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<document id="9E4EE011FF2F4484F23A5ECE191D8FED" ID-CLB-Dataset="29418" ID-DOI="10.11646/zootaxa.4482.3.2" ID-GBIF-Dataset="fd24ef18-76ac-46f1-aec2-2faf5b0ff325" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440707" ID-ZooBank="3FBE11E2-9F97-4A29-92A9-67780C782E0D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538375301724" checkinUser="plazi" docAuthor="Li, Jheng-Jhang, Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2018" docId="376A1B4DFFA9FF89D1CDFC534107FE9A" docLanguage="en" docName="zootaxa.4482.3.2.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parasesarma macaco Li &amp; Rahayu &amp; Ng 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="466" masterDocId="CB536335FFA4FF86D15AFFF74432FFDA" masterDocTitle="Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific" masterLastPageNumber="490" masterPageNumber="451" pageNumber="464" updateTime="1698559507815" updateUser="ExternalLinkService">
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<mods:title id="967CD8F3A441E878A2AA4B632F922E1F">Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific</mods:title>
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<mods:namePart id="90CEDADBFAB0F20EC7D23DF124D3AE69">Li, Jheng-Jhang</mods:namePart>
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<mods:namePart id="6383F687CDC8151487896E5E19BB61F9">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="726F1383595B29EDD1E00FA72810A856">Ng, Peter K. L.</mods:namePart>
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<treatment id="376A1B4DFFA9FF89D1CDFC534107FE9A" ID-DOI="http://doi.org/10.5281/zenodo.5986789" ID-GBIF-Taxon="148421727" ID-Zenodo-Dep="5986789" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A" httpUri="http://treatment.plazi.org/id/376A1B4DFFA9FF89D1CDFC534107FE9A" lastPageId="15" lastPageNumber="466" pageId="13" pageNumber="464">
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<paragraph id="BF7CAA5BFFA9FF8BD1CDFC5345D0FC65" blockId="13.[151,733,932,992]" box="[151,482,932,959]" pageId="13" pageNumber="464">
<heading id="E4341D37FFA9FF8BD1CDFC5345D0FC65" bold="true" box="[151,482,932,959]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
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<taxonomicName id="78C3D1D8FFA9FF8BD1CDFC5345ABFC64" ID-CoL="84G4C" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,409,932,958]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD1CDFC5345ABFC64" bold="true" box="[151,409,932,958]" italics="true" pageId="13" pageNumber="464">Parasesarma macaco</emphasis>
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<taxonomicNameLabel id="9684CB32FFA9FF8BD0FAFC5245D0FC65" box="[416,482,933,959]" pageId="13" pageNumber="464" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="BF7CAA5BFFA9FF8BD1CDFC3046EFFC3A" blockId="13.[151,733,932,992]" box="[151,733,967,992]" pageId="13" pageNumber="464">
(
<figureCitation id="27F8B6DEFFA9FF8BD1C5FC3044DCFC05" box="[159,238,967,992]" captionStart="FIGURE 7" captionStartId="14.[151,250,1353,1375]" captionTargetBox="[152,1435,193,1332]" captionTargetId="figure@14.[152,1435,193,1332]" captionTargetPageId="14" captionText="FIGURE 7. Parasesarma macaco n. sp.. AD, holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; E, F, paratype female (13.3×11.1 mm) (NMNS-7779-013), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440723/files/figure.png" pageId="13" pageNumber="464">Figs. 7</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD1A0FC3F4538FC3A" box="[250,266,968,992]" captionStart="FIGURE 8" captionStartId="15.[151,250,1572,1594]" captionTargetBox="[196,1307,452,1520]" captionTargetId="figure@15.[152,1435,375,1550]" captionTargetPageId="15" captionText="FIGURE 8. Parasesarma macaco n. sp., holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan. A, dorsal view of carapace; BD, tip of left cheliped; E, F, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, dorsal (sternal) view; F, ventral (pleonal) view. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440725/files/figure.png" pageId="13" pageNumber="464">8</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD04CFC3F4575FC3A" box="[278,327,968,992]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="13" pageNumber="464">19C</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD009FC3F45B7FC3A" box="[339,389,968,992]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="13" pageNumber="464">20B</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD0CAFC3F45F3FC3A" box="[400,449,968,992]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="13" pageNumber="464">21C</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD097FC3F45CCFC3A" box="[461,510,968,992]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="13" pageNumber="464">22B</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD353FC3F4608FC3A" box="[521,570,968,992]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="13" pageNumber="464">23B</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD31CFC3F4645FC3A" box="[582,631,968,992]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="13" pageNumber="464">24C</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD3D8FC3046E7FC3A" box="[642,725,967,992]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. AD, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="13" pageNumber="464">25BD</figureCitation>
)
</paragraph>
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<paragraph id="BF7CAA5BFFA9FF8BD1CDFBF84739F9FA" blockId="13.[151,1437,1039,2037]" pageId="13" pageNumber="464">
<materialsCitation id="0FABA006FFA9FF8BD1CDFBF840A2FB97" ID-GBIF-Occurrence="1913889743" collectingDate="2015-09-29" collectorName="J. Li" country="Taiwan" county="Material" location="Gaertn." municipality="Paoli River" pageId="13" pageNumber="464" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="8DB77649FFA9FF8BD1CDFBF845B0FBF2" bold="true" box="[151,386,1039,1064]" pageId="13" pageNumber="464">
<collectingCounty id="561DD2D7FFA9FF8BD1CDFBF844CDFBF2" box="[151,255,1039,1064]" pageId="13" pageNumber="464">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="607814F9FFA9FF8BD0D1FBE74611FBF2" box="[395,547,1040,1064]" pageId="13" pageNumber="464" type="holotype">HOLOTYPE</typeStatus>
: male (13.8×
<quantity id="783B07BEFFA9FF8BD3E2FBE74712FBF2" box="[696,800,1040,1064]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="13" pageNumber="464" unit="mm" value="12.3">12.3 mm</quantity>
) (NMNS-7779-005),
<collectingCountry id="C7D4EACBFFA9FF8BD543FBF8405DFBF2" box="[1049,1135,1039,1064]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD526FBF840D4FBF2" box="[1148,1254,1039,1064]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<collectingMunicipality id="5F183021FFA9FF8BD438FBF844E5FB97" pageId="13" pageNumber="464">Paoli River</collectingMunicipality>
, north bank (
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on
<taxonomicName id="78C3D1D8FFA9FF8BD0C4FBC346ABFB96" box="[414,665,1076,1101]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="464" phylum="Tracheophyta" rank="species" species="inerme">
<emphasis id="8DB77649FFA9FF8BD0C4FBC346ABFB96" box="[414,665,1076,1101]" italics="true" pageId="13" pageNumber="464">Clerodendrum inerme</emphasis>
</taxonomicName>
</collectedFrom>
(L.)
