325 lines
38 KiB
XML
325 lines
38 KiB
XML
<document id="2BAFFD15F892E32125BEBE2F4C0064E0" ID-DOI="10.1649/0010-065X-69.mo4.55" ID-ISSN="1938-4394" ID-Zenodo-Dep="4908445" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623096069005" checkinUser="carolina" docAuthor="Smith, Aaron D. & Cifuentes-Ruiz, Paulina" docDate="2015" docId="372E87BF25005D39FF0DFD12FB738031" docLanguage="en" docName="TheColeopBull.69.mo14.55-72.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 14) 69" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Diceroderes cusucoensis Smith 2015, new species" docType="treatment" docVersion="3" lastPageNumber="63" masterDocId="CB17FFC725075D31FFA1FFD5FFE68613" masterDocTitle="Revision of Diceroderes Solier (Coleoptera: Tenebrionidae: Toxicini: Eudysantina), with Descriptions of Four New Species" masterLastPageNumber="72" masterPageNumber="55" pageNumber="62" updateTime="1698956517230" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="BED91B01A389BCDB2ED477DCCC386D78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AC4C88751AEE925D6DD70B44A5D7AC96">
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<mods:title id="2D08EE16555D9211BBB7A63FAF2F200D">Revision of Diceroderes Solier (Coleoptera: Tenebrionidae: Toxicini: Eudysantina), with Descriptions of Four New Species</mods:title>
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</mods:titleInfo>
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<mods:name id="82A67AFC5B3AFE8D46F91584742BF12C" type="personal">
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<mods:role id="983F4EE02C4297EABF5AF9BF716D1D28">
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<mods:roleTerm id="D585447A61B09D14EE4B5FAF386B5A82">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8624D65E27225E39F4E097B85E4673A4">Smith, Aaron D.</mods:namePart>
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<mods:affiliation id="AEAB1ECD8D84A9961331938FD45A632A">Department of Biological Sciences, Northern Arizona University PO Box 5640, Flagstaff, AZ, 86011 - 5640, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="4782C14B8CAC743AB04A82D26EB2DA78" type="email">aaron.smith@nau.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="9DCDE2EC416F00945511E1BCE9F07F9E" type="personal">
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<mods:role id="092C34A317606CE26FDCAF53B7521BC1">
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<mods:roleTerm id="3CB57919967D03BD018E0AB332CF2765">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="38A4DDC17E71E17BC8710951AC939E5C">Cifuentes-Ruiz, Paulina</mods:namePart>
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<mods:affiliation id="499CF537B240810918373263DC46C94C">Instituto de Biología, Departamento de Zoología Universidad Nacional Autónoma de México, Apartado Postal 70 - 153 Delegación Coyoacán, C. P. 04510, DF, MEXICO</mods:affiliation>
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<mods:nameIdentifier id="916C1F26387A8B8051C3ADC9E9DDB6F1" type="email">paulinacifruz@yahoo.com.mx</mods:nameIdentifier>
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<mods:typeOfResource id="4CBCB03C3D4AD599D8ADDF1334F3739D">text</mods:typeOfResource>
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<mods:titleInfo id="FE0DEFD1B702C5586039C4AD38EE395A">
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<mods:title id="07E12BA689EA4CB0554D416255CF84E6">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="E040ED14A3C7C0621BD43D74C9909D7D">
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<mods:date id="38641D6ED17D5A3CFF47FD48EE0A3FA5">2015</mods:date>
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<mods:detail id="C5E84B78D2728F3213AD91149DCA8D3B" type="series">
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<mods:title id="3F7DB6E13697708AE1C0039C1563C1E6">mo 14</mods:title>
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</mods:detail>
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<mods:number id="F53CA4452B6CF23E177AF9638FD5C2B5">2015-12-18</mods:number>
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<mods:detail id="E3AD6F74114A642AFFEC21E96ED32632" type="volume">
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<mods:number id="BE76151D0836DDEFDED5EFCA0C24A342">69</mods:number>
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<mods:start id="82FC1D6E8EBA46DA91C11D756C9DB096">55</mods:start>
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<mods:classification id="F4912BA28F9AACE2C5A8D2B7FAE674C5">journal article</mods:classification>
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<mods:identifier id="D31C6C8FF3155730BC30525F59A4BCF2" type="DOI">10.1649/0010-065X-69.mo4.55</mods:identifier>
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<mods:identifier id="BB960415CE808CCF784D29AF73E600A9" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="F79D652225005D36FF0DFD12FD8E84CE" box="[172,616,711,733]" pageId="7" pageNumber="62" type="nomenclature">
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<paragraph id="BF3836A925005D36FF0DFD12FD8E84CE" blockId="7.[172,616,711,760]" box="[172,616,711,733]" pageId="7" pageNumber="62">
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<heading id="E47081C525005D36FF0DFD12FD8E84CE" box="[172,616,711,733]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="62" reason="2">
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<emphasis id="8DF3EABB25005D36FF0DFD12FD8E84CE" bold="true" box="[172,616,711,733]" pageId="7" pageNumber="62">
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<taxonomicName id="78874D2A25005D36FF0DFD12FE0384CE" authority="Smith" authorityName="Smith" authorityYear="2015" box="[172,485,711,733]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cusucoensis" status="sp. nov.">
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<emphasis id="8DF3EABB25005D36FF0DFD12FE7884CE" bold="true" box="[172,414,711,733]" italics="true" pageId="7" pageNumber="62">Diceroderes cusucoensis</emphasis>
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Smith
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</taxonomicName>
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,
