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<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak &amp; Sutcharit, Chirasak" docDate="2023" docId="37F4588D4F345A08AE01AC391B2D01C7" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Rhiostoma abletti Thach 2016" docType="treatment" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:title id="E2D3450CEBF6F58BA9FE5A401229B31F">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:name id="490DCA8E42DFEA97CFA9FC92C673E12E" type="personal">
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<mods:namePart id="C308904BD5160E90826713F14C5B693B">Tongkerd, Piyoros</mods:namePart>
<mods:affiliation id="B7B5F3CF3F8920FA7ED2258A3359137E">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="F12684B784C606B247948EFEB02DABDB">Tumpeesuwan, Sakboworn</mods:namePart>
<mods:affiliation id="DE3319CB1E6908C72AAA01AA3E2ED1D2">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:namePart id="44C22B19E89D3F7D770C275984B5A0C3">Inkhavilay, Khamla</mods:namePart>
<mods:affiliation id="292370952862C34794AFBA8BE3B95EBC">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:namePart id="CFFA5B856ED56CE591EA6AC8A44304D0">Prasankok, Pongpun</mods:namePart>
<mods:affiliation id="117F497E581C6E3C99FE34B70D90C503">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="AF0F1611C76B159369DDE20442A1BCD4">Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation id="FA7AB7CC0B7880B57EEB75D344B83FCF">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="BC27946036269E4DAE36A3C6A0786AD3">Panha, Somsak</mods:namePart>
<mods:affiliation id="5D0CFAA38AE851B9D83CBFECBCE8E21D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="B57D28D631FBE4A9EBE41BB1A1FEC028" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="7E6329FF8D9EAB7014E631C25BF4C6A4">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="96888E4E23F3884F8637DEE9E1CA604D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="CCD8FC19AA1DD36145A4C926DBE23EAD" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:date id="B55CF1ACDE9BF2BDEDDA4148411BA02D">2023</mods:date>
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<mods:number id="D1953AE055555D4CB581E8EF9294BE1C">1142</mods:number>
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<treatment id="37F4588D4F345A08AE01AC391B2D01C7" ID-GBIF-Taxon="209102290" LSID="urn:lsid:plazi:treatment:37F4588D4F345A08AE01AC391B2D01C7" httpUri="http://treatment.plazi.org/id/37F4588D4F345A08AE01AC391B2D01C7" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
<subSubSection id="86404FA33262D830DD2A313035B81108" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="A34FF84C16756EAD6070B256125E55AF" pageId="0" pageNumber="1">
16.
<taxonomicName id="CA8046EC7C92F7E4796CD4720B39BF7C" ID-CoL="78TFX" LSID="37F4588D-4F34-5A08-AE01-AC391B2D01C7" authority="Thach, 2016" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma abletti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="abletti">Rhiostoma abletti Thach, 2016</taxonomicName>
</paragraph>
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<subSubSection id="624D3D5AE3CB6632B0071FAE0652C40D" pageId="0" pageNumber="1" type="description">
<paragraph id="BECA3641C870FDF9DB6A2DEB1D5ED71D" pageId="0" pageNumber="1">
<figureCitation id="D15FC99BBD6C5CE8A03965140B41B7C4" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Distribution map of Rhiostoma asiphon - species group." figureDoi="10.3897/zookeys.1142.90097.figure34" httpUri="https://binary.pensoft.net/fig/799892" pageId="0" pageNumber="1">Figs 34</figureCitation>
<figureCitation id="2CBA2E0324CA17A4C69122B51E74F7D4" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">, 37</figureCitation>
</paragraph>
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<subSubSection id="80A0BC7BCCEDDBA9C648CDA8EF30AA58" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="69FB6E6382BB167655E57C1B09B68B96" pageId="0" pageNumber="1">
<taxonomicName id="9F5A20CA7D6196A436E1FCCC5966F60E" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma abletti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="abletti">Rhiostoma abletti</taxonomicName>
Thach, 2016: 37, 38, figs 53, 122-124. Type locality: Northwest of Lai Chau city, on the way going to Paso, Lai Chau Province, North Vietnam.
