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<document id="06A7B3098B06529185E16D5701D9DA55" ID-DOI="10.26107/RBZ-2022-0017" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174641" ID-ZooBank="6A65DD81-A22F-4901-8B2A-029695E4AB43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665291496255" checkinUser="felipe" docAuthor="Rahayu, Dwi Listyo" docDate="2022" docId="37778780FFC1E940DA52F588FAFEF30E" docLanguage="en" docName="RafflesBZool.70.329363.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Diogenes fasciatus Rahayu &amp; Forest 1995" docType="treatment" docVersion="4" lastPageNumber="343" masterDocId="CB4EFFF8FFCCE94ED92AF135FF96F21B" masterDocTitle="Hermit crabs of Singapore (Crustacea: Decapoda: Anomura: Diogenidae, Paguridae), with description of two new species" masterLastPageNumber="363" masterPageNumber="329" pageNumber="342" updateTime="1699380117419" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="51806205CF39F5C5A54443662AE38BD1">Hermit crabs of Singapore (Crustacea: Decapoda: Anomura: Diogenidae, Paguridae), with description of two new species</mods:title>
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<mods:namePart id="E21FB2ABBD1625CE769E802FA43AD2C9">Rahayu, Dwi Listyo</mods:namePart>
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<mods:date id="29AE4ECEF86457FA39CD4EFD797C39D8">2022</mods:date>
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<paragraph id="BF613696FFC1E943DA52F588FAC9F6CE" blockId="13.[888,1375,1213,1269]" box="[888,1375,1213,1237]" pageId="13" pageNumber="342">
<heading id="E42981FAFFC1E943DA52F588FAC9F6CE" bold="true" box="[888,1375,1213,1237]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="342" reason="2">
<taxonomicName id="78DE4D15FFC1E943DA52F588FAC9F6CE" ID-CoL="cae50cdf-168d-4e97-893b-422e0e1af9a7" authority="Rahayu &amp; Forest, 1995" authorityName="Rahayu &amp; Forest" authorityYear="1995" box="[888,1375,1213,1237]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="342" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC1E943DA52F588FAC9F6CE" bold="true" box="[888,1375,1213,1237]" pageId="13" pageNumber="342">
<emphasis id="8DAAEA84FFC1E943DA52F588FBDFF6CE" bold="true" box="[888,1097,1213,1237]" italics="true" pageId="13" pageNumber="342">Diogenes fasciatus</emphasis>
<bibRefCitation id="DB4F4B67FFC1E943DD78F588FAC9F6CE" author="Rahayu DL &amp; Forest J" box="[1106,1375,1213,1237]" pageId="13" pageNumber="342" pagination="383 - 415" refId="ref32714" refString="Rahayu DL &amp; Forest J (1995) Le genre Diogenes (Crustacea, Decapoda, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin Museum National d'Histoire Naturelle, Paris, 16 (2 - 4): 383 - 415." type="journal article" year="1995">Rahayu &amp; Forest, 1995</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="BF613696FFC1E943DD10F5E8FB0BF6EE" blockId="13.[888,1375,1213,1269]" box="[1082,1181,1245,1269]" pageId="13" pageNumber="342">
(
<figureCitation id="27E52A13FFC1E943DD68F5E8FB00F6EE" box="[1090,1174,1245,1269]" captionStart="Fig" captionStartId="12.[135,170,1931,1953]" captionTargetBox="[135,1473,236,1916]" captionTargetId="figure-7@12.[135,1473,236,1916]" captionTargetPageId="12" captionText="Fig. 8. A, Diogenes avarus Heller, 1852, not collected, IT86, Cyrene Reef, 1°15.374N 103°44.816E, 27 May 2013; B, Diogenes fasciatus Rahayu &amp; Forest, 1995, male, 5.3 mm (ZRC 2021.0132); C, Diogenes klaasi Rahayu &amp; Forest, 1995, male, not collected, Singapore Strait; D, Diogenes laevicarpus Rahayu, 1996, male, 5.1 mm (ZRC 2021.0139); E, Diogenes moosai Rahayu &amp; Forest, 1995, male, 1.8 mm (ZRC 2021.0160); F, Diogenes rectimanus Miers, 1884, male, not collected, DW58, East of Pulau Tekong, 01°25.064N 104°04.992E, 10.911.3 m, 22 October 2012." figureDoi="http://doi.org/10.5281/zenodo.7174661" httpUri="https://zenodo.org/record/7174661/files/figure.png" pageId="13" pageNumber="342">Fig. 8B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7C4651DFFC1E943DA07F428FAD8F770" pageId="13" pageNumber="342" type="reference_group">