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),
<collectingDate id="DB397573FFA9FF8BD212FBC3401FFB97" box="[840,1069,1076,1101]" pageId="13" pageNumber="464" value="2015-09-29">
<date id="CB7D8C9BFFA9FF8BD212FBC3401FFB97" box="[840,1069,1076,1101]" pageId="13" pageNumber="464" value="2015-09-29">29 September 2015</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD50DFBC240A2FB97" box="[1111,1168,1076,1101]" pageId="13" pageNumber="464">J. Li</collectorName>
</materialsCitation>
.
<materialsCitation id="0FABA006FFA9FF8BD5C5FBC3411DFBAA" ID-GBIF-Occurrence="1913889737" collectingDate="2016-07-05" collectionCode="ZRC" collectorName="J. Li." country="Taiwan" location="Paoli River" pageId="13" pageNumber="464" specimenCode="ZRC 2018.0788" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<typeStatus id="607814F9FFA9FF8BD5C5FBC3410CFB96" box="[1183,1342,1076,1100]" pageId="13" pageNumber="464" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="A9C561D2FFA9FF8BD417FBC341A9FB97" box="[1357,1435,1076,1101]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(12.3×
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) (
<specimenCode id="EF650220FFA9FF8BD03DFBAF461AFBAA" box="[359,552,1112,1136]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="464">ZRC 2018.0788</specimenCode>
),
<collectingCountry id="C7D4EACBFFA9FF8BD31BFBA046AAFBAA" box="[577,664,1111,1136]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD3F2FBA04723FBAA" box="[680,785,1111,1136]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD2C0FBA04013FBAA" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD2C0FBA04013FBAA" box="[922,1057,1111,1136]" country="Taiwan" name="Paoli River" pageId="13" pageNumber="464" stateProvince="Pingtung">Paoli River</location>
,
<collectingDate id="DB397573FFA9FF8BD568FBA040F6FBAA" box="[1074,1220,1111,1136]" pageId="13" pageNumber="464" value="2016-07-05">
<date id="CB7D8C9BFFA9FF8BD568FBA040F6FBAA" box="[1074,1220,1111,1136]" pageId="13" pageNumber="464" value="2016-07-05">5 July 2016</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD5AAFBAF411DFBAA" box="[1264,1327,1111,1136]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD415FBA04107FB4F" ID-GBIF-Occurrence="1913889755" collectingDate="2015-09-30" collectionCode="ZRC" collectorName="J. Li." country="Taiwan" location="Paoli River" pageId="13" pageNumber="464" specimenCode="ZRC 2018.0789" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD415FBA041AEFBAA" box="[1359,1436,1111,1136]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(11.6×10.0 mm) (
<specimenCode id="EF650220FFA9FF8BD006FB8B4624FB4E" box="[348,534,1148,1172]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="464">ZRC 2018.0789</specimenCode>
),
<collectingCountry id="C7D4EACBFFA9FF8BD370FB8B46B2FB4F" box="[554,640,1148,1173]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD3D1FB8B46C6FB4F" box="[651,756,1148,1173]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD236FB8B47DFFB4F" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD236FB8B47DFFB4F" box="[876,1005,1148,1173]" country="Taiwan" name="Paoli River" pageId="13" pageNumber="464" stateProvince="Pingtung">Paoli River</location>
,
<collectingDate id="DB397573FFA9FF8BD2A2FB8B40E7FB4F" box="[1016,1237,1148,1173]" pageId="13" pageNumber="464" value="2015-09-30">
<date id="CB7D8C9BFFA9FF8BD2A2FB8B40E7FB4F" box="[1016,1237,1148,1173]" pageId="13" pageNumber="464" value="2015-09-30">30 September 2015</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD5A1FB8A4107FB4F" box="[1275,1333,1148,1173]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD40FFB8B4102FB62" ID-GBIF-Occurrence="1913889756" collectingDate="2016-06-06" collectorName="J. Li." country="Taiwan" location="Paoli River" pageId="13" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD40FFB8B41AEFB4F" box="[1365,1436,1148,1173]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(15.2×
<quantity id="783B07BEFFA9FF8BD1BAFB684579FB62" box="[224,331,1183,1208]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="13" pageNumber="464" unit="mm" value="13.4">13.4 mm</quantity>
) (NMNS-7779-008),
<collectingCountry id="C7D4EACBFFA9FF8BD313FB684692FB62" box="[585,672,1183,1208]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD3F5FB68472BFB62" box="[687,793,1183,1208]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD2C4FB684011FB62" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD2C4FB684011FB62" box="[926,1059,1183,1208]" country="Taiwan" name="Paoli River" pageId="13" pageNumber="464" stateProvince="Pingtung">Paoli River</location>
,
<collectingDate id="DB397573FFA9FF8BD569FB6840F5FB62" box="[1075,1223,1183,1208]" pageId="13" pageNumber="464" value="2016-06-06">
<date id="CB7D8C9BFFA9FF8BD569FB6840F5FB62" box="[1075,1223,1183,1208]" pageId="13" pageNumber="464" value="2016-06-06">6 June 2016</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD5A8FB574102FB62" box="[1266,1328,1183,1208]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD40AFB684106FB07" ID-GBIF-Occurrence="1913889754" collectingDate="2016-06-22" collectorName="J. Li." country="Taiwan" location="Kangkou River" pageId="13" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD40AFB6841AEFB62" box="[1360,1436,1183,1208]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(13.9×
<quantity id="783B07BEFFA9FF8BD1BAFB334575FB06" box="[224,327,1220,1245]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="464" unit="mm" value="12.5">12.5 mm</quantity>
) (NMNS-7779-009),