<bibRefCitation id="36D9625079574EDF6FCDE18DDE0695EF" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
: 5, fig. 1a.
</paragraph>
<paragraph id="6764D246A70AE480BEA32536D89F13D4" pageId="0" pageNumber="1">
<taxonomicName id="9C185E0B38857F39C4AE6434028C64E7" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma christae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="christae">Rhiostoma christae</taxonomicName>
Thach, 2016: 38, figs 51, 130-133. Type locality: Near the road No. 6 to Chieng Ngan, Son La Province, North Vietnam.
<bibRefCitation id="BF40ADC15FCF8AEC6199C1222522C489" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
: 17, fig. 3l. New synonym.
</paragraph>
<paragraph id="11D3BC9B872A5C9573446783C102B105" pageId="0" pageNumber="1">
<taxonomicName id="3442DBA8C73AC96900BD68A0334A7149" authorityName="D. S. Do, T. S. Nguyen &amp; H. L. Do" authorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma ninhbien" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ninhbien">Rhiostoma ninhbien</taxonomicName>
Do, Nguyen &amp; Do, 2020b: 169, 170, fig. 2a-c. Type locality: near Tay Trang international border gate, Na U Commune, Dien Bien District, Dien Bien Province, Vietnam. New synonym.
</paragraph>
<paragraph id="63F40CBDF4244C7F09EF48E8EA0F09B7" pageId="0" pageNumber="1">
<taxonomicName id="B7D4F85AB6BA5CA780EB60217BE5953C" baseAuthorityName="Ancey" baseAuthorityYear="1898" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma marioni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="marioni">Rhiostoma marioni</taxonomicName>
-
<bibRefCitation id="167D30FC1427AAE4BB184545DA6C9387" DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
: 168 (in part), fig. 1e, f (not
<bibRefCitation id="A69947A43E17CAB641F7A6EE5FA8793F" author="Ancey, C-F" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille" pageId="0" pageNumber="1" pagination="125 - 150" refId="B3" refString="Ancey, C-F, 1898. Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region. Bulletin du Musee ďHistoire Naturelle de Marseille 1: 125 - 150" title="Observations sur les mollusques terrestres et fluviatiles recueillies dans I'Indo-Chine et particulaierement au Laos, par M. Henri Counillon, accompagnees de remarques sur ďautres mollusques de cette region." volume="1" year="1898">Ancey 1898</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8080B6A401AFA4043B34E610A1A8C184" pageId="0" pageNumber="1" type="type material">
<paragraph id="DD4CCE10C9118EDCDD1FC682FBB3B97A" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="B1D07FA2A02929D0B6CAB3ECE5C43DF9" pageId="0" pageNumber="1">
<emphasis id="50EBE5451FA64B9A67CC09EF3465FE58" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
NHMUK 20160307 (Fig.
<figureCitation id="A3EA0696AE4E3DEF9F40AC148F3DD689" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37A</figureCitation>
) from Northwest of Lai Chau city, on the way going to Paso, Lai Chau Province, Vietnam.
<emphasis id="37B2608E873713DBADDA681B73A18293" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
NHMUK 20160306 (Fig.
<figureCitation id="0DB33AB1FB390B0C2B2E7568D07F1076" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37B</figureCitation>
) of
<taxonomicName id="4151E01051450B928E0257757D6470E4" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma christae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="christae">
<emphasis id="F5AEC6E953E9BAF4B7CAA8D010045D81" italics="true" pageId="0" pageNumber="1">Rhiostoma christae</emphasis>
</taxonomicName>
Thach, 2016, from Near the road No. 6 to Chieng Ngan, Son La Province (North Vietnam).