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<taxonomicNameLabel id="96C057C025005D36FE53FD1DFD8E84CE" box="[498,616,712,733]" pageId="7" pageNumber="62" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<caption id="EBF8662125005D36FD64FD17FB7584C6" ID-DOI="http://doi.org/10.5281/zenodo.4908469" ID-Zenodo-Dep="4908469" box="[709,1171,705,725]" httpUri="https://zenodo.org/record/4908469/files/figure.png" pageId="7" pageNumber="62" startId="7.[709,751,706,725]" targetBox="[685,1171,194,682]" targetPageId="7">
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<paragraph id="BF3836A925005D36FD64FD17FB7584C6" blockId="7.[685,1171,705,749]" box="[709,1171,705,725]" pageId="7" pageNumber="62">
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<emphasis id="8DF3EABB25005D36FD64FD17FCDB84C6" bold="true" box="[709,829,705,725]" pageId="7" pageNumber="62">Figs. 26–27.</emphasis>
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<taxonomicName id="78874D2A25005D36FCF5FD14FBDF84C7" authorityName="Smith" authorityYear="2015" box="[852,1081,705,724]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cusucoensis">
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<emphasis id="8DF3EABB25005D36FCF5FD14FBDF84C7" box="[852,1081,705,724]" italics="true" pageId="7" pageNumber="62">Diceroderes cusucoensis</emphasis>
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</taxonomicName>
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, allotype
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="F79D652225005D36FEBCFD36FE1E84EB" box="[285,504,738,760]" pageId="7" pageNumber="62" type="description">
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<paragraph id="BF3836A925005D36FEBCFD36FE1E84EB" blockId="7.[172,616,711,760]" box="[285,504,738,760]" pageId="7" pageNumber="62">
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(
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<figureCitation id="27BC2A2C25005D36FE85FD36FE6584EB" box="[292,387,738,760]" captionStart="Figs" captionStartId="3.[175,215,577,596]" captionTargetBox="[157,1165,202,555]" captionTargetId="figure-539@3.[151,1173,183,559]" captionTargetPageId="3" captionText="Figs. 3–12. Diceroderes species, aedeagi, dorsal and lateral views. 3–4) D. mexicanus, TB# 14740; 5–6) D. cusucoensis, holotype, TB# 14688; 7–8) D. ocozocoautlaensis, holotype, TB# 14689; 9–10) D. skelleyi, holotype, TB# 13105; 11–12) D. subtriplehorni, holotype, TB# 13109." figureDoi="http://doi.org/10.5281/zenodo.4908451" httpUri="https://zenodo.org/record/4908451/files/figure.png" pageId="7" pageNumber="62">Figs. 5–6</figureCitation>
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,
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<figureCitation id="27BC2A2C25005D36FE2EFD36FE4F84EB" box="[399,425,739,760]" captionStart="Fig" captionStartId="6.[197,229,774,793]" captionTargetBox="[173,1193,194,751]" captionTargetId="figure-447@6.[172,1194,193,751]" captionTargetPageId="6" captionText="Fig. 23. Diceroderes species, distribution map. Triangles = D. mexicanus, white triangles are for morph one, gray/blue triangles for morph two. Stars = D. cusucoensis. Squares = D. ocozocoautlaensis. Diamonds = D. skelleyi. Circles = D. subtriplehorni." figureDoi="http://doi.org/10.5281/zenodo.4908467" httpUri="https://zenodo.org/record/4908467/files/figure.png" pageId="7" pageNumber="62">23</figureCitation>
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,
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<figureCitation id="27BC2A2C25005D36FE15FD36FE1784EB" box="[436,497,738,760]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="7.[175,215,1686,1705]" captionStartId-1="7.[709,751,706,725]" captionTargetBox-0="[151,637,1153,1663]" captionTargetBox-1="[685,1171,194,682]" captionTargetId-0="figure-534@7.[151,638,1152,1663]" captionTargetId-1="figure-551@7.[685,1171,194,683]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figs. 24–25. Diceroderes cusucoensis, holotype (male)." captionText-1="Figs. 26–27. Diceroderes cusucoensis, allotype" figureDoi-0="http://doi.org/10.5281/zenodo.4908473" figureDoi-1="http://doi.org/10.5281/zenodo.4908469" httpUri-0="https://zenodo.org/record/4908473/files/figure.png" httpUri-1="https://zenodo.org/record/4908469/files/figure.png" pageId="7" pageNumber="62">24–27</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F79D652225005D36FF0EFCD8FBEB82DA" pageId="7" pageNumber="62" type="materials_examined">
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<paragraph id="BF3836A925005D36FF0EFCD8FD988255" blockId="7.[151,638,781,1095]" pageId="7" pageNumber="62">
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<emphasis id="8DF3EABB25005D36FF0EFCD8FEA08531" bold="true" box="[175,326,781,802]" pageId="7" pageNumber="62">Type Material.</emphasis>
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<materialsCitation id="0FEF3CF425005D36FEF5FCD8FE6F85CF" collectingDate="1994-08-26" collectionCode="CMNC" collectorName="N. Cofradia & W. Buenos Aires & R. Anderson & Aaron D. Smith" country="Honduras" county="Cusuco" elevation="1650" location="Cerro" municipality="Buenos Aires" pageId="7" pageNumber="62" specimenCount="2" stateProvince="Cortes" typeStatus="holotype">
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<typeStatus id="603C880B25005D36FEF5FCD8FE318531" box="[340,471,781,802]" pageId="7" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
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(male) labeled: (a) “
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<collectingCountry id="C790763925005D36FF6AFCFDFEBB852E" box="[203,349,808,829]" name="Honduras" pageId="7" pageNumber="62">HONDURAS</collectingCountry>
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:
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<collectingRegion id="7D43F84B25005D36FECDFCFDFE36852E" box="[364,464,808,829]" country="Honduras" name="Cortes" pageId="7" pageNumber="62">CORTES</collectingRegion>
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, P.N. /
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<collectingCounty id="56594E2525005D36FD89FCFDFD9F852E" box="[552,633,808,829]" pageId="7" pageNumber="62">Cusuco</collectingCounty>
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,
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<quantity id="787F9B4C25005D36FF36FC97FF028544" box="[151,228,834,855]" metricMagnitude="4" metricUnit="m" metricValue="1.8699999999999999" pageId="7" pageNumber="62" unit="km" value="18.7">18.7km</quantity>
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.