<collectingCountry id="C7D4EACBFFA9FF8BD367FB3346A6FB07" box="[573,660,1220,1245]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD3C4FB334735FB07" box="[670,775,1220,1245]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD2DAFB33401DFB07" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD2DAFB33401DFB07" box="[896,1071,1220,1245]" country="Taiwan" name="Kangkou River" pageId="13" pageNumber="464" stateProvince="Pingtung">Kangkou River</location>
,
<collectingDate id="DB397573FFA9FF8BD560FB3340E6FB07" box="[1082,1236,1220,1245]" pageId="13" pageNumber="464" value="2016-06-22">
<date id="CB7D8C9BFFA9FF8BD560FB3340E6FB07" box="[1082,1236,1220,1245]" pageId="13" pageNumber="464" value="2016-06-22">22 June 2016</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD5A1FB324106FB07" box="[1275,1332,1220,1245]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD40FFB334043FADA" ID-GBIF-Occurrence="1913889740" collectingDate="2016-03-05" collectorName="B. Chan." country="Philippines" location="Mindoro" pageId="13" pageNumber="464" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD40FFB3341AEFB07" box="[1365,1436,1220,1245]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(9.7×
<quantity id="783B07BEFFA9FF8BD188FB1F451CFADA" box="[210,302,1256,1280]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="13" pageNumber="464" unit="mm" value="8.4">8.4 mm</quantity>
) (NMNS-7779-010),
<location id="BA1CFC80FFA9FF8BD370FB1046BDFADA" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD370FB1046BDFADA" box="[554,655,1255,1280]" country="Philippines" name="Mindoro" pageId="13" pageNumber="464">Mindoro</location>
,
<collectingCountry id="C7D4EACBFFA9FF8BD3C7FB10472CFADA" box="[669,798,1255,1280]" name="Philippines" pageId="13" pageNumber="464">Philippines</collectingCountry>
,
<collectingDate id="DB397573FFA9FF8BD277FB1047E6FADA" box="[813,980,1255,1280]" pageId="13" pageNumber="464" value="2016-03-05">
<date id="CB7D8C9BFFA9FF8BD277FB1047E6FADA" box="[813,980,1255,1280]" pageId="13" pageNumber="464" value="2016-03-05">5 March 2016</date>
</collectingDate>
, W.-
<collectorName id="1236CF8DFFA9FF8BD554FB1F4043FADA" box="[1038,1137,1255,1280]" pageId="13" pageNumber="464">B. Chan.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD5D4FB104100FAFF" ID-GBIF-Occurrence="1913889751" collectingDate="2016-03-19" collectorName="J. Li." country="Taiwan" location="Paoli River" pageId="13" pageNumber="464" specimenCount="2" specimenCount-male="2" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD5D4FB1040D5FADA" box="[1166,1255,1255,1280]" pageId="13" pageNumber="464" type="male">2 males</specimenCount>
(10.1×
<quantity id="783B07BEFFA9FF8BD460FB1F41A5FB25" box="[1338,1431,1256,1280]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="464" unit="mm" value="8.9">8.9 mm</quantity>
, 12.9×
<quantity id="783B07BEFFA9FF8BD18DFAFB450DFAFE" box="[215,319,1292,1316]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="13" pageNumber="464" unit="mm" value="11.8">11.8 mm</quantity>
) (NMNS-7779-012),
<collectingCountry id="C7D4EACBFFA9FF8BD360FAFB46A2FAFF" box="[570,656,1292,1317]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD3C7FAFB4735FAFF" box="[669,775,1292,1317]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD2DDFAFB4039FAFF" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD2DDFAFB4039FAFF" box="[903,1035,1292,1317]" country="Taiwan" name="Paoli River" pageId="13" pageNumber="464" stateProvince="Pingtung">Paoli River</location>
,
<collectingDate id="DB397573FFA9FF8BD543FAFB40FFFAFF" box="[1049,1229,1292,1317]" pageId="13" pageNumber="464" value="2016-03-19">
<date id="CB7D8C9BFFA9FF8BD543FAFB40FFFAFF" box="[1049,1229,1292,1317]" pageId="13" pageNumber="464" value="2016-03-19">19 March 2016</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD5ACFAFA4100FAFF" box="[1270,1330,1292,1317]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
—1 male (12.8×
<quantity id="783B07BEFFA9FF8BD1BAFAC74561FA92" box="[224,339,1328,1352]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="13" pageNumber="464" unit="mm" value="11.3">11.3 mm</quantity>
),
<materialsCitation id="0FABA006FFA9FF8BD02AFAC7479CFAB7" ID-GBIF-Occurrence="1913889742" collectingDate="2015-10-04" collectorName="J. Li." country="Taiwan" location="Taijiang National Park" pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="Tainan" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD02AFAC745EEFA92" box="[368,476,1327,1352]" pageId="13" pageNumber="464" type="female">1 female</specimenCount>
(14.4×13.0 mm) (NMNS-7779-013),
<collectingCountry id="C7D4EACBFFA9FF8BD2EBFAD84035FA92" box="[945,1031,1327,1352]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD547FAD8405CFA92" box="[1053,1134,1327,1352]" country="Taiwan" name="Tainan" pageId="13" pageNumber="464">Tainan</collectingRegion>
:
<location id="BA1CFC80FFA9FF8BD5D8FAD841ABFA92" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD5D8FAD841ABFA92" box="[1154,1433,1327,1352]" country="Taiwan" name="Taijiang National Park" pageId="13" pageNumber="464" stateProvince="Tainan">Taijiang National Park</location>
: Sicao Mangrove Green Tunnel, South bank,
<collectingDate id="DB397573FFA9FF8BD3CFFAA34779FAB7" box="[661,843,1364,1389]" pageId="13" pageNumber="464" value="2015-10-04">