</paragraph>
<caption id="4642B6C252FEC7DFF708057FED2E8795" doi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
<paragraph id="ABDCF1DF9CC67C2FEC174F5A3B7552A5" pageId="0" pageNumber="1">
<emphasis id="97F66E4F753D838CD65AF3AB77E52742" bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
Shell of
<taxonomicName id="6B4B5A431D888099D9CFE316FFA05B9C" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma abletti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="abletti">
<emphasis id="D4F2EDFDBD22A13A67DBE75E3687C03E" italics="true" pageId="0" pageNumber="1">Rhiostoma abletti</emphasis>
</taxonomicName>
<emphasis id="7DB64068F79F785F41AA7808472D6FE6" bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype NHMUK 20160307 from Lai Chau Province, Vietnam
<emphasis id="E7267B7012210B0D63C6C068F7ECC33F" bold="true" pageId="0" pageNumber="1">B</emphasis>
holotype of
<taxonomicName id="83EAB8C153C83598C585C96CAE5C4085" authorityName="Thach" authorityYear="2016" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma christae" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="christae">
<emphasis id="41860BB23DAC253E2ACC58EDBBF2055B" italics="true" pageId="0" pageNumber="1">Rhiostoma christae</emphasis>
</taxonomicName>
Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam
<emphasis id="3C1F05430FF90FBAEB7E580147906C85" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos
<emphasis id="3FB62CE87D6820277300E58B1C124CD9" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F3ACEA99A3C7340330D5A6D1E544C86C" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="F7FC470223A8FED5EDF6BAA9884C658F" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="E79C8F600B44B5731A39031B7761494F" pageId="0" pageNumber="1">
<materialsCitation id="7D5995AA862A46BE85FC4ECD8F4EA780" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Laos" county="Meuang Hiam District" location="Meuang Hiam District" municipality="Near Ban Na Puek" specimenCount="1" stateProvince="Houaphanh Province">
<emphasis id="BF56E6A480F54FC67518D0E6C80E2E0F" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="E56DEF882DCE221A28172B52C06A0D13" name="Laos">Laos</collectingCountry>
</emphasis>
: Hot Springs (Vieng Thong),
<collectingCounty id="16E14E3673827530FCC6952F7DDB6981">Meuang Hiam District</collectingCounty>
,
<collectingRegion id="8CA6B15433749E7D2D2C9B97046EB54C" country="Laos" name="Houaphan">Houaphanh Province</collectingRegion>
: CUMZ 10206 (3 shells; Fig.
<figureCitation id="5A737528D92DAAF147433F9DDF0AD091" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37E, F</figureCitation>
).
<collectingMunicipality id="CCB8305E18486421DD6F19B71CD45F07">Near Ban Na Puek</collectingMunicipality>
,
<location id="56A66339B8F349466EFC161B4325008E" LSID="urn:lsid:plazi:treatment:37F4588D4F345A08AE01AC391B2D01C7:56A66339B8F349466EFC161B4325008E" country="Laos" county="Meuang Hiam District" municipality="Near Ban Na Puek" name="Meuang Hiam District" stateProvince="Houaphanh Province">Meuang Hiam District</location>
,
<collectingRegion id="A756F21C53B864C486CBE67C17583873" country="Laos" name="Houaphan">Houaphanh Province</collectingRegion>
: CUMZ 10207 (2 shells; Fig.
