,723,296,319]" class="Insecta" family="Dolichopodidae" genus="Tenuopus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="taitensis">
<emphasis box="[602,723,296,319]" italics="true" pageId="16" pageNumber="17">T. taitensis</emphasis>
</taxonomicName>
due to enlarged surstylus, differing from the latter in male cercus subtriangular, without process, surstylus about as large as cercus (see key above).
</paragraph>
</subSubSection>
</treatment>
</document>
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