<date id="CB7D8C9BFFA9FF8BD3CFFAA34779FAB7" box="[661,843,1364,1389]" pageId="13" pageNumber="464" value="2015-10-04">4 October 2015</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD229FAA2479CFAB7" box="[883,942,1364,1389]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD295FAA347F7FA4A" ID-GBIF-Occurrence="1913889745" collectingDate="2016-10-04" collectorName="J. Li." country="Taiwan" location="Paoli River" pageId="13" pageNumber="464" specimenCount="1" specimenCount-female="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD295FAA3401FFAB7" box="[975,1069,1364,1389]" pageId="13" pageNumber="464" type="female">1 female</specimenCount>
(12.7×
<quantity id="783B07BEFFA9FF8BD524FAA340D5FAB6" box="[1150,1255,1364,1388]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="13" pageNumber="464" unit="mm" value="11.2">11.2 mm</quantity>
) (NMNS-7779- 014),
<collectingCountry id="C7D4EACBFFA9FF8BD180FA804502FA4A" box="[218,304,1399,1424]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD066FA804597FA4A" box="[316,421,1399,1424]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
: mouth of
<location id="BA1CFC80FFA9FF8BD37BFA804691FA4A" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD37BFA804691FA4A" box="[545,675,1399,1424]" country="Taiwan" name="Paoli River" pageId="13" pageNumber="464" stateProvince="Pingtung">Paoli River</location>
,
<collectingDate id="DB397573FFA9FF8BD3F5FA8F4751FA4A" box="[687,867,1399,1424]" pageId="13" pageNumber="464" value="2016-10-04">
<date id="CB7D8C9BFFA9FF8BD3F5FA8F4751FA4A" box="[687,867,1399,1424]" pageId="13" pageNumber="464" value="2016-10-04">4 October 2016</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD2D1FA8F47F7FA4A" box="[907,965,1399,1424]" pageId="13" pageNumber="464">J. Li.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD2BCFA8044C5FA02" ID-GBIF-Occurrence="1913889741" collectingDate="2015-09-26" collectorName="J. Li. Other" country="Taiwan" location="Kangkou River" municipality="Kenting National Park" pageId="13" pageNumber="464" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD2BCFA80401FFA4A" box="[998,1069,1399,1424]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(12.4×
<quantity id="783B07BEFFA9FF8BD5DAFA8F40DAFA4A" box="[1152,1256,1400,1424]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="13" pageNumber="464" unit="mm" value="10.8">10.8 mm</quantity>
) (NMNS-7779- 007),
<collectingCountry id="C7D4EACBFFA9FF8BD186FA6B4500FA6F" box="[220,306,1436,1461]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD01BFA6B4598FA6F" box="[321,426,1436,1461]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="464">Pingtung</collectingRegion>
:
<collectingMunicipality id="5F183021FFA9FF8BD0E3FA6B46F0FA6F" box="[441,706,1436,1461]" pageId="13" pageNumber="464">Kenting National Park</collectingMunicipality>
: mouth of
<location id="BA1CFC80FFA9FF8BD21FFA6B47C4FA6F" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD21FFA6B47C4FA6F" box="[837,1014,1436,1461]" country="Taiwan" municipality="Kenting National Park" name="Kangkou River" pageId="13" pageNumber="464" stateProvince="Pingtung">Kangkou River</location>
,
<collectingDate id="DB397573FFA9FF8BD55CFA6B40D8FA6F" box="[1030,1258,1436,1461]" pageId="13" pageNumber="464" value="2015-09-26">
<date id="CB7D8C9BFFA9FF8BD55CFA6B40D8FA6F" box="[1030,1258,1436,1461]" pageId="13" pageNumber="464" value="2015-09-26">26 September 2015</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFA9FF8BD44EFA6A41AEFA6F" box="[1300,1436,1436,1461]" pageId="13" pageNumber="464">J. Li. Other</collectorName>
material
</materialsCitation>
: 2 males (12.7×
<quantity id="783B07BEFFA9FF8BD0FCFA37463EFA0D" box="[422,524,1472,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="13" pageNumber="464" unit="mm" value="10.8">10.8 mm</quantity>
, 9.5×
<quantity id="783B07BEFFA9FF8BD310FA484680FA02" box="[586,690,1471,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="13" pageNumber="464" unit="mm" value="10.9">10.9 mm</quantity>
),
<materialsCitation id="0FABA006FFA9FF8BD39EFA374020FA27" ID-GBIF-Occurrence="1913889744" collectingDate="2004-05-18" collectorName="C. Liu." country="Philippines" location="Loboc River" pageId="13" pageNumber="464" specimenCount="3" specimenCount-female="3" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD39EFA374702FA02" box="[708,816,1471,1496]" pageId="13" pageNumber="464" type="female">3 females</specimenCount>
(12.3×
<quantity id="783B07BEFFA9FF8BD225FA3747D4FA0D" box="[895,998,1472,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="13" pageNumber="464" unit="mm" value="10.8">10.8 mm</quantity>
, 11.8×10.0 mm, 8.4×
<quantity id="783B07BEFFA9FF8BD589FA484118FA02" box="[1235,1322,1471,1496]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="464" unit="mm" value="7.5">7.5 mm</quantity>
) (NMNS- 7779-020),
<collectingCountry id="C7D4EACBFFA9FF8BD046FA1345ADFA27" box="[284,415,1508,1533]" name="Philippines" pageId="13" pageNumber="464">Philippines</collectingCountry>
:
<collectingRegion id="7D0764B9FFA9FF8BD0F6FA1345C2FA27" box="[428,496,1508,1533]" country="Philippines" name="Bohol" pageId="13" pageNumber="464">Bohol</collectingRegion>
Island:
<location id="BA1CFC80FFA9FF8BD314FA1346EDFA27" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD314FA1346EDFA27" box="[590,735,1508,1533]" country="Philippines" name="Loboc River" pageId="13" pageNumber="464" stateProvince="Bohol">Loboc River</location>
,
<collectingDate id="DB397573FFA9FF8BD3B4FA1347BEFA26" box="[750,908,1508,1533]" pageId="13" pageNumber="464" value="2004-05-18">
<date id="CB7D8C9BFFA9FF8BD3B4FA1347BEFA26" box="[750,908,1508,1533]" pageId="13" pageNumber="464" value="2004-05-18">18 May 2004</date>
</collectingDate>
, H.-
<collectorName id="1236CF8DFFA9FF8BD2E5FA134020FA27" box="[959,1042,1508,1533]" pageId="13" pageNumber="464">C. Liu.</collectorName>
</materialsCitation>
<materialsCitation id="0FABA006FFA9FF8BD56BFA134739F9FA" ID-GBIF-Occurrence="1913889738" collectingDate="2016-03-05" collectorName="B. Chan." country="Philippines" location="Mindoro" pageId="13" pageNumber="464" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFA9FF8BD56BFA134048FA27" box="[1073,1146,1508,1533]" pageId="13" pageNumber="464" type="male">1 male</specimenCount>
(9.9×8.4) (NMNS-7779- 011),
<collectingCountry id="C7D4EACBFFA9FF8BD18DF9F04568F9FA" box="[215,346,1543,1568]" name="Philippines" pageId="13" pageNumber="464">Philippines</collectingCountry>
:
<location id="BA1CFC80FFA9FF8BD03EF9F045FBF9FA" LSID="urn:lsid:plazi:treatment:376A1B4DFFA9FF89D1CDFC534107FE9A:BA1CFC80FFA9FF8BD03EF9F045FBF9FA" box="[356,457,1543,1568]" country="Philippines" name="Mindoro" pageId="13" pageNumber="464">Mindoro</location>
,
<collectingDate id="DB397573FFA9FF8BD08EF9F04646F9FA" box="[468,628,1543,1568]" pageId="13" pageNumber="464" value="2016-03-05">
<date id="CB7D8C9BFFA9FF8BD08EF9F04646F9FA" box="[468,628,1543,1568]" pageId="13" pageNumber="464" value="2016-03-05">5 March 2016</date>
</collectingDate>
, W.-
<collectorName id="1236CF8DFFA9FF8BD3F0F9FF4739F9FA" box="[682,779,1543,1568]" pageId="13" pageNumber="464">B. Chan.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA9FF88D19DF9DB40D9F9DC" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464" type="diagnosis">
<paragraph id="BF7CAA5BFFA9FF8BD19DF9DB4022F82E" blockId="13.[151,1437,1039,2037]" pageId="13" pageNumber="464">
<emphasis id="8DB77649FFA9FF8BD19DF9DB4573F99F" bold="true" box="[199,321,1580,1605]" pageId="13" pageNumber="464">Diagnosis.</emphasis>
Carapace (
<figureCitation id="27F8B6DEFFA9FF8BD098F9DB4675F99E" box="[450,583,1580,1605]" captionStart="FIGURE 7" captionStartId="14.[151,250,1353,1375]" captionTargetBox="[152,1435,193,1332]" captionTargetId="figure@14.[152,1435,193,1332]" captionTargetPageId="14" captionText="FIGURE 7. Parasesarma macaco n. sp.. AD, holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; E, F, paratype female (13.3×11.1 mm) (NMNS-7779-013), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440723/files/figure.png" pageId="13" pageNumber="464">Figs. 7A, E</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD30EF9DB464AF99E" box="[596,632,1580,1604]" captionStart="FIGURE 8" captionStartId="15.[151,250,1572,1594]" captionTargetBox="[196,1307,452,1520]" captionTargetId="figure@15.[152,1435,375,1550]" captionTargetPageId="15" captionText="FIGURE 8. Parasesarma macaco n. sp., holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan. A, dorsal view of carapace; BD, tip of left cheliped; E, F, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, dorsal (sternal) view; F, ventral (pleonal) view. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440725/files/figure.png" pageId="13" pageNumber="464">8A</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD3DCF9DB4687F99E" box="[646,693,1580,1604]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="13" pageNumber="464">22B</figureCitation>
) squarish in general outline, 1.1 times broader than long; regions well defined, separated by deep grooves; postfrontal region distinct, separated into 4 lobes by deep grooves; front deflexed downwards (
<figureCitation id="27F8B6DEFFA9FF8BD0D0F9834638F956" box="[394,522,1652,1677]" captionStart="FIGURE 7" captionStartId="14.[151,250,1353,1375]" captionTargetBox="[152,1435,193,1332]" captionTargetId="figure@14.[152,1435,193,1332]" captionTargetPageId="14" captionText="FIGURE 7. Parasesarma macaco n. sp.. AD, holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; E, F, paratype female (13.3×11.1 mm) (NMNS-7779-013), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440723/files/figure.png" pageId="13" pageNumber="464">Figs. 7C, D</figureCitation>
) margin intensely concave in dorsal view; lateral margin straight, subparallel along most of length before curving to join almost straight posterior carapace margin; cornea extending or just reaching tip of external orbital tooth (Figs. 77A, E, 8A, 22B). Ischium of third maxilliped with shallow median sulcus, merus with distinct submedian ridge; exopod slender, tip reaching half-length of outer margin of merus, flagellum long. Male cheliped palm with 2 transverse pectinate crests (13 and 10 corneous teeth, respectively) on upper surface; upper surface of dactylus with 6 to 8 (average 7) symmetrical, obliquely elongate dactylar tubercles, proximal 3 tubercles steep, sharp, the others tubercles large, distalmost 2 tubercles indistinct (
<figureCitation id="27F8B6DEFFA9FF8BD2E5F8BB4012F8BE" box="[959,1056,1868,1893]" captionStart="FIGURE 8" captionStartId="15.[151,250,1572,1594]" captionTargetBox="[196,1307,452,1520]" captionTargetId="figure@15.[152,1435,375,1550]" captionTargetPageId="15" captionText="FIGURE 8. Parasesarma macaco n. sp., holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan. A, dorsal view of carapace; BD, tip of left cheliped; E, F, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, dorsal (sternal) view; F, ventral (pleonal) view. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440725/files/figure.png" pageId="13" pageNumber="464">Figs. 8C</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD577F8BB406DF8BE" box="[1069,1119,1868,1892]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="13" pageNumber="464">19C</figureCitation>