<paragraph id="BF613696FFC1E943DA07F428FAD8F770" blockId="13.[813,1453,1308,1387]" pageId="13" pageNumber="342">
<treatmentCitationGroup id="9FCE11B8FFC1E943DA07F428FAD8F770" pageId="13" pageNumber="342">
<treatmentCitation id="3E7F1087FFC1E943DA07F428FB7BF72A" author="Rahayu DL &amp; Forest J" box="[813,1261,1308,1330]" page="388" pageId="13" pageNumber="342" year="1995">
<taxonomicName id="78DE4D15FFC1E943DA07F428FB7BF72A" ID-CoL="cae50cdf-168d-4e97-893b-422e0e1af9a7" authority="Rahayu &amp; Forest, 1995: 388" authorityName="Rahayu &amp; Forest" authorityPageNumber="388" authorityYear="1995" box="[813,1261,1308,1330]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="342" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC1E943DA07F428FC49F72A" box="[813,991,1308,1330]" italics="true" pageId="13" pageNumber="342">Diogenes fasciatus</emphasis>
<bibRefCitation id="DB4F4B67FFC1E943DAC9F429FB7BF72A" author="Rahayu DL &amp; Forest J" box="[995,1261,1308,1330]" pageId="13" pageNumber="342" pagination="383 - 415" refId="ref32714" refString="Rahayu DL &amp; Forest J (1995) Le genre Diogenes (Crustacea, Decapoda, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin Museum National d'Histoire Naturelle, Paris, 16 (2 - 4): 383 - 415." type="journal article" year="1995">Rahayu &amp; Forest, 1995: 388</bibRefCitation>
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, fig. 1 (
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locality: Muara Karang,
<collectingRegion id="7D1AF874FFC1E943DADAF40CFBA2F754" box="[1008,1076,1337,1359]" country="Indonesia" name="Jakarta Raya" pageId="13" pageNumber="342">Jakarta</collectingRegion>
Bay,
<collectingCountry id="C7C97606FFC1E943DD59F40CFB43F754" box="[1139,1237,1337,1359]" name="Indonesia" pageId="13" pageNumber="342">Indonesia</collectingCountry>
);
<treatmentCitation id="3E7F1087FFC1E943DDC2F40CFA31F755" author="Rahayu DL" box="[1256,1447,1337,1359]" page="340" pageId="13" pageNumber="342" year="1996">
<bibRefCitation id="DB4F4B67FFC1E943DDC2F40CFA31F755" author="Rahayu DL" box="[1256,1447,1337,1359]" pageId="13" pageNumber="342" pagination="335 - 355" refId="ref32232" refString="Rahayu DL (1996) Notes on littoral hermit crabs (excluding Coenobitidae) (Crustacea: Decapoda: Anomura) mainly from Singapore and Peninsular Malaysia. Raffles Bulletin of Zoology, 44 (2): 335 - 355." type="journal article" year="1996">Rahayu, 1996: 340</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="3E7F1087FFC1E943DA78F460FB6BF770" author="Siddiqui FA &amp; Kazmi OB &amp; McLaughlin PA" box="[850,1277,1365,1387]" page="185" pageId="13" pageNumber="342" year="2004">
<bibRefCitation id="DB4F4B67FFC1E943DA78F460FB6BF770" author="Siddiqui FA &amp; Kazmi OB &amp; McLaughlin PA" box="[850,1277,1365,1387]" pageId="13" pageNumber="342" pagination="155 - 200" refId="ref33312" refString="Siddiqui FA, Kazmi OB &amp; McLaughlin PA (2004) Review of the Pakistani species of Diogenes Dana, 1951 (Decapoda Anomura Paguroidea Diogenidae). Tropical Zoology, 17: 155 - 200." type="journal article" year="2004">Siddiqui, Kazmi &amp; McLaughlin, 2004: 185</bibRefCitation>
</treatmentCitation>
, fig. 13.