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<collectorName id="1272537F25005D36FF51FC97FE808544" box="[240,358,834,855]" pageId="7" pageNumber="62">N. Cofradia</collectorName>
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, /
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<quantity id="787F9B4C25005D36FE22FC97FE258544" box="[387,451,834,855]" metricMagnitude="3" metricUnit="m" metricValue="5.4" pageId="7" pageNumber="62" unit="km" value="5.4">5.4km</quantity>
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.
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<collectorName id="1272537F25005D36FE6EFC97FD9C8544" box="[463,634,834,855]" pageId="7" pageNumber="62">
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W.
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<collectingMunicipality id="5F5CACD325005D36FE53FC97FD9C8544" box="[498,634,834,855]" pageId="7" pageNumber="62">Buenos Aires</collectingMunicipality>
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</collectorName>
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,
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<location id="BA58607225005D36FF36FC88FF298561" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FF36FC88FF298561" box="[151,207,861,882]" country="Honduras" county="Cusuco" municipality="Buenos Aires" name="Cerro" pageId="7" pageNumber="62" stateProvince="Cortes">Cerro</location>
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/
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<location id="BA58607225005D36FF48FC88FEC88561" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FF48FC88FEC88561" box="[233,302,861,882]" country="Honduras" county="Cusuco" municipality="Buenos Aires" name="Jilinco" pageId="7" pageNumber="62" stateProvince="Cortes">Jilinco</location>
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,
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<quantity id="787F9B4C25005D36FE9AFC88FE648561" box="[315,386,861,882]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="62" unit="m" value="1650.0">
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<elevation id="34AAD19A25005D36FE9AFC88FE648561" box="[315,386,861,882]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="62" unit="m" value="1650.0">1650m</elevation>
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</quantity>
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,
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<date id="CB39106925005D36FE2EFC88FDF48561" box="[399,530,861,882]" pageId="7" pageNumber="62" value="1994-08-26">
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<collectingDate id="DB7DE98125005D36FE2EFC88FDF48561" box="[399,530,861,882]" pageId="7" pageNumber="62" value="1994-08-26">26.VIII.1994</collectingDate>
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</date>
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- / 225,
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<collectorName id="1272537F25005D36FDC6FC88FF12859F" pageId="7" pageNumber="62">R. Anderson</collectorName>
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, beating”; (b) on blue paper “Tenebrionid Base /
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<collectorName id="1272537F25005D36FF77FC47FE8E85B4" box="[214,360,914,935]" pageId="7" pageNumber="62">Aaron D. Smith</collectorName>
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/ Catalog # 14688”; (f) on red paper, “
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<typeStatus id="603C880B25005D36FF42FC78FEB985D1" box="[227,351,941,962]" pageId="7" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="78874D2A25005D36FED6FC78FE0185D1" authorityName="Solier" authorityYear="1841" box="[375,487,941,962]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Diceroderes</taxonomicName>
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/ cusucoensis / Smith 2015” (
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<collectionCode id="D996AE6C25005D36FE90FC12FE6485CF" box="[305,386,967,988]" pageId="7" pageNumber="62">CMNC</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="0FEF3CF425005D36FE39FC12FD8F8255" collectingDate="1994-08-26" collectingDateMax="1994-08-27" collectingDateMin="1994-08-26" collectorName="RD Cave & Tenebrionid Base & Aaron D. Smith & Catalog" country="Honduras" location="Cortes" municipality="Parq Nac Cusuco" pageId="7" pageNumber="62" specimenCount="1" typeStatus="allotype">
|
||
<typeStatus id="603C880B25005D36FE39FC12FDFD85CF" box="[408,539,967,988]" pageId="7" pageNumber="62" type="allotype">ALLOTYPE</typeStatus>
|
||
(female) labeled: (a) “
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||
<collectingCountry id="C790763925005D36FE85FC37FE5485E4" box="[292,434,994,1015]" name="Honduras" pageId="7" pageNumber="62">HONDURAS</collectingCountry>
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:
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<location id="BA58607225005D36FE1EFC37FDE685E4" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FE1EFC37FDE685E4" box="[447,512,994,1015]" country="Honduras" municipality="Parq Nac Cusuco" name="Cortes" pageId="7" pageNumber="62">Cortés</location>
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/
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<collectingMunicipality id="5F5CACD325005D36FDBFFC37FF398202" pageId="7" pageNumber="62">Parq Nac Cusuco</collectingMunicipality>