). Ambulatory legs relatively slender for this species-complex, P3 and P4 about 1.5 times carapace width; P3 and P4 coxae without dense setae; P3 merus 2.5 times as long as broad; P3 propodus 5.0 times as long as broad; P3 dactylus 0.4 times length of propodus (
<figureCitation id="27F8B6DEFFA9FF8BD1C5F84F4536F80A" box="[159,260,1976,2001]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="13" pageNumber="464">Fig. 20B</figureCitation>
). G1 relatively slender; apical process corneous, elongated, slightly bent at angle of 45°, long, stout, ending in rounded tip (
<figureCitation id="27F8B6DEFFA9FF8BD065F82B458AF82E" box="[319,440,2012,2037]" captionStart="FIGURE 8" captionStartId="15.[151,250,1572,1594]" captionTargetBox="[196,1307,452,1520]" captionTargetId="figure@15.[152,1435,375,1550]" captionTargetPageId="15" captionText="FIGURE 8. Parasesarma macaco n. sp., holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan. A, dorsal view of carapace; BD, tip of left cheliped; E, F, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, dorsal (sternal) view; F, ventral (pleonal) view. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440725/files/figure.png" pageId="13" pageNumber="464">Figs. 8E, F</figureCitation>
,
<figureCitation id="27F8B6DEFFA9FF8BD099F82B45C6F82E" box="[451,500,2012,2036]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="13" pageNumber="464">21C</figureCitation>
). G2 longer than quarter length of G1 (
<figureCitation id="27F8B6DEFFA9FF8BD2C5F82B4031F82E" box="[927,1027,2012,2037]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="13" pageNumber="464">Fig. 21C</figureCitation>
).
</paragraph>
<caption id="EBBCFAD3FFAAFF88D1CDFABE41AEFAA4" httpUri="https://zenodo.org/record/1440723/files/figure.png" pageId="14" pageNumber="465" startId="14.[151,250,1353,1375]" targetBox="[152,1435,193,1332]" targetPageId="14">
<paragraph id="BF7CAA5BFFAAFF88D1CDFABE41AEFAA4" blockId="14.[151,1436,1353,1406]" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D1CDFABE453DFA85" bold="true" box="[151,271,1353,1375]" pageId="14" pageNumber="465">FIGURE 7</emphasis>
.
<taxonomicName id="78C3D1D8FFAAFF88D047FABD45C9FA85" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[285,507,1354,1375]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFAAFF88D047FABD45C9FA85" box="[285,507,1354,1375]" italics="true" pageId="14" pageNumber="465">Parasesarma macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFAAFF88D359FABD460CFA85" box="[515,574,1354,1375]" pageId="14" pageNumber="465" rank="species">
<emphasis id="8DB77649FFAAFF88D359FABD460CFA85" bold="true" box="[515,574,1354,1375]" pageId="14" pageNumber="465">n. sp.</emphasis>
</taxonomicNameLabel>
. AD, holotype male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; E, F, paratype female (13.3×11.1 mm) (NMNS-7779-013), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views.
</paragraph>
</caption>
<paragraph id="BF7CAA5BFFAAFF88D19DFA5240D9F9DC" blockId="14.[151,1437,1445,2010]" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D19DFA52455FFA64" bold="true" box="[199,365,1445,1470]" pageId="14" pageNumber="465">Colour in life</emphasis>
. In large individuals, the carapace is dark brown, mottled with darker and lighter blotches; females and juveniles occasionally have light orange blotches on the gastric region (
<figureCitation id="27F8B6DEFFAAFF88D521FA3E40D4FA38" box="[1147,1254,1481,1506]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. AD, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="14" pageNumber="465">Fig. 25C</figureCitation>
). Chelipeds are yellowish orange, but brighter in males. Ambulatory legs are gray to brown (
<figureCitation id="27F8B6DEFFAAFF88D2A9FA1A4056F9DC" box="[1011,1124,1517,1542]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="14" pageNumber="465">Figs. 22B</figureCitation>
,
<figureCitation id="27F8B6DEFFAAFF88D52AFA194092F9DC" box="[1136,1184,1518,1542]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="14" pageNumber="465">23B</figureCitation>
,
<figureCitation id="27F8B6DEFFAAFF88D5F6FA1940EDF9DC" box="[1196,1247,1518,1542]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="14" pageNumber="465">24C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFAAFF88D19DF9E647C8F960" pageId="14" pageNumber="465" type="discussion">
<paragraph id="BF7CAA5BFFAAFF88D19DF9E647C8F960" blockId="14.[151,1437,1445,2010]" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D19DF9E64509F9F0" bold="true" box="[199,315,1553,1578]" pageId="14" pageNumber="465">Remarks.</emphasis>
<taxonomicName id="78C3D1D8FFAAFF88D01BF9E44606F9F0" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[321,564,1555,1578]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFAAFF88D01BF9E44606F9F0" box="[321,564,1555,1578]" italics="true" pageId="14" pageNumber="465">Parasesarma macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFAAFF88D361F9E6464BF9F0" box="[571,633,1553,1578]" pageId="14" pageNumber="465" rank="species">
<emphasis id="8DB77649FFAAFF88D361F9E6464BF9F0" bold="true" box="[571,633,1553,1578]" pageId="14" pageNumber="465">n. sp.</emphasis>
</taxonomicNameLabel>
is most easily separated from other members of the species-complex by the medially concave frontal margin, and in having 68 dactylar tubercles of the chela (all tubercles are counted, including the small proximal and obsolescent distal ones). The specimens from