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</paragraph>
</subSubSection>
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<paragraph id="BF613696FFC1E943DA07F4A6FA3AF597" blockId="13.[812,1453,1427,1932]" pageId="13" pageNumber="342">
<emphasis id="8DAAEA84FFC1E943DA07F4A6FB81F7B0" bold="true" box="[813,1047,1427,1451]" pageId="13" pageNumber="342">Material examined.</emphasis>
<specimenCount id="A9D8FD1FFFC1E943DD08F4A1FB1DF7B7" box="[1058,1163,1428,1452]" pageId="13" pageNumber="342" type="male">96 males</specimenCount>
,
<quantity id="78269B73FFC1E943DDB1F4A1FAB0F7B0" box="[1179,1318,1428,1452]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.9" metricValueMin="0.7" pageId="13" pageNumber="342" unit="mm" value="1.7999999999999998" valueMax="2.9" valueMin="0.7">0.72.9 mm</quantity>
,
<materialsCitation id="0FB63CCBFFC1E943DC1CF4A1FB40F7F7" collectingDate="1995-01-18" collectionCode="ZRC" location="Changi" pageId="13" pageNumber="342" specimenCode="ZRC 2021.0130" specimenCount="5" specimenCount-female="5">
<specimenCount id="A9D8FD1FFFC1E943DC1CF4A1FA31F7B7" box="[1334,1447,1428,1452]" pageId="13" pageNumber="342" type="female">5 females</specimenCount>
,
<quantity id="78269B73FFC1E943DA07F481FC2DF7D0" box="[813,955,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="13" pageNumber="342" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.82.5 mm</quantity>
, 20 ovigerous females,
<quantity id="78269B73FFC1E943DDFCF481FAF7F7D0" box="[1238,1377,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="13" pageNumber="342" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.82.5 mm</quantity>
(
<specimenCode id="EF789EEDFFC1E943DC5EF481FC32F7F7" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="342">ZRC 2021.0130</specimenCode>
),
<location id="BA01604DFFC1E943DA90F4E1FB9DF7F7" LSID="urn:lsid:plazi:treatment:37778780FFC1E940DA52F588FAFEF30E:BA01604DFFC1E943DA90F4E1FB9DF7F7" box="[954,1035,1492,1516]" name="Changi" pageId="13" pageNumber="342">Changi</location>
,
<date id="CB601056FFC1E943DD33F4E1FB40F7F7" box="[1049,1238,1492,1516]" pageId="13" pageNumber="342" value="1995-01-18">
<collectingDate id="DB24E9BEFFC1E943DD33F4E1FB40F7F7" box="[1049,1238,1492,1516]" pageId="13" pageNumber="342" value="1995-01-18">18 January 1995</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="A9D8FD1FFFC1E943DDCEF4E1FADAF7F7" box="[1252,1356,1492,1516]" pageId="13" pageNumber="342" type="male">20 males</specimenCount>
,
<quantity id="78269B73FFC1E943DC70F4E1FCCDF410" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.2" metricValueMin="2.4" pageId="13" pageNumber="342" unit="mm" value="3.3" valueMax="4.2" valueMin="2.4">2.44.2 mm</quantity>
,
<materialsCitation id="0FB63CCBFFC1E943DA41F4C1FA31F437" collectingDate="1995-02-16" collectionCode="ZRC" location="Changi" pageId="13" pageNumber="342" specimenCode="ZRC 2021.0131" specimenCount="12" specimenCount-female="12">
<specimenCount id="A9D8FD1FFFC1E943DA41F4C1FC66F417" box="[875,1008,1524,1548]" pageId="13" pageNumber="342" type="female">12 females</specimenCount>
,
<quantity id="78269B73FFC1E943DD2BF4C1FB05F410" box="[1025,1171,1524,1548]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="13" pageNumber="342" unit="mm" value="2.0" valueMax="2.4" valueMin="1.6">1.62.4 mm</quantity>
, 20 ovigerous females,
<quantity id="78269B73FFC1E943DA07F721FC38F430" box="[813,942,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.9" metricValueMin="1.6" pageId="13" pageNumber="342" unit="mm" value="2.25" valueMax="2.9" valueMin="1.6">1.62.9 mm</quantity>
(
<specimenCode id="EF789EEDFFC1E943DA94F721FBE5F437" box="[958,1139,1556,1580]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="342">ZRC 2021.0131</specimenCode>
),
<location id="BA01604DFFC1E943DDACF721FB40F437" LSID="urn:lsid:plazi:treatment:37778780FFC1E940DA52F588FAFEF30E:BA01604DFFC1E943DDACF721FB40F437" box="[1158,1238,1556,1580]" name="Changi" pageId="13" pageNumber="342">Changi</location>
,
<date id="CB601056FFC1E943DDC8F721FA31F437" box="[1250,1447,1556,1580]" pageId="13" pageNumber="342" value="1995-02-16">