<figureCitation id="11C156584D31EBB83FB2BD5BAA1607F2" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37C, D</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D7BA34DF2025D464F8F8DE443C940462" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="02C53F3660493C7B2627A951C7C91FAA" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="54DBBFBA9A6C7BFD6877011953F841C2" pageId="0" pageNumber="1">Shell medium, thin and flattened; detached whorl shorter than apertural width. Shell colour with brownish zigzag patterns. Breathing device with incomplete tube shape and nearly perpendicular to detached whorl.</paragraph>
</subSubSection>
<subSubSection id="B60D2C8FC5EC6715D974A89C53CC1B01" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="85AAF4FA67D86A63678CA94027C55CBC" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="5C16AB036C0273483D108C7FEB90A84C" pageId="0" pageNumber="1">
This species can be distinguished from
<taxonomicName id="040B7FD8A4EDB59D4438802934EC8357" lsidName="R. anceyi" pageId="0" pageNumber="1" rank="species" species="anceyi">
<emphasis id="9581697F7B1192E28C28F105739023D7" italics="true" pageId="0" pageNumber="1">R. anceyi</emphasis>
</taxonomicName>
sp. nov. by having an incomplete tube breathing device that is nearly perpendicular to detached whorl, and translucent periostracum. In contrast, the new species has a tubular breathing device, and thickened and reddish brown periostracum.
</paragraph>
</subSubSection>
<subSubSection id="34D92236BDE8AD358B772875820D27EC" pageId="0" pageNumber="1" type="description">
<paragraph id="76C87B92EF0563DE6870DB166B6F927A" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="081ADE1511CA1B9E5A6CD44BD478160D" pageId="0" pageNumber="1">
<emphasis id="D5D1133DF843792205E0A4C3F4D08172" bold="true" pageId="0" pageNumber="1">
<emphasis id="04AF0486A953ED4CEA9339C8DFDC3FE5" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell medium, width 38.5 mm, height 14.5 mm, thin, and flattened to sub-discoidal shape. Apex acute; spire slightly elevated to flattened. Whorls 5 to 6, convex, increasing regularly; suture wide and depressed; last whorl rounded and stout. Shell surface with fine growth lines. Periostracum thick corneous and translucent. Shell colour with dark brown zigzag pattern; with narrow dark spiral band on periphery. Detached whorl shorter than apertural width. Peristome circular and double; lip thickened and slightly expanded. Aperture opened sub-laterally. Breathing device with incomplete tube shape and nearly perpendicular to detached whorl; outer lip forming a short to long nearly closed tube; inner lip with deep incision or small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, low cup-shaped, and multispiral (Fig.
<figureCitation id="66F24BE063538C6B29EC4F0C478645A6" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2D552637481DC58575BC79DA6EEE39F0" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A8474617246E1C210A9F0F38C9D47D60" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="CCAD25566C942A2E1B571BA7B207C2FE" pageId="0" pageNumber="1">
This species is known from Lai Chau, Son La and Dien Bien provinces, northern Vietnam. The current study recorded specimens from a few localities in Hua Phan Province, northern Laos (Fig.
<figureCitation id="959B7E46F8D57B079CB90081D044B195" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Distribution map of Rhiostoma asiphon - species group." figureDoi="10.3897/zookeys.1142.90097.figure34" httpUri="https://binary.pensoft.net/fig/799892" pageId="0" pageNumber="1">34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4ADD85F0440E43A1F42B102D522D6A0A" pageId="0" pageNumber="1" type="remarks">
<paragraph id="2A3E5A3FEF66976DA4ABAFA3CB5BDEBC" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="EFD9D82916267C77B8EC18F7ABAE9F4E" pageId="0" pageNumber="1">
The holotypes of
<taxonomicName id="03485943805FE6437CB87EDC19366584" lsidName="R. abletti" pageId="0" pageNumber="1" rank="species" species="abletti">
<emphasis id="3FFBB7448CFE2CCF5307D7F289521B4E" italics="true" pageId="0" pageNumber="1">R. abletti</emphasis>
</taxonomicName>
from Lai Chau Province (Fig.
<figureCitation id="3C313CEA43452D664404DC1CF9A53DE7" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37A</figureCitation>
), &quot;
<taxonomicName id="3161FBCAB3B0C692778737B6F899B1AC" lsidName="R. christae" pageId="0" pageNumber="1" rank="species" species="christae">
<emphasis id="A9ACACB46065FD1344054B135129FD1C" italics="true" pageId="0" pageNumber="1">R. christae</emphasis>
</taxonomicName>
&quot; from Son La Province (Fig.