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/
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<locationDeviation id="AE8A2F8B25005D36FF59FC29FE3C8202" box="[248,474,1020,1041]" location="Buenos Aires" pageId="7" pageNumber="62">
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<quantity id="787F9B4C25005D36FF59FC29FECA8202" box="[248,300,1020,1041]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="62" unit="km" value="5.0">5 km</quantity>
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W Buenos Aires
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</locationDeviation>
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/
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<date id="CB39106925005D36FE53FC29FF2F823F" pageId="7" pageNumber="62" value="1994-08-26" valueMax="1994-08-27" valueMin="1994-08-26">
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<collectingDate id="DB7DE98125005D36FE53FC29FF2F823F" pageId="7" pageNumber="62" value="1994-08-26" valueMax="1994-08-27" valueMin="1994-08-26">26–27 August 1994</collectingDate>
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</date>
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/
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<collectorName id="1272537F25005D36FF47FBC2FEA3823F" box="[230,325,1047,1068]" pageId="7" pageNumber="62">RD Cave</collectorName>
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collector”; (b) on blue paper “
|
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<collectorName id="1272537F25005D36FF03FBE4FEB68255" box="[162,336,1073,1094]" pageId="7" pageNumber="62">Tenebrionid Base</collectorName>
|
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/
|
||
<collectorName id="1272537F25005D36FEC6FBE4FDE08255" box="[359,518,1073,1094]" pageId="7" pageNumber="62">Aaron D. Smith</collectorName>
|
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/
|
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<collectorName id="1272537F25005D36FDBCFBE4FD8F8255" box="[541,617,1073,1094]" pageId="7" pageNumber="62">Catalog</collectorName>
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</materialsCitation>
|
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#
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||
</paragraph>
|
||
<caption id="EBF8662125005D36FF0EF943FD9980BA" ID-DOI="http://doi.org/10.5281/zenodo.4908473" ID-Zenodo-Dep="4908473" box="[175,639,1686,1705]" httpUri="https://zenodo.org/record/4908473/files/figure.png" pageId="7" pageNumber="62" startId="7.[175,215,1686,1705]" targetBox="[151,637,1153,1663]" targetPageId="7">
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<paragraph id="BF3836A925005D36FF0EF943FD9980BA" blockId="7.[151,639,1686,1729]" box="[175,639,1686,1705]" pageId="7" pageNumber="62">
|
||
<emphasis id="8DF3EABB25005D36FF0EF943FEC680BA" bold="true" box="[175,288,1686,1705]" pageId="7" pageNumber="62">Figs. 24–25.</emphasis>
|
||
<taxonomicName id="78874D2A25005D36FE97F943FE1380BA" authorityName="Smith" authorityYear="2015" box="[310,501,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cusucoensis">
|
||
<emphasis id="8DF3EABB25005D36FE97F943FE1380BA" box="[310,501,1686,1705]" italics="true" pageId="7" pageNumber="62">Diceroderes cusucoensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype (male).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BF3836A925005D36FF36F97BFE0080D2" blockId="7.[151,639,1686,1729]" box="[151,486,1710,1729]" pageId="7" pageNumber="62">
|
||
<emphasis id="8DF3EABB25005D36FF36F97BFF5380D2" bold="true" box="[151,181,1710,1729]" pageId="7" pageNumber="62">24)</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis id="8DF3EABB25005D36FEE1F97BFEBB80D2" bold="true" box="[320,349,1710,1729]" pageId="7" pageNumber="62">25)</emphasis>
|
||
Lateral habitus.
|
||
</paragraph>
|
||
<paragraph id="BF3836A925005D36FD0CFD0FFB8E84FE" blockId="7.[685,1171,705,749]" box="[685,1128,730,749]" pageId="7" pageNumber="62">
|
||
(female).
|
||
<emphasis id="8DF3EABB25005D36FCA2FD0FFCC584FE" bold="true" box="[771,803,730,749]" pageId="7" pageNumber="62">26)</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis id="8DF3EABB25005D36FC16FD0FFC3084FE" bold="true" box="[951,982,730,749]" pageId="7" pageNumber="62">27)</emphasis>
|
||
Lateral habitus.
|
||
</paragraph>
|
||
<paragraph id="BF3836A925005D36FD0CFCF0FBEB82DA" blockId="7.[685,1172,805,1729]" pageId="7" pageNumber="62">
|
||
<materialsCitation id="0FEF3CF425005D36FD0CFCF0FBEC82DA" collectingDate="1990-08-10" collectionCode="ADSC, RDCC, BMNH, UVGC" collectorName="Aaron D. Smith & J. Monzon E. Smith" country="Guatemala" county="Smith" location="Catalog" municipality="Tenebrionid Base" pageId="7" pageNumber="62" specimenCount="8" specimenCount-female="1" specimenCount-male="3" stateProvince="Izabal" typeStatus="allotype">
|
||
14949”; (c) on red paper, “
|
||
<typeStatus id="603C880B25005D36FC38FCF0FBE88529" box="[921,1038,805,826]" pageId="7" pageNumber="62" type="allotype">ALLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="78874D2A25005D36FBBFFCF0FB6E8529" authorityName="Solier" authorityYear="1841" box="[1054,1160,805,826]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Diceroderes</taxonomicName>
|
||
/ cusucoensis / Smith 2015” (
|
||
<collectionCode id="D996AE6C25005D36FC1DFC95FBE68546" box="[956,1024,832,853]" pageId="7" pageNumber="62">ADSC</collectionCode>
|
||
).