<collectingDate id="DB24E9BEFFC1E943DDC8F721FA31F437" box="[1250,1447,1556,1580]" pageId="13" pageNumber="342" value="1995-02-16">16 February 1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0FB63CCBFFC1E943DA05F701FC50F497" collectingDate="2012-10-22" collectionCode="ZRC" elevation="11" latitude="1.4177333" location="East of Pulau Tekong" longLatPrecision="1" longitude="104.0832" pageId="13" pageNumber="342" specimenCode="ZRC 2021.0132" specimenCount="8" specimenCount-male="8">
<specimenCount id="A9D8FD1FFFC1E943DA05F701FC1CF457" box="[815,906,1588,1612]" pageId="13" pageNumber="342" type="male">8 males</specimenCount>
,
<quantity id="78269B73FFC1E943DAB2F701FBB6F450" box="[920,1056,1588,1612]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="5.3" metricValueMin="2.3" pageId="13" pageNumber="342" unit="mm" value="3.8" valueMax="5.3" valueMin="2.3">2.35.3 mm</quantity>
(
<specimenCode id="EF789EEDFFC1E943DD18F701FB7AF457" box="[1074,1260,1588,1612]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="342">ZRC 2021.0132</specimenCode>
), DW58,
<location id="BA01604DFFC1E943DC77F701FC53F477" LSID="urn:lsid:plazi:treatment:37778780FFC1E940DA52F588FAFEF30E:BA01604DFFC1E943DC77F701FC53F477" latitude="1.4177333" longLatPrecision="1" longitude="104.0832" name="East of Pulau Tekong" pageId="13" pageNumber="342">East of Pulau Tekong</location>
,
<geoCoordinate id="DAEA5051FFC1E943DAE5F767FBC1F470" box="[975,1111,1618,1643]" degrees="01" direction="north" minutes="25.064" orientation="latitude" pageId="13" pageNumber="342" precision="1" value="1.4177333">01°25.064N</geoCoordinate>
<geoCoordinate id="DAEA5051FFC1E943DD74F767FB65F470" box="[1118,1267,1618,1643]" degrees="104" direction="east" minutes="04.992" orientation="longitude" pageId="13" pageNumber="342" precision="1" value="104.0832">104°04.992E</geoCoordinate>
,
<quantity id="78269B73FFC1E943DDD7F761FA10F470" box="[1277,1414,1620,1644]" metricMagnitude="1" metricUnit="m" metricValue="1.11" metricValueMax="1.13" metricValueMin="1.09" pageId="13" pageNumber="342" unit="m" value="11.1" valueMax="11.3" valueMin="10.9">
<elevation id="34F3D1A5FFC1E943DDD7F761FA10F470" box="[1277,1414,1620,1644]" metricMagnitude="1" metricUnit="m" metricValue="1.11" metricValueMax="1.13" metricValueMin="1.09" pageId="13" pageNumber="342" unit="m" value="11.1" valueMax="11.3" valueMin="10.9">10.911.3 m</elevation>
</quantity>
,
<date id="CB601056FFC1E943DCBBF761FC50F497" pageId="13" pageNumber="342" value="2012-10-22">
<collectingDate id="DB24E9BEFFC1E943DCBBF761FC50F497" pageId="13" pageNumber="342" value="2012-10-22">22 October 2012</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="A9D8FD1FFFC1E943DAFFF741FBADF497" box="[981,1083,1652,1676]" pageId="13" pageNumber="342" type="male">15 males</specimenCount>
,
<quantity id="78269B73FFC1E943DD63F741FB46F490" box="[1097,1232,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="13" pageNumber="342" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.62.2 mm</quantity>
,
<materialsCitation id="0FB63CCBFFC1E943DDF5F741FB0DF4D7" collectingDate="2001-09-21" collectionCode="ZRC" location="Chek Jawa" pageId="13" pageNumber="342" specimenCode="ZRC 2021.0133" specimenCount="4" specimenCount-female="4">
<specimenCount id="A9D8FD1FFFC1E943DDF5F741FADAF497" box="[1247,1356,1652,1676]" pageId="13" pageNumber="342" type="female">4 females</specimenCount>
, 1.62.0 mm, 5 ovigerous females, 1.82.0 mm (
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),
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, 1.0 mm (
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(
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), st. M24,
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1 ovigerous female,
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(
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), st. M13,
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, 4.56.0 mm (
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),
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, beach seine,
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,
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,
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.