<figureCitation id="1DA1B62680D8EC7866798412C533D521" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Shell of Rhiostoma abletti A holotype NHMUK 20160307 from Lai Chau Province, Vietnam B holotype of Rhiostoma christae Thach, 2016, NHMUK 20160306 from Son La Province, Vietnam C, D specimens CUMZ 10207 from Ban Na Puek, Meuang Hiam, Houaphanh, Laos E, F specimens CUMZ 10206 from Hot Springs, Meuang Hiam, Houaphanh, Laos." figureDoi="10.3897/zookeys.1142.90097.figure37" httpUri="https://binary.pensoft.net/fig/799895" pageId="0" pageNumber="1">37B</figureCitation>
) and &quot;
<taxonomicName id="84D1B2572E5EFCF8BA3CCDF4FA1AEBF2" lsidName="R. ninhbien" pageId="0" pageNumber="1" rank="species" species="ninhbien">
<emphasis id="29D20AD4BF825ED6CCB9D6DF1858891D" italics="true" pageId="0" pageNumber="1">R. ninhbien</emphasis>
</taxonomicName>
&quot; from Dien Bien Province (see
<bibRefCitation id="35CE07AF3240669041BD03777E0E714D" DOI="https://doi.org/10.35885/ruthenica.2021.30(3).4" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="165 - 172" refId="B36" refString="Do, DS, Nguyen, TS, Do, HL, 2020b. The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species. Ruthenica 30 (3): 165 - 172, DOI: https://doi.org/10.35885/ruthenica.2021.30(3).4" title="The land snail genus Rhiostoma Benson, 1860 from Vietnam (Gastropoda: Caenogastropoda: Cyclophoridae), with description of a new species." url="https://doi.org/10.35885/ruthenica.2021.30(3).4" volume="30" year="2020 b">Do et al. 2020b</bibRefCitation>
: fig. 2a-c) are nearly identical in appearance. They have a depressed shell, with short detached whorl and incomplete tube shape, but are slightly different in the shell colour pattern, which could not be used as a diagnostic trait. In addition, the type localities of these three species are in neighbouring areas of northern Vietnam, less than 150 km apart. Therefore, we recognise
<taxonomicName id="C69778DCE2A82685B6A670FF3591DA9C" lsidName="R. christae" pageId="0" pageNumber="1" rank="species" species="christae">
<emphasis id="AF05C4CED75F56DAF425A25B6CDAE16B" italics="true" pageId="0" pageNumber="1">R. christae</emphasis>
</taxonomicName>
and
<taxonomicName id="4009187EC5558DBB2755368AAC77CBCB" lsidName="R. ninhbien" pageId="0" pageNumber="1" rank="species" species="ninhbien">
<emphasis id="94C48C45166F0B174B8725B66A530188" italics="true" pageId="0" pageNumber="1">R. ninhbien</emphasis>
</taxonomicName>
as junior subjective synonyms of
<taxonomicName id="940B0BE00CAA0265F8AB844ADEFAE115" lsidName="R. abletti" pageId="0" pageNumber="1" rank="species" species="abletti">
<emphasis id="C8F007452208A30CAFB463F4B86AABF0" italics="true" pageId="0" pageNumber="1">R. abletti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7C541D6CA815701416BDD919C9349DEC" pageId="0" pageNumber="1">
The specimens from Hua Phan Province, Laos, tend to have thicker periostracum and more coarse growth lines than the type specimen. However, a short detached whorl and incomplete tube as breathing device suggest they are more closely related to
<taxonomicName id="5564B1ED40F7C550CA8AD5684166D179" lsidName="R. abletti" pageId="0" pageNumber="1" rank="species" species="abletti">
<emphasis id="8218592E0F32E3B1E86E9546AC5B2B99" italics="true" pageId="0" pageNumber="1">R. abletti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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