|
||
<typeStatus id="603C880B25005D36FBB0FC95FB728546" box="[1041,1172,832,853]" pageId="7" pageNumber="62" type="paratype">PARATYPES</typeStatus>
|
||
(
|
||
<specimenCount id="A981FD2025005D36FD15FC8FFCC8857C" box="[692,814,858,879]" pageId="7" pageNumber="62" type="generic">4 specimens</specimenCount>
|
||
, excluding
|
||
<typeStatus id="603C880B25005D36FC03FC8FFC09857C" box="[930,1007,858,879]" pageId="7" pageNumber="62" type="allotype">allotype</typeStatus>
|
||
) (all bearing the label “
|
||
<typeStatus id="603C880B25005D36FD51FCA0FC818599" box="[752,871,885,906]" pageId="7" pageNumber="62" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="78874D2A25005D36FC23FCA0FC138599" authorityName="Solier" authorityYear="1841" box="[898,1013,885,906]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Diceroderes</taxonomicName>
|
||
/ cusucoensis /
|
||
<collectingCounty id="56594E2525005D36FD0CFC5AFD0285B7" box="[685,740,911,932]" pageId="7" pageNumber="62">Smith</collectingCounty>
|
||
2015” on yellow paper and the database label “
|
||
<collectingMunicipality id="5F5CACD325005D36FD19FC7FFCBB85AC" box="[696,861,938,959]" pageId="7" pageNumber="62">Tenebrionid Base</collectingMunicipality>
|
||
/
|
||
<collectorName id="1272537F25005D36FCD3FC7FFBED85AC" box="[882,1035,938,959]" pageId="7" pageNumber="62">Aaron D. Smith</collectorName>
|
||
/
|
||
<location id="BA58607225005D36FB81FC7FFB8E85AC" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FB81FC7FFB8E85AC" box="[1056,1128,938,959]" country="Guatemala" county="Smith" municipality="Tenebrionid Base" name="Catalog" pageId="7" pageNumber="62" stateProvince="Izabal">Catalog</location>
|
||
# ”, for convenience tenebrioniDBase catalog numbers are listed as TB # without quotations):
|
||
<location id="BA58607225005D36FB92FC0AFBBD85E7" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FB92FC0AFBBD85E7" box="[1075,1115,991,1012]" country="Guatemala" county="Smith" municipality="Tenebrionid Base" name="One" pageId="7" pageNumber="62" stateProvince="Izabal">One</location>
|
||
<typeStatus id="603C880B25005D36FBC5FC0AFD3E821C" pageId="7" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="D996AE6C25005D36FD4BFC2FFCD4821C" box="[746,818,1018,1039]" pageId="7" pageNumber="62">RDCC</collectionCode>
|
||
,
|
||
<specimenCount id="A981FD2025005D36FCE1FC2CFCB4821C" box="[832,850,1017,1039]" pageId="7" pageNumber="62" type="male">♂</specimenCount>
|
||
) labeled (a) same as
|
||
<typeStatus id="603C880B25005D36FB9DFC2FFB76821C" box="[1084,1168,1018,1039]" pageId="7" pageNumber="62" type="allotype">allotype</typeStatus>
|
||
; (b) TB #s 14951.
|
||
<location id="BA58607225005D36FCF5FBC1FC9D823A" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FCF5FBC1FC9D823A" box="[852,891,1044,1065]" country="Guatemala" county="Smith" municipality="Tenebrionid Base" name="One" pageId="7" pageNumber="62" stateProvince="Izabal">One</location>
|
||
<typeStatus id="603C880B25005D36FC20FBC1FC37823A" box="[897,977,1044,1065]" pageId="7" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="D996AE6C25005D36FC7FFBC1FBCE823A" box="[990,1064,1044,1065]" pageId="7" pageNumber="62">BMNH</collectionCode>
|
||
,
|
||
<specimenCount id="A981FD2025005D36FB90FBC1FBA58239" box="[1073,1091,1044,1066]" pageId="7" pageNumber="62" type="male">♂</specimenCount>
|
||
) labeled (a) same as
|
||
<typeStatus id="603C880B25005D36FC87FBFAFC928257" box="[806,884,1071,1092]" pageId="7" pageNumber="62" type="allotype">allotype</typeStatus>
|
||
; (b) TB #s 14952.
|
||
<location id="BA58607225005D36FB95FBFAFBBD8257" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FB95FBFAFBBD8257" box="[1076,1115,1071,1092]" country="Guatemala" county="Smith" municipality="Tenebrionid Base" name="One" pageId="7" pageNumber="62" stateProvince="Izabal">One</location>
|
||
<typeStatus id="603C880B25005D36FBC5FBFAFD33824D" pageId="7" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="D996AE6C25005D36FD47FB9CFCCE824D" box="[742,808,1097,1118]" pageId="7" pageNumber="62">ADSC</collectionCode>
|
||
,
|
||
<specimenCount id="A981FD2025005D36FC94FB9CFCA1824C" box="[821,839,1097,1119]" pageId="7" pageNumber="62" type="male">♂</specimenCount>
|
||
) labeled (a) same as
|
||
<typeStatus id="603C880B25005D36FBBFFB9CFB8A824D" box="[1054,1132,1097,1118]" pageId="7" pageNumber="62" type="allotype">allotype</typeStatus>
|
||
; (b) TB # 14950.