</paragraph>
<paragraph id="BF613696FFC1E940DA07F686FC91F3CE" blockId="13.[813,1452,1971,2060]" lastBlockId="14.[136,775,189,469]" lastPageId="14" lastPageNumber="343" pageId="13" pageNumber="342">
<emphasis id="8DAAEA84FFC1E943DA07F686FC4EF5D0" bold="true" box="[813,984,1971,1995]" pageId="13" pageNumber="342">Colour in life.</emphasis>
Shield mottled with dark grey and light brown. Ocular peduncle white with greyish brown blotch on dorsomesial face distally, light brown patch proximally. Antennular peduncle generally greyish brown, with dark brown band on distal margin of ultimate and penultimate segments, flagella white; first to fourth segments and antennal acicle of antennal peduncle greyish brown, fifth segment and flagella white. Chelipeds palm whitish grey; dactyl and fixed finger with tint of light brown, carpi and meri dark brown or dark grey. P2 and P3 dark brown or dark grey, lighter grey dorsally and ventrally on propodi and carpi, with white spot on articulation dactyl, propodus and carpus (
<figureCitation id="27E52A13FFC2E940DB8EF088FD6FF3CE" box="[676,761,445,469]" captionStart="Fig" captionStartId="12.[135,170,1931,1953]" captionTargetBox="[135,1473,236,1916]" captionTargetId="figure-7@12.[135,1473,236,1916]" captionTargetPageId="12" captionText="Fig. 8. A, Diogenes avarus Heller, 1852, not collected, IT86, Cyrene Reef, 1°15.374N 103°44.816E, 27 May 2013; B, Diogenes fasciatus Rahayu &amp; Forest, 1995, male, 5.3 mm (ZRC 2021.0132); C, Diogenes klaasi Rahayu &amp; Forest, 1995, male, not collected, Singapore Strait; D, Diogenes laevicarpus Rahayu, 1996, male, 5.1 mm (ZRC 2021.0139); E, Diogenes moosai Rahayu &amp; Forest, 1995, male, 1.8 mm (ZRC 2021.0160); F, Diogenes rectimanus Miers, 1884, male, not collected, DW58, East of Pulau Tekong, 01°25.064N 104°04.992E, 10.911.3 m, 22 October 2012." figureDoi="http://doi.org/10.5281/zenodo.7174661" httpUri="https://zenodo.org/record/7174661/files/figure.png" pageId="14" pageNumber="343">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="BF613696FFC2E940D9A2F0C8FC91F02E" blockId="14.[136,775,509,566]" pageId="14" pageNumber="343">The general colouration in life varies from yellowish light brown to dark brown or dark grey, but the pattern is the same.</paragraph>
</subSubSection>
<subSubSection id="F7C4651DFFC2E940D9A2F368FDA3F08E" pageId="14" pageNumber="343" type="distribution">
<paragraph id="BF613696FFC2E940D9A2F368FDA3F08E" blockId="14.[135,775,605,661]" pageId="14" pageNumber="343">
<emphasis id="8DAAEA84FFC2E940D9A2F368FE8CF06E" bold="true" box="[136,282,605,629]" pageId="14" pageNumber="343">Distribution.</emphasis>
<collectingCountry id="C7C97606FFC2E940D835F368FEEAF06E" box="[287,380,605,629]" name="Pakistan" pageId="14" pageNumber="343">Pakistan</collectingCountry>
,
<collectingCountry id="C7C97606FFC2E940D8ACF368FE63F06E" box="[390,501,605,629]" name="Singapore" pageId="14" pageNumber="343">Singapore</collectingCountry>
,
<collectingCountry id="C7C97606FFC2E940D8D4F368FDF4F06E" box="[510,610,605,629]" name="Malaysia" pageId="14" pageNumber="343">Malaysia</collectingCountry>
, and
<collectingCountry id="C7C97606FFC2E940DBB2F368FC95F06E" box="[664,771,605,629]" name="Indonesia" pageId="14" pageNumber="343">Indonesia</collectingCountry>
;
<quantity id="78269B73FFC2E940D9ADF348FF58F08E" box="[135,206,637,661]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="7.0" metricValueMin="0.0" pageId="14" pageNumber="343" unit="m" value="3.5" valueMax="7.0" valueMin="0.0">07 m</quantity>
, inhabit sandy, muddy substrate.