<collectingRegion id="7D0764B9FFAAFF88D574F9AE4040F9A8" box="[1070,1138,1625,1650]" country="Philippines" name="Bohol" pageId="14" pageNumber="465">Bohol</collectingRegion>
Island,
<collectingCountry id="C7D4EACBFFAAFF88D58AF9AE4163F9A8" box="[1232,1361,1625,1650]" name="Philippines" pageId="14" pageNumber="465">Philippines</collectingCountry>
, differ from the other specimens in having a slightly longer G1 but the tip is of the same shape. The features that distinguish this species from its closest congeners are summarized in
<tableCitation id="F2419FE0FFAAFF88D2FAF95647C6F960" box="[928,1012,1697,1722]" captionStart="TABLE 1" captionStartId="7.[147,224,586,606]" captionText="TABLE 1. Comparison of major differences between members of the Parasesarma leptosoma species-complex." pageId="14" pageNumber="465">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFAAFF88D19DF9324134F8D8" pageId="14" pageNumber="465" type="etymology">
<paragraph id="BF7CAA5BFFAAFF88D19DF9324134F8D8" blockId="14.[151,1437,1445,2010]" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D19DF932457CF904" bold="true" box="[199,334,1733,1758]" pageId="14" pageNumber="465">Etymology.</emphasis>
Derived from the Portuguese &quot;
<taxonomicName id="78C3D1D8FFAAFF88D3EEF9304739F904" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[692,779,1735,1758]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macaco">
<emphasis id="8DB77649FFAAFF88D3EEF9304739F904" box="[692,779,1735,1758]" italics="true" pageId="14" pageNumber="465">macaco</emphasis>
</taxonomicName>
&quot; meaning “monkey”. It alludes to the agility of this treeclimbing species and its habit of jumping around branches. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFAAFF88D19DF8FA45F5F890" pageId="14" pageNumber="465" type="distribution">
<paragraph id="BF7CAA5BFFAAFF88D19DF8FA45F5F890" blockId="14.[151,1437,1445,2010]" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D19DF8FA4552F8FC" bold="true" box="[199,352,1805,1830]" pageId="14" pageNumber="465">Distribution.</emphasis>
<collectingCountry id="C7D4EACBFFAAFF88D03DF8FA458CF8FC" box="[359,446,1805,1830]" name="Taiwan" pageId="14" pageNumber="465">Taiwan</collectingCountry>
: Kangkou River and Paoli River (
<collectingRegion id="7D0764B9FFAAFF88D262F8FA47C6F8FC" box="[824,1012,1805,1830]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="465">Pingtung county</collectingRegion>
) and Sicao (
<collectingRegion id="7D0764B9FFAAFF88D525F8FA40FFF8FC" box="[1151,1229,1805,1830]" country="Taiwan" name="Tainan" pageId="14" pageNumber="465">Tainan</collectingRegion>
city);
<collectingCountry id="C7D4EACBFFAAFF88D44CF8FA41ABF8FC" box="[1302,1433,1805,1830]" name="Philippines" pageId="14" pageNumber="465">Philippines</collectingCountry>
: Mindoro and
<collectingRegion id="7D0764B9FFAAFF88D075F8C64541F890" box="[303,371,1841,1866]" country="Philippines" name="Bohol" pageId="14" pageNumber="465">Bohol</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFAAFF89D19DF8A24107FE9A" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465" type="biology_ecology">
<paragraph id="BF7CAA5BFFAAFF89D19DF8A24108FF22" blockId="14.[151,1437,1445,2010]" lastBlockId="15.[151,1436,151,320]" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465">
<emphasis id="8DB77649FFAAFF88D19DF8A24519F8B4" bold="true" box="[199,299,1877,1902]" pageId="14" pageNumber="465">Ecology.</emphasis>
<taxonomicName id="78C3D1D8FFAAFF88D069F8A04614F8B4" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[307,550,1879,1902]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFAAFF88D069F8A04614F8B4" box="[307,550,1879,1902]" italics="true" pageId="14" pageNumber="465">Parasesarma macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFAAFF88D374F8A2465FF8B4" box="[558,621,1877,1902]" pageId="14" pageNumber="465" rank="species">
<emphasis id="8DB77649FFAAFF88D374F8A2465FF8B4" bold="true" box="[558,621,1877,1902]" pageId="14" pageNumber="465">n. sp.</emphasis>
</taxonomicNameLabel>
is a common species in Paoli River,
<collectingCountry id="C7D4EACBFFAAFF88D555F8A24057F8B4" box="[1039,1125,1877,1902]" name="Taiwan" pageId="14" pageNumber="465">Taiwan</collectingCountry>
. In
<collectingCountry id="C7D4EACBFFAAFF88D5D4F8A240D2F8B4" box="[1166,1248,1877,1902]" name="Taiwan" pageId="14" pageNumber="465">Taiwan</collectingCountry>
and
<collectingCountry id="C7D4EACBFFAAFF88D44DF8A241AAF8B4" box="[1303,1432,1877,1902]" name="Philippines" pageId="14" pageNumber="465">Philippines</collectingCountry>
, most of specimens of
<taxonomicName id="78C3D1D8FFAAFF88D0D0F88C45CEF848" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[394,508,1915,1938]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="macaco">
<emphasis id="8DB77649FFAAFF88D0D0F88C45CEF848" box="[394,508,1915,1938]" italics="true" pageId="14" pageNumber="465">P. macaco</emphasis>
</taxonomicName>
were collected at night, with specimens observed climbing the trees along the river banks of the estuary. The common host plants are
<taxonomicName id="78C3D1D8FFAAFF88D3B6F86A47F5F86C" box="[748,967,1949,1974]" class="Magnoliopsida" family="Malvaceae" genus="Talipariti" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="465" phylum="Tracheophyta" rank="species" species="tiliaceum">
<emphasis id="8DB77649FFAAFF88D3B6F86A47F5F86C" box="[748,967,1949,1974]" italics="true" pageId="14" pageNumber="465">Talipariti tiliaceum</emphasis>
</taxonomicName>
(L.) Fryxell,
<taxonomicName id="78C3D1D8FFAAFF88D531F86A4154F86F" box="[1131,1382,1949,1974]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="465" phylum="Tracheophyta" rank="species" species="inerme">
<emphasis id="8DB77649FFAAFF88D531F86A4154F86F" box="[1131,1382,1949,1974]" italics="true" pageId="14" pageNumber="465">Clerodendrum inerme</emphasis>
</taxonomicName>
(L.) Gaertn. (in
<collectingCountry id="C7D4EACBFFAAFF88D07AF8364546F800" box="[288,372,1985,2010]" name="Taiwan" pageId="14" pageNumber="465">Taiwan</collectingCountry>
) (
<figureCitation id="27F8B6DEFFAAFF88D0D5F836461AF800" box="[399,552,1985,2010]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. AD, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="14" pageNumber="465">Figs. 25AC</figureCitation>
),
<taxonomicName id="78C3D1D8FFAAFF88D31AF834470CF800" box="[576,830,1985,2010]" class="Magnoliopsida" family="Rhizophoraceae" genus="Bruguiera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="14" pageNumber="465" phylum="Tracheophyta" rank="species" species="gymnorhiza">
<emphasis id="8DB77649FFAAFF88D31AF834470CF800" box="[576,830,1985,2010]" italics="true" pageId="14" pageNumber="465">Bruguiera gymnorhiza</emphasis>
</taxonomicName>
(L.) Lam. and
<taxonomicName id="78C3D1D8FFAAFF88D2AFF8354140F800" authority="(Forsk.) Vierh." authorityName="(Forsk.) Vierh." box="[1013,1394,1985,2010]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="465" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="8DB77649FFAAFF88D2AFF83540F2F800" box="[1013,1216,1986,2010]" italics="true" pageId="14" pageNumber="465">Avicennia marina</emphasis>
(Forsk.) Vierh.
</taxonomicName>
(in
<collectingCountry id="C7D4EACBFFABFF89D1CDFF604528FF6A" box="[151,282,151,176]" name="Philippines" pageId="15" pageNumber="466">Philippines</collectingCountry>
). When the water levels (from the rising tide and rainfall) reach the host plant, the crabs would climb downward and submerge in the water.
<taxonomicName id="78C3D1D8FFABFF89D304FF4A4767FF0E" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[606,853,189,212]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFABFF89D304FF4A4767FF0E" box="[606,853,189,212]" italics="true" pageId="15" pageNumber="466">Parasesarma macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFABFF89D23AFF4B4791FF0F" box="[864,931,188,213]" pageId="15" pageNumber="466" rank="species">
<emphasis id="8DB77649FFABFF89D23AFF4B4791FF0F" bold="true" box="[864,931,188,213]" pageId="15" pageNumber="466">n. sp.</emphasis>
</taxonomicNameLabel>
rarely ventures onto the ground except for breeding (
<figureCitation id="27F8B6DEFFABFF89D052FF284543FF22" box="[264,369,223,248]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. AD, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="15" pageNumber="466">Fig. 25D</figureCitation>
). Ovigerous females were found almost all year around, but mainly in July to October.
</paragraph>
<paragraph id="BF7CAA5BFFABFF89D19DFEF24107FE9A" blockId="15.[151,1436,151,320]" pageId="15" pageNumber="466">
<taxonomicName id="78C3D1D8FFABFF89D19DFEF2458CFEC6" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[199,446,261,284]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFABFF89D19DFEF2458CFEC6" box="[199,446,261,284]" italics="true" pageId="15" pageNumber="466">Parasesarma macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFABFF89D092FEF34639FEC7" box="[456,523,260,285]" pageId="15" pageNumber="466" rank="species">
<emphasis id="8DB77649FFABFF89D092FEF34639FEC7" bold="true" box="[456,523,260,285]" pageId="15" pageNumber="466">n. sp.</emphasis>
</taxonomicNameLabel>
occurs sympatrically with
<taxonomicName id="78C3D1D8FFABFF89D213FEF247B8FEC7" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[841,906,260,285]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">
<emphasis id="8DB77649FFABFF89D213FEF247B8FEC7" box="[841,906,260,285]" italics="true" pageId="15" pageNumber="466">P. kui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFABFF89D2CFFEF347E5FEC7" box="[917,983,260,285]" pageId="15" pageNumber="466" rank="species">
<emphasis id="8DB77649FFABFF89D2CFFEF347E5FEC7" bold="true" box="[917,983,260,285]" pageId="15" pageNumber="466">n. sp.</emphasis>
</taxonomicNameLabel>
(
<collectingCountry id="C7D4EACBFFABFF89D2B0FEF34072FEC7" box="[1002,1088,260,285]" name="Taiwan" pageId="15" pageNumber="466">Taiwan</collectingCountry>
) and
<taxonomicName id="78C3D1D8FFABFF89D5DFFEF241AEFEC7" baseAuthorityName="White" baseAuthorityYear="1847" box="[1157,1436,260,285]" class="Malacostraca" family="Grapsidae" genus="Metopograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="8DB77649FFABFF89D5DFFEF241AEFEC7" box="[1157,1436,260,285]" italics="true" pageId="15" pageNumber="466">Metopograpsus latifrons</emphasis>
</taxonomicName>
(
<collectingCountry id="C7D4EACBFFABFF89D1C4FED044C0FE9A" box="[158,242,295,320]" name="Taiwan" pageId="15" pageNumber="466">Taiwan</collectingCountry>
and
<collectingCountry id="C7D4EACBFFABFF89D073FED04599FE9A" box="[297,427,295,320]" name="Philippines" pageId="15" pageNumber="466">Philippines</collectingCountry>
), the niche and ecology appear to be generally similar (see “Ecology” for
<taxonomicName id="78C3D1D8FFABFF89D5BCFEDE4113FE9A" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[1254,1313,295,320]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="kui">
<emphasis id="8DB77649FFABFF89D5BCFEDE4113FE9A" box="[1254,1313,295,320]" italics="true" pageId="15" pageNumber="466">P. kui</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>