|
||
<location id="BA58607225005D36FC9AFBB1FC85826A" LSID="urn:lsid:plazi:treatment:372E87BF25005D39FF0DFD12FB738031:BA58607225005D36FC9AFBB1FC85826A" box="[827,867,1124,1145]" country="Guatemala" county="Smith" municipality="Tenebrionid Base" name="One" pageId="7" pageNumber="62" stateProvince="Izabal">One</location>
|
||
<typeStatus id="603C880B25005D36FCCCFBB1FC27826A" box="[877,961,1124,1145]" pageId="7" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="D996AE6C25005D36FC73FBB1FBFD826A" box="[978,1051,1124,1145]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="7" pageNumber="62">UVGC</collectionCode>
|
||
,
|
||
<specimenCount id="A981FD2025005D36FB88FBB6FBDD826A" box="[1065,1083,1123,1145]" pageId="7" pageNumber="62" type="female">♀</specimenCount>
|
||
) labeled (a) “
|
||
<collectingCountry id="C790763925005D36FD41FBABFC9B8280" box="[736,893,1150,1171]" name="Guatemala" pageId="7" pageNumber="62">GUATEMALA</collectingCountry>
|
||
.
|
||
<collectingRegion id="7D43F84B25005D36FC2CFBABFC2C8280" box="[909,970,1150,1171]" country="Guatemala" name="Izabal" pageId="7" pageNumber="62">Izabal</collectingRegion>
|
||
/ Los Amates, Sn Antonio / Sierra Espiritu Santo /
|
||
<date id="CB39106925005D36FC51FB4CFB6282BD" box="[1008,1156,1177,1198]" pageId="7" pageNumber="62" value="1990-08-10">
|
||
<collectingDate id="DB7DE98125005D36FC51FB4CFB6282BD" box="[1008,1156,1177,1198]" pageId="7" pageNumber="62" value="1990-08-10">10 agosto 1990</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="1272537F25005D36FD0CFB61FCBC82DA" box="[685,858,1204,1225]" pageId="7" pageNumber="62">J.Monzon E.Smith</collectorName>
|
||
”; (b) TB # 14953
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F79D652225005D36FD64FB1BFCAB83C1" pageId="7" pageNumber="62" type="diagnosis">
|
||
<paragraph id="BF3836A925005D36FD64FB1BFCAB83C1" blockId="7.[685,1172,805,1729]" pageId="7" pageNumber="62">
|
||
<emphasis id="8DF3EABB25005D36FD64FB1BFCC882F0" bold="true" box="[709,814,1230,1251]" pageId="7" pageNumber="62">Diagnosis.</emphasis>
|
||
<taxonomicName id="78874D2A25005D36FC9DFB1BFBA582F0" authorityName="Smith" authorityYear="2015" box="[828,1091,1230,1251]" class="Insecta" family="Tenebrionidae" genus="Diceroderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cusucoensis">
|
||
<emphasis id="8DF3EABB25005D36FC9DFB1BFBA582F0" box="[828,1091,1230,1251]" italics="true" pageId="7" pageNumber="62">Diceroderes cusucoensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from all other known members of the genus based on the following character combination: Supraorbital costae, transverse depressions, and medial tubercle on frons all absent; elytra in lateral view nearly straight from base to near apex then sharply declivous; prosternal process raised between coxae; males without apical spine on tibia; tubercles clothed in golden micro-pubescence, including apices.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F79D652225005D39FD64FA0DFB8A838E" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62" type="description">
|
||
<paragraph id="BF3836A925005D39FD64FA0DFBBC835E" blockId="7.[685,1172,805,1729]" lastBlockId="8.[707,1194,194,1570]" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">
|
||
<emphasis id="8DF3EABB25005D36FD64FA0DFC6F83FE" bold="true" box="[709,905,1496,1517]" pageId="7" pageNumber="62">Description. Male.</emphasis>
|
||
Length
|
||
<quantity id="787F9B4C25005D36FC7DFA0DFBB483FE" box="[988,1106,1495,1517]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.5" metricValueMin="8.6" pageId="7" pageNumber="62" unit="mm" value="9.05" valueMax="9.5" valueMin="8.6">8.6–9.5 mm</quantity>
|
||
, width
|
||
<quantity id="787F9B4C25005D36FD0CFA27FCC48014" box="[685,802,1522,1544]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.9" metricValueMin="4.3" pageId="7" pageNumber="62" unit="mm" value="4.6" valueMax="4.9" valueMin="4.3">4.3–4.9 mm</quantity>
|
||
(
|
||
<emphasis id="8DF3EABB25005D36FC93FA27FCD88014" box="[818,830,1522,1543]" italics="true" pageId="7" pageNumber="62">n</emphasis>
|
||
=
|
||
<specimenCount id="A981FD2025005D36FCFDFA27FC318014" box="[860,983,1522,1543]" pageId="7" pageNumber="62" type="generic">4 specimens</specimenCount>
|
||
). Color black, each fovea or puncture with 1 minute, decumbent, golden seta and all tubercles clothed in golden, tomentose pubescence unless otherwise noted.