</paragraph>
</subSubSection>
<subSubSection id="F7C4651DFFC2E940D9A2F388FAFEF30E" pageId="14" pageNumber="343" type="discussion">
<paragraph id="BF613696FFC2E940D9A2F388FE63F76E" blockId="14.[135,777,701,1397]" pageId="14" pageNumber="343">
<emphasis id="8DAAEA84FFC2E940D9A2F388FF61F0CE" bold="true" box="[136,247,701,725]" pageId="14" pageNumber="343">Remarks.</emphasis>
Among the species in the
<taxonomicName id="78DE4D15FFC2E940DB0AF388FD1FF0CE" baseAuthorityName="De Haan" baseAuthorityYear="1849" box="[544,649,701,725]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="8DAAEA84FFC2E940DB0AF388FD1FF0CE" box="[544,649,701,725]" italics="true" pageId="14" pageNumber="343">edwardsii</emphasis>
</taxonomicName>
group, this species is readily recognised by an ensemble of characters as follows: antennal peduncles stout, flagellum with abundant setae; left cheliped dactyl broad, flattened, covered with small tubercls; fixed finger outer surface tuberculate and devoid of longitudinal ridge; and palm with row of strong spines on upper and lower margins, at proximoventral angle row of strong spines parallel to carpal articulation, reaching to distal quarter, then continued as longitudinal row/rows of small spines on outer surface. The differences between
<taxonomicName id="78DE4D15FFC2E940D9A2F2CBFE9CF60E" authorityName="Rahayu &amp; Forest" authorityYear="1995" box="[136,266,1021,1045]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC2E940D9A2F2CBFE9CF60E" box="[136,266,1021,1045]" italics="true" pageId="14" pageNumber="343">D. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="78DE4D15FFC2E940D833F2CBFD81F60E" authority="Heller, 1865" authorityName="Heller" authorityYear="1865" box="[281,535,1021,1045]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="avarus">
<emphasis id="8DAAEA84FFC2E940D833F2CBFE13F60E" box="[281,389,1022,1045]" italics="true" pageId="14" pageNumber="343">D. avarus</emphasis>
<bibRefCitation id="DB4F4B67FFC2E940D8A7F2C8FD81F60E" author="Heller C" box="[397,535,1021,1045]" pageId="14" pageNumber="343" refId="ref29241" refString="Heller C (1865) Crustaceen. In: Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodors B. von Wullerstorf-Urbair. Zoologischer Theil, Bd. 2, Abt. 3. Kaiserlich-koniglichen Hof- und Staatsdruckerei, Wien, 280 pp., 25 pls." type="book" year="1865">Heller, 1865</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="78DE4D15FFC2E940DB7FF2CBFEB0F62E" authority="Alcock, 1905 b" authorityName="Alcock" authorityYear="1905" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="investigatoris">
<emphasis id="8DAAEA84FFC2E940DB7FF2CBFC91F60E" box="[597,775,1022,1045]" italics="true" pageId="14" pageNumber="343">D. investigatoris</emphasis>
<bibRefCitation id="DB4F4B67FFC2E940D9A2F528FEB0F62E" author="Alcock A" box="[136,294,1053,1077]" pageId="14" pageNumber="343" refId="ref27802" refString="Alcock A (1905 b) Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum. Part 2. Anomura. Fasc. 1. Pagurides. Indian Museum, Calcutta, xi + 197 pp." type="book" year="1905">Alcock, 1905b</bibRefCitation>
</taxonomicName>
were already discussed by
<bibRefCitation id="DB4F4B67FFC2E940DB65F528FF45F64E" author="Rahayu DL &amp; Forest J" pageId="14" pageNumber="343" pagination="383 - 415" refId="ref32714" refString="Rahayu DL &amp; Forest J (1995) Le genre Diogenes (Crustacea, Decapoda, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin Museum National d'Histoire Naturelle, Paris, 16 (2 - 4): 383 - 415." type="journal article" year="1995">Rahayu &amp; Forest (1995)</bibRefCitation>
, and the significance of those characters has been confirmed. The flattened dactyl of the left cheliped and the presence of a row or rows of spines on the propodi of P2 and P3 placed
<taxonomicName id="78DE4D15FFC2E940D9D2F5ABFEEEF6AE" authorityName="Rahayu &amp; Forest" authorityYear="1995" box="[248,376,1181,1205]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC2E940D9D2F5ABFEEEF6AE" box="[248,376,1181,1205]" italics="true" pageId="14" pageNumber="343">D. fasciatus</emphasis>
</taxonomicName>
close to
<taxonomicName id="78DE4D15FFC2E940D8F3F5ABFC92F6AE" authority="Morgan, 1989" authorityName="Morgan" authorityYear="1989" box="[473,772,1181,1206]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="lophochir">
<emphasis id="8DAAEA84FFC2E940D8F3F5ABFDF4F6AE" box="[473,610,1181,1205]" italics="true" pageId="14" pageNumber="343">D. lophochir</emphasis>
<bibRefCitation id="DB4F4B67FFC2E940DB43F5ABFC92F6AE" author="Morgan GJ" box="[617,772,1181,1206]" pageId="14" pageNumber="343" pagination="391 - 417" refId="ref31577" refString="Morgan GJ (1989) The hermit crabs (Decapoda: Anomura: Diogenidae, Paguridae) of southwestern Australia, with description of two new species. Records of the Western Australian Museum, 14 (3): 391 - 417." type="journal article" year="1989">Morgan, 1989</bibRefCitation>
</taxonomicName>
. But these two species differ in the presence of pronounced ridge on the fixed finger of the left cheliped in
<taxonomicName id="78DE4D15FFC2E940DBA0F5EBFC90F6EE" authorityName="Morgan" authorityYear="1989" box="[650,774,1245,1269]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="lopochir">
<emphasis id="8DAAEA84FFC2E940DBA0F5EBFC90F6EE" box="[650,774,1245,1269]" italics="true" pageId="14" pageNumber="343">D. lopochir</emphasis>
</taxonomicName>
(absence in
<taxonomicName id="78DE4D15FFC2E940D826F5CBFE1DF70E" authorityName="Rahayu &amp; Forest" authorityYear="1995" box="[268,395,1277,1301]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC2E940D826F5CBFE1DF70E" box="[268,395,1277,1301]" italics="true" pageId="14" pageNumber="343">D. fasciatus</emphasis>
</taxonomicName>
), the tuberculate crest on the outer face of the palm of the left cheliped is positioned in the midline in
<taxonomicName id="78DE4D15FFC2E940D828F40BFE18F74E" authorityName="Morgan" authorityYear="1989" box="[258,398,1341,1365]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="lophochir">
<emphasis id="8DAAEA84FFC2E940D828F40BFE18F74E" box="[258,398,1341,1365]" italics="true" pageId="14" pageNumber="343">D. lophochir</emphasis>
</taxonomicName>
while in
<taxonomicName id="78DE4D15FFC2E940D8D6F40BFDE8F74E" authorityName="Rahayu &amp; Forest" authorityYear="1995" box="[508,638,1341,1365]" class="Malacostraca" family="Diogenidae" genus="Diogenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="343" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="8DAAEA84FFC2E940D8D6F40BFDE8F74E" box="[508,638,1341,1365]" italics="true" pageId="14" pageNumber="343">D. fasciatus</emphasis>
</taxonomicName>
this crest is located adjacent to upper margin.
</paragraph>
<paragraph id="BF613696FFC2E940D9A2F4A8FAFEF30E" blockId="14.[135,778,1437,2037]" lastBlockId="14.[835,1474,189,277]" pageId="14" pageNumber="343">
<bibRefCitation id="DB4F4B67FFC2E940D9A2F4A8FE0EF7AE" author="Rahayu DL &amp; Forest J" box="[136,408,1437,1461]" pageId="14" pageNumber="343" pagination="383 - 415" refId="ref32714" refString="Rahayu DL &amp; Forest J (1995) Le genre Diogenes (Crustacea, Decapoda, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin Museum National d'Histoire Naturelle, Paris, 16 (2 - 4): 383 - 415." type="journal article" year="1995">Rahayu &amp; Forest (1995)</bibRefCitation>
did not mention the presence of row of spines on the dorsal margin of the propodi of P2 and P3 but indicated that the spines on the carpi are present only in the large individual.
<bibRefCitation id="DB4F4B67FFC2E940D8E7F4C8FD28F40E" author="Siddiqui FA &amp; Kazmi OB &amp; McLaughlin PA" box="[461,702,1533,1557]" pageId="14" pageNumber="343" pagination="155 - 200" refId="ref33312" refString="Siddiqui FA, Kazmi OB &amp; McLaughlin PA (2004) Review of the Pakistani species of Diogenes Dana, 1951 (Decapoda Anomura Paguroidea Diogenidae). Tropical Zoology, 17: 155 - 200." type="journal article" year="2004">Siddiqui et al. (2004)</bibRefCitation>
stated that these spines are present or absent in their material and that the specimens from
<collectingCountry id="C7C97606FFC2E940D8B2F708FE64F44E" box="[408,498,1597,1621]" name="Pakistan" pageId="14" pageNumber="343">Pakistan</collectingCountry>
have one or two rows of spines on carpi, while
<bibRefCitation id="DB4F4B67FFC2E940D8A6F768FDACF46E" author="Asakura A" box="[396,570,1629,1653]" pageId="14" pageNumber="343" pagination="1 - 4" refId="ref27914" refString="Asakura A (2020) Hermit crabs of the genus Diogenes Dana, 1851 (Crustacea: Decapoda: Diogenidae) collected during the Albatross Philippine Expedition, 1907 - 1910, including descriptions of three new species. Publications of the Seto Marine Biological Laboratory, 45: 1 - 4." type="journal article" year="2020">Asakura (2020)</bibRefCitation>
stated that, in the material from
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and
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, the dorsal margins of propodi of ambulatory legs have a row of spines, and also figured the spines on the dorsal margin of carpi (
<bibRefCitation id="DB4F4B67FFC2E940DB82F788FF57F4EE" author="Asakura A" pageId="14" pageNumber="343" pagination="1 - 4" refId="ref27914" refString="Asakura A (2020) Hermit crabs of the genus Diogenes Dana, 1851 (Crustacea: Decapoda: Diogenidae) collected during the Albatross Philippine Expedition, 1907 - 1910, including descriptions of three new species. Publications of the Seto Marine Biological Laboratory, 45: 1 - 4." type="journal article" year="2020">Asakura, 2020</bibRefCitation>
: fig. 11A, C, E). In some specimens in this study, the propodi and carpi of P2 and P3 have one or two rows of spines on the dorsal margin. Additionally, a tiny spine on the dorsodistal margin of the fourth segment of the antennal peduncles was also detected in some individuals. The spines on the mesial margin of the antennal acicle varies from very prominent to moderately large, and one additional spine is sometimes present at the midlength of the lateral margin. The same character was also observed by
<bibRefCitation id="DB4F4B67FFC2E940DB77F6E8FC90F5EE" author="Asakura A" box="[605,774,2013,2037]" pageId="14" pageNumber="343" pagination="1 - 4" refId="ref27914" refString="Asakura A (2020) Hermit crabs of the genus Diogenes Dana, 1851 (Crustacea: Decapoda: Diogenidae) collected during the Albatross Philippine Expedition, 1907 - 1910, including descriptions of three new species. Publications of the Seto Marine Biological Laboratory, 45: 1 - 4." type="journal article" year="2020">Asakura (2020)</bibRefCitation>
in his specimens. Further study is necessary to confirm that these characters mentioned above are merely attributed to intraspecific variations or interspecific differences.
</paragraph>
</subSubSection>
</treatment>
</document>