|
||
<emphasis id="8DF3EABB25005D36FBB4F997FBB58044" bold="true" box="[1045,1107,1602,1623]" pageId="7" pageNumber="62">Head:</emphasis>
|
||
Frons densely foveate throughout, without low supraorbital ridge or medial tubercle, posteriorly without transverse impression or nearly confluent foveae; vertex nearly on same plane as frons and densely foveate; frontoclypeal suture deeply and broadly impressed; clypeus moderately setose and with golden pubescence, anterior margin nearly straight, with shallow submarginal impression; gena between eye and clypeus raised and with golden pubescence; impression absent around eye from gena to apex. Eye elongate oval, approximately facets across; large U-shaped lobe present behind eye, lobe moderately punctate and pubescent. Labrum without transverse medial ridge, semi-erect golden setae present on anterior half, margin rounded with setae along edges of vertical surface; mandible bifid at apex, dorsal and lateral surfaces smooth; mentum trapezoidal, rugofoveate with short decumbent setae otherwise, widest at anterior margin, medial longitudinal ridge absent; submentum small, as wide as basal margin of mentum, triangular. Antennomeres 9 and 10 subequal in size, antennomere 11 slightly smaller than preceding segments; antennomere 3 approximately 1.2X length of antennomere 4, antennomeres 4–8 subequal in length.
|
||
<emphasis id="8DF3EABB250F5D39FE25FD00FE1B84F9" bold="true" box="[388,509,725,746]" pageId="8" pageNumber="63">Prothorax:</emphasis>
|
||
Pronotal disc almost flat, widest near middle then constricted in basal third; moderately foveate, without medial impunctate region, tuberculate, each tubercle clothed in golden pubescence and decumbent minute setae curved towards apex of tubercle, apex pubescent and rounded; discal tubercles dense and random, some forming 2 very small submedial patches of nearly confluent tubercles connecting to 2 longitudinal tuberculate ridges running from outer base of horns to near basal margin; anterior half of pronotum bearing 2 sublateral anteriorly directed horns, horns divergent near base then curved towards midline, dorsal and outer surfaces densely tuberculate, ventral surfaces densely foveate, inner surfaces rugose and foveate, apices recurved; pronotum laterally from horn to margin densely tuberculate near base of horn, moderately foveate otherwise, lateral costa present near anterior margin, lateral margin with row of regularly spaced tubercles medially, anterior apices slightly produced and acute, posterior apices acute, not projecting; anterior margin straight, with transverse costa absent, longitudinal costae on underside of horns weakly indicated; posterior margin slightly bisinuate with medial emargination receiving scutellum. Hypomeron moderately foveate, fovea lacking or with a minute setae. Prosternum anterior to coxa approximately as long as coxal cavity, densely foveate; prosternal process raised between coxa, weakly impressed, apex acute.
|
||
<emphasis id="8DF3EABB250F5D39FE28FA0DFDE083FE" bold="true" box="[393,518,1496,1517]" pageId="8" pageNumber="63">Pterothorax:</emphasis>
|
||
Dorsal outline elongate oval, widest behind middle. Elytron dorsally nearly flat to slightly concave, medially depressed posterior to scutellum, sharply sloping and tapering caudad; stria indicated by irregular rows of fovea, interstria with tubercles and somewhat regularly spaced minute decumbent scale-like setae; each elytral disc with 1 short tuberculate submarginal costa from humeral angle to nearly half of elytral length and 1 closer to elytral suture in caudal third, additional short tuberculate costa or protrusions somewhat randomly occurring in interstrial regions; lateral portion of elytron between costae and poorly defined pseudepipleural margin densely tuberculate; pseudepipleura regularly moderately foveate, lacking tubercles; ventral margin of epiplura with deep groove for reception of abdominal ventrites 3–5. Scutellum with minute decumbent scale-like setae, subtriangular, posterior portion V-shaped, dorsal margin rounded and extending past elytral base,
|
||
<emphasis id="8DF3EABB250F5D39FD62FE33FD3787EF" box="[707,721,486,508]" italics="true" pageId="8" pageNumber="63">∼</emphasis>
|
||
1.6X as wide as long. Mesosternum short, anteriorly weakly emarginate behind prosternal process, mesocoxal cavities open. Metasternum short, separating meso- and metacoxal cavities by less than mesocoxal cavity length. All visible ventrites on the pterothorax moderately foveate.
|
||
<emphasis id="8DF3EABB250F5D39FB80FDBEFBBF8493" bold="true" box="[1057,1113,619,640]" pageId="8" pageNumber="63">Legs:</emphasis>
|
||
Femora lacking spines or other protrusions, densely micropubescent with minute decumbent setae; tibia densely clothed in micro-pubescence and minute golden decumbent setae, inner surface with longer straight spine-like setae, outer margin without low pubescent tubercles, apical margin with 2 distinct socketed spurs, apical spine between spurs absent; distal tarsomere length subequal to length of combined preceding segments on meso- and meta-legs, with sparse golden setae, venter of all other tarsomeres clothed with dense long golden setae.
|
||
<emphasis id="8DF3EABB250F5D39FBC6FC5AFD1E85AC" bold="true" pageId="8" pageNumber="63">Abdomen:</emphasis>
|
||
Visible ventrites densely finely punctate medially, each puncture with one minute decumbent seta, relatively clean of pruinescence, lateral margins moderately setose, often obscured by pruinescence; abdominal intercoxal process broad, with submarginal impression, anterior margin rounded; intersegmental membranes concealed, connection between ventrites 3–5 deeply medially depressed; ventrite 5 lacking submarginal groove; abdominal defensive reservoirs present; sternite 8 weakly sclerotized and setose, deeply medially emarginate, emargination V-shaped; aedeagus with parameres fused, much longer than basal piece, dorsally flattened, nearly parallel sided to apical fifth of length, apical fifth sharply acuminate, partially dehisced at tip and weakly curved ventrad (
|
||
<figureCitation id="27BC2A2C250F5D39FC53FAEDFBA8835E" box="[1010,1102,1336,1358]" captionStart="Figs" captionStartId="3.[175,215,577,596]" captionTargetBox="[157,1165,202,555]" captionTargetId="figure-539@3.[151,1173,183,559]" captionTargetPageId="3" captionText="Figs. 3–12. Diceroderes species, aedeagi, dorsal and lateral views. 3–4) D. mexicanus, TB# 14740; 5–6) D. cusucoensis, holotype, TB# 14688; 7–8) D. ocozocoautlaensis, holotype, TB# 14689; 9–10) D. skelleyi, holotype, TB# 13105; 11–12) D. subtriplehorni, holotype, TB# 13109." figureDoi="http://doi.org/10.5281/zenodo.4908451" httpUri="https://zenodo.org/record/4908451/files/figure.png" pageId="8" pageNumber="63">Figs. 5–6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BF3836A9250F5D39FD7AFA86FB8A838E" blockId="8.[707,1194,194,1570]" pageId="8" pageNumber="63">
|
||
<emphasis id="8DF3EABB250F5D39FD7AFA86FCCD837B" bold="true" box="[731,811,1363,1384]" pageId="8" pageNumber="63">Female.</emphasis>
|
||
Similar to male, except with shorter, stouter pronotal horns, horns curved towards midline but not recurved at apex (
|
||
<figureCitation id="27BC2A2C250F5D39FC4BFA5DFB86838E" box="[1002,1120,1416,1438]" captionStart="Figs" captionStartId="7.[709,751,706,725]" captionTargetBox="[685,1171,194,682]" captionTargetId="figure-551@7.[685,1171,194,683]" captionTargetPageId="7" captionText="Figs. 26–27. Diceroderes cusucoensis, allotype" figureDoi="http://doi.org/10.5281/zenodo.4908469" httpUri="https://zenodo.org/record/4908469/files/figure.png" pageId="8" pageNumber="63">Figs. 26–27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F79D6522250F5D39FD7AFA76FC8683C1" pageId="8" pageNumber="63" type="distribution">
|
||
<paragraph id="BF3836A9250F5D39FD7AFA76FC8683C1" blockId="8.[707,1194,194,1570]" pageId="8" pageNumber="63">
|
||
<emphasis id="8DF3EABB250F5D39FD7AFA76FC8483AB" bold="true" box="[731,866,1443,1464]" pageId="8" pageNumber="63">Distribution.</emphasis>
|
||
<collectingCountry id="C7907639250F5D39FCD1FA76FC3383AB" box="[880,981,1443,1464]" name="Honduras" pageId="8" pageNumber="63">Honduras</collectingCountry>
|
||
:
|
||
<collectingRegion id="7D43F84B250F5D39FC42FA76FBCE83AB" box="[995,1064,1443,1464]" country="Honduras" name="Cortes" pageId="8" pageNumber="63">Cortés</collectingRegion>
|
||
;
|
||
<collectingCountry id="C7907639250F5D39FB97FA76FB4083AB" box="[1078,1190,1443,1464]" name="Guatemala" pageId="8" pageNumber="63">Guatemala</collectingCountry>
|
||
:
|
||
<collectingRegion id="7D43F84B250F5D39FD62FA68FD1A83C1" box="[707,764,1469,1490]" country="Guatemala" name="Izabal" pageId="8" pageNumber="63">Izabal</collectingRegion>
|
||
(
|
||
<figureCitation id="27BC2A2C250F5D39FCAAFA68FCB283C1" box="[779,852,1469,1490]" captionStart="Fig" captionStartId="6.[197,229,774,793]" captionTargetBox="[173,1193,194,751]" captionTargetId="figure-447@6.[172,1194,193,751]" captionTargetPageId="6" captionText="Fig. 23. Diceroderes species, distribution map. Triangles = D. mexicanus, white triangles are for morph one, gray/blue triangles for morph two. Stars = D. cusucoensis. Squares = D. ocozocoautlaensis. Diamonds = D. skelleyi. Circles = D. subtriplehorni." figureDoi="http://doi.org/10.5281/zenodo.4908467" httpUri="https://zenodo.org/record/4908467/files/figure.png" pageId="8" pageNumber="63">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F79D6522250F5D39FD7AFA0DFB738031" pageId="8" pageNumber="63" type="etymology">
|
||
<paragraph id="BF3836A9250F5D39FD7AFA0DFB738031" blockId="8.[707,1194,194,1570]" pageId="8" pageNumber="63">
|
||
<emphasis id="8DF3EABB250F5D39FD7AFA0DFCAB83FE" bold="true" box="[731,845,1496,1517]" pageId="8" pageNumber="63">Etymology.</emphasis>
|
||
The species epithet refers to the Cusuco National Park in
|
||
<collectingCountry id="C7907639250F5D39FC13FA27FBE88014" box="[946,1038,1522,1543]" name="Honduras" pageId="8" pageNumber="63">Honduras</collectingCountry>
|
||
where the
|
||
<typeStatus id="603C880B250F5D39FBD6FA27FD0A8031" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
and most of the type series were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |