255 lines
34 KiB
XML
255 lines
34 KiB
XML
<document id="618C836F891BDC1BF8C38E606F4EE102" ID-DOI="10.11646/palaeoentomology.4.5.15" ID-ISSN="2624-2834" ID-Zenodo-Dep="5530240" ID-ZooBank="3747A7EE-21F8-4FC5-9B57-4B726A0AFBD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632740553429" checkinUser="plazi" docAuthor="STRELNIKOVA, OLESYA D. & YAN, EVGENY V." docDate="2021" docId="370687D49131FFE41B1506FEFC87FAF9" docLanguage="en" docName="palaeoentomology.4.5.15..pdf" docOrigin="Palaeoentomology 4 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.5.15" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Notocupes excellens Ponomarenko 1966" docType="treatment" docVersion="3" lastPageNumber="505" masterDocId="CB3FFFAC9132FFE2183F0445FF8DFF92" masterDocTitle="Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia" masterLastPageNumber="514" masterPageNumber="499" pageNumber="502" updateTime="1699415070130" updateUser="plazi">
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<mods:titleInfo id="5EB8D8CE9BA769ADA9B3942663170726">
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<mods:title id="F0369C392A9B66C13709A36C036AE622">Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia</mods:title>
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<mods:namePart id="942CC76E24874992127539954CED18FE">STRELNIKOVA, OLESYA D.</mods:namePart>
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<mods:name id="EED8E384DF99E1CBA492F26F874ED4DB" type="personal">
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<mods:roleTerm id="CD1DEA8FCAAAAE1641A011D261556AE3">Author</mods:roleTerm>
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<mods:namePart id="ACC754B1F3CF0B3B9B27588608798162">YAN, EVGENY V.</mods:namePart>
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<mods:title id="3FCCDFBAB93CE752F334BD5151FDF34E">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="FD4C217A60D6162B6905D10E8844275F">
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<mods:date id="CE83D69920B49BA6ED0720BF6FFCC96C">2021</mods:date>
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<mods:number id="1971059AED378282DDDE9D2D99EC6A8A">2021-09-24</mods:number>
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<mods:number id="ED1307DD88CE029ED62D38214E85E1F6">4</mods:number>
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<mods:number id="FF224D52C55955BA1C763DAF91ECBE6B">5</mods:number>
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<mods:url id="0CE8DFA9439933DAE46CE1F0FD5CAEBD">http://dx.doi.org/10.11646/palaeoentomology.4.5.15</mods:url>
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<mods:classification id="A95CBFDE79A140BA1F6070A7C8CFE62E">journal article</mods:classification>
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<mods:identifier id="185C239F09F6629FBC32E457F1F8942A" type="DOI">10.11646/palaeoentomology.4.5.15</mods:identifier>
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<mods:identifier id="51E0369622D6BB2CED45BD2071F21503" type="ISSN">2624-2834</mods:identifier>
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<treatment id="370687D49131FFE41B1506FEFC87FAF9" LSID="urn:lsid:plazi:treatment:370687D49131FFE41B1506FEFC87FAF9" httpUri="http://treatment.plazi.org/id/370687D49131FFE41B1506FEFC87FAF9" lastPageId="6" lastPageNumber="505" pageId="3" pageNumber="502">
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<subSubSection id="F7B565499131FFE11B1506FEFA85FD46" box="[810,1288,698,725]" pageId="3" pageNumber="502" type="nomenclature">
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<paragraph id="BF1036C29131FFE11B1506FEFA85FD46" blockId="3.[810,1288,698,760]" box="[810,1288,698,725]" pageId="3" pageNumber="502">
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<heading id="E45881AE9131FFE11B1506FEFA85FD46" bold="true" box="[810,1288,698,725]" fontSize="10" level="2" pageId="3" pageNumber="502" reason="3">
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<taxonomicName id="78AF4D419131FFE11B1506FEFA85FD46" authority="Ponomarenko, 1966" authorityName="Ponomarenko" authorityYear="1966" box="[810,1288,698,725]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="502" phylum="Arthropoda" rank="species" species="excellens">
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<emphasis id="8DDBEAD09131FFE11B1506FEFA85FD46" bold="true" box="[810,1288,698,725]" pageId="3" pageNumber="502">
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<emphasis id="8DDBEAD09131FFE11B1506FEFB99FD47" bold="true" box="[810,1044,698,725]" italics="true" pageId="3" pageNumber="502">Notocupes excellens</emphasis>
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Ponomarenko, 1966
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F7B565499131FFE11B15069BFC13FD6A" box="[810,926,734,760]" pageId="3" pageNumber="502" type="description">
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<paragraph id="BF1036C29131FFE11B15069BFC13FD6A" blockId="3.[810,1288,698,760]" box="[810,926,734,760]" pageId="3" pageNumber="502">
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(
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<figureCitation id="27942A479131FFE11B0D069BFC18FD6A" box="[818,917,734,760]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[136,235,1884,1908]" captionStartId-1="4.[136,235,1080,1104]" captionStartId-2="5.[136,235,1354,1378]" captionTargetBox-0="[160,1426,1092,1852]" captionTargetBox-1="[278,1310,197,1049]" captionTargetBox-2="[430,1149,205,1313]" captionTargetId-0="figure-381@3.[151,1436,1085,1861]" captionTargetId-1="figure-395@4.[269,1318,189,1057]" captionTargetId-2="figure-251@5.[418,1170,189,1330]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Notocupes excellens (PIN, 1989/3007). A, Wetted with 95% ethanol. B, Dry. C, Interpretative line drawing (here and on the further figures square “a” indicate area with small tubercles and square “b” indicate area with larger tubercles).Abbreviations here and in the following figures: dep, depressions; epl, epipleura; gen, genae; hypo, hypomeron; lab, labrum; me+sme+gu, fused mentum, submentum and gular plate; mnd, mandibles; pl, pleura; pra, praementum; prst, prosternum; prtr, protrochantin; s.gu, sutura gularis; s.n.pl., sutura notopleuralis; s.pl.st., sutura pleurasternalis." captionText-1="FIGURE 3. Counterpart of Notocupes excellens (PIN, 1989/3007). A, Counterpart wetted with 95% ethanol. B, Interpretative line drawing." captionText-2="FIGURE 4. Line drawings of elytra of Notocupes excellens. A, Mirrored line drawing of left elytra of part. B, Line drawing of right elytra of part. C, Line drawing of left elytra of elongate-oval cells, counterpart. D, Mirrored line drawing of right elytra of counterpart. E, General scheme of Notocupes excellens elytra. Labels here and in the following: Roman numerals—fields; Arabic numerals—veins." figureDoi-0="http://doi.org/10.5281/zenodo.5530244" figureDoi-1="http://doi.org/10.5281/zenodo.5530246" figureDoi-2="http://doi.org/10.5281/zenodo.5530248" httpUri-0="https://zenodo.org/record/5530244/files/figure.png" httpUri-1="https://zenodo.org/record/5530246/files/figure.png" httpUri-2="https://zenodo.org/record/5530248/files/figure.png" pageId="3" pageNumber="502">Figs 2–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F7B565499131FFE11B150763FACCFCF6" pageId="3" pageNumber="502" type="materials_examined">
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<paragraph id="BF1036C29131FFE11B150763FACCFCF6" blockId="3.[810,1452,806,1048]" pageId="3" pageNumber="502">
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<emphasis id="8DDBEAD09131FFE11B150763FC14FCD2" bold="true" box="[810,921,806,832]" pageId="3" pageNumber="502">Material.</emphasis>
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<materialsCitation id="0FC73C9F9131FFE11BA00763FACCFCF6" collectionCode="PIN" location="Baissa" municipality="Lower Cretaceous." pageId="3" pageNumber="502" specimenCount="1" typeStatus="holotype">
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<typeStatus id="601488609131FFE11BA00763FB8AFCD2" box="[927,1031,806,832]" pageId="3" pageNumber="502" type="holotype">Holotype</typeStatus>
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<collectionCode id="D9BEAE079131FFE11C320762FBB0FCD3" box="[1037,1085,807,833]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="3" pageNumber="502">PIN</collectionCode>
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, 1989/3007, part and counterpart of full body,
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<location id="BA7060199131FFE11B85070FFB8EFCF6" LSID="urn:lsid:plazi:treatment:370687D49131FFE41B1506FEFC87FAF9:BA7060199131FFE11B85070FFB8EFCF6" box="[954,1027,842,868]" municipality="Lower Cretaceous." name="Baissa" pageId="3" pageNumber="502">Baissa</location>
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locality;
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<collectingMunicipality id="5F74ACB89131FFE11C52070EFACCFCF6" box="[1133,1345,842,869]" pageId="3" pageNumber="502">Lower Cretaceous.</collectingMunicipality>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F7B565499131FFE61B60072BFD42FAA4" lastPageId="4" lastPageNumber="503" pageId="3" pageNumber="502" type="diagnosis">
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<paragraph id="BF1036C29131FFE61B60072BFD42FAA4" blockId="3.[810,1452,806,1048]" lastBlockId="4.[136,778,1199,2093]" lastPageId="4" lastPageNumber="503" pageId="3" pageNumber="502">
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<emphasis id="8DDBEAD09131FFE11B60072BFC54FC1A" bold="true" box="[863,985,878,904]" pageId="3" pageNumber="502">Diagnosis.</emphasis>
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Genae protruding anteriorly as pair of acute processes, lateral to mandibles. Pronotum with dentate lateral margins.Antennae reaching slightly beyond pronotal base, weakly moniliform. Elytra with rectangular cells with somewhat rounded angles. Maximum of cells in a row 26. Black maculae absent. Cells occupy more than half distance between veins, often reaching lateral margin of fields, forming weak zigzag pattern. Epipleural margin dentate near elytral base, epipleuron narrow.
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</paragraph>
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</subSubSection>
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<caption id="EBD0664A9131FFE118B70319FCE1F797" ID-DOI="http://doi.org/10.5281/zenodo.5530244" ID-Zenodo-Dep="5530244" httpUri="https://zenodo.org/record/5530244/files/figure.png" pageId="3" pageNumber="502" startId="3.[136,235,1884,1908]" targetBox="[160,1426,1092,1852]" targetPageId="3">
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<paragraph id="BF1036C29131FFE118B70319FCE1F797" blockId="3.[136,1452,1884,2053]" pageId="3" pageNumber="502">
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<emphasis id="8DDBEAD09131FFE118B70319FE89F8E6" bold="true" box="[136,260,1884,1908]" pageId="3" pageNumber="502">FIGURE 2.</emphasis>
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<taxonomicName id="78AF4D419131FFE119350318FE55F8E7" authorityName="Ponomarenko" authorityYear="1966" box="[266,472,1885,1909]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="502" phylum="Arthropoda" rank="species" species="excellens">
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<emphasis id="8DDBEAD09131FFE119350318FE55F8E7" box="[266,472,1885,1909]" italics="true" pageId="3" pageNumber="502">Notocupes excellens</emphasis>
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</taxonomicName>
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(PIN, 1989/3007
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<emphasis id="8DDBEAD09131FFE11AB60318FD1CF8E6" box="[649,657,1885,1908]" italics="true" pageId="3" pageNumber="502">)</emphasis>
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.
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<emphasis id="8DDBEAD09131FFE11AA10319FD3DF8E6" bold="true" box="[670,688,1884,1908]" pageId="3" pageNumber="502">A</emphasis>
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, Wetted with 95% ethanol.
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<emphasis id="8DDBEAD09131FFE11BF60318FC57F8E7" bold="true" box="[969,986,1885,1909]" pageId="3" pageNumber="502">B</emphasis>
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, Dry.
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<emphasis id="8DDBEAD09131FFE11C260319FBA6F8E6" bold="true" box="[1049,1067,1884,1908]" pageId="3" pageNumber="502">C</emphasis>
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, Interpretative line drawing (here and on the further figures square “a” indicate area with small tubercles and square “b” indicate area with larger tubercles). Abbreviations here and in the following figures: dep, depressions; epl, epipleura; gen, genae; hypo, hypomeron; lab, labrum; me+sme+gu, fused mentum, submentum and gular plate; mnd, mandibles; pl, pleura; pra, praementum; prst, prosternum; prtr, protrochantin; s.gu, sutura gularis; s.n.pl., sutura notopleuralis; s.pl.st., sutura pleurasternalis.
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</paragraph>
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</caption>
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<caption id="EBD0664A9136FFE618B7007DFF6FFBE6" ID-DOI="http://doi.org/10.5281/zenodo.5530246" ID-Zenodo-Dep="5530246" httpUri="https://zenodo.org/record/5530246/files/figure.png" pageId="4" pageNumber="503" startId="4.[136,235,1080,1104]" targetBox="[278,1310,197,1049]" targetPageId="4">
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<paragraph id="BF1036C29136FFE618B7007DFF6FFBE6" blockId="4.[136,1451,1080,1140]" pageId="4" pageNumber="503">
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<emphasis id="8DDBEAD09136FFE618B7007DFE8EFBC2" bold="true" box="[136,259,1080,1104]" pageId="4" pageNumber="503">FIGURE 3.</emphasis>
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Counterpart of
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<taxonomicName id="78AF4D419136FFE6199C007CFDFCFBC2" authorityName="Ponomarenko" authorityYear="1966" box="[419,625,1080,1104]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="503" phylum="Arthropoda" rank="species" species="excellens">
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<emphasis id="8DDBEAD09136FFE6199C007CFDFCFBC2" box="[419,625,1080,1104]" italics="true" pageId="4" pageNumber="503">Notocupes excellens</emphasis>
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</taxonomicName>
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(PIN, 1989/3007
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<emphasis id="8DDBEAD09136FFE61B1E007DFCA4FBDD" box="[801,809,1080,1103]" italics="true" pageId="4" pageNumber="503">)</emphasis>
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.
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<emphasis id="8DDBEAD09136FFE61B0A007DFCCAFBC2" bold="true" box="[821,839,1080,1104]" pageId="4" pageNumber="503">A</emphasis>
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, Counterpart wetted with 95% ethanol.
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<emphasis id="8DDBEAD09136FFE61CE5007DFB66FBC2" bold="true" box="[1242,1259,1080,1104]" pageId="4" pageNumber="503">B</emphasis>
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, Interpretative line drawing.
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</paragraph>
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||
</caption>
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||
<subSubSection id="F7B565499136FFE41882017AFD2BFCD3" lastPageId="6" lastPageNumber="505" pageId="4" pageNumber="503" type="description">
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<paragraph id="BF1036C29136FFE61882017AFC87F97A" blockId="4.[136,778,1199,2093]" pageId="4" pageNumber="503">
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<emphasis id="8DDBEAD09136FFE61882017AFEC2FACB" bold="true" box="[189,335,1343,1369]" pageId="4" pageNumber="503">Description.</emphasis>
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Body length/width ratio 2.8. Cuticular tubercles roundish, uniformly covering entire body, on most regions around
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<quantity id="78579B279136FFE6198001CDFDA0FA30" box="[447,557,1416,1442]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="503" unit="mm" value="0.02">0.02 mm</quantity>
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in diameter with density of 720 tubercles per mm² (
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<figureCitation id="27942A479136FFE61A0901E9FDF2FA55" box="[566,639,1452,1479]" captionStart="FIGURE 2" captionStartId="3.[136,235,1884,1908]" captionTargetBox="[160,1426,1092,1852]" captionTargetId="figure-381@3.[151,1436,1085,1861]" captionTargetPageId="3" captionText="FIGURE 2. Notocupes excellens (PIN, 1989/3007). A, Wetted with 95% ethanol. B, Dry. C, Interpretative line drawing (here and on the further figures square “a” indicate area with small tubercles and square “b” indicate area with larger tubercles).Abbreviations here and in the following figures: dep, depressions; epl, epipleura; gen, genae; hypo, hypomeron; lab, labrum; me+sme+gu, fused mentum, submentum and gular plate; mnd, mandibles; pl, pleura; pra, praementum; prst, prosternum; prtr, protrochantin; s.gu, sutura gularis; s.n.pl., sutura notopleuralis; s.pl.st., sutura pleurasternalis." figureDoi="http://doi.org/10.5281/zenodo.5530244" httpUri="https://zenodo.org/record/5530244/files/figure.png" pageId="4" pageNumber="503">Fig. 2</figureCitation>
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C-b); larger on meso- and metaventrite, apical abdominal sternite and on depressed areas of abdominal sternites III–VI, with
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<quantity id="78579B279136FFE618F7025DFEBEF9A1" box="[200,307,1560,1587]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="503" unit="mm" value="0.06">0.06 mm</quantity>
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||
diameter and density of 228 per mm² (
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||
<figureCitation id="27942A479136FFE618AF0279FF54F9C5" box="[144,217,1596,1623]" captionStart="FIGURE 2" captionStartId="3.[136,235,1884,1908]" captionTargetBox="[160,1426,1092,1852]" captionTargetId="figure-381@3.[151,1436,1085,1861]" captionTargetPageId="3" captionText="FIGURE 2. Notocupes excellens (PIN, 1989/3007). A, Wetted with 95% ethanol. B, Dry. C, Interpretative line drawing (here and on the further figures square “a” indicate area with small tubercles and square “b” indicate area with larger tubercles).Abbreviations here and in the following figures: dep, depressions; epl, epipleura; gen, genae; hypo, hypomeron; lab, labrum; me+sme+gu, fused mentum, submentum and gular plate; mnd, mandibles; pl, pleura; pra, praementum; prst, prosternum; prtr, protrochantin; s.gu, sutura gularis; s.n.pl., sutura notopleuralis; s.pl.st., sutura pleurasternalis." figureDoi="http://doi.org/10.5281/zenodo.5530244" httpUri="https://zenodo.org/record/5530244/files/figure.png" pageId="4" pageNumber="503">Fig. 2</figureCitation>
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C-a). Distance between tubercles on areas with smaller ones
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<quantity id="78579B279136FFE6190F0224FE53F9E9" box="[304,478,1633,1659]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="503" unit="mm" value="0.015" valueMax="0.02" valueMin="0.01">0.01–0.02 mm</quantity>
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, approximately equal to single tubercle diameter; on areas with large tubercles distance at most
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<quantity id="78579B279136FFE6194C02ECFE67F951" box="[371,490,1705,1731]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="503" unit="mm" value="0.01">0.01 mm</quantity>
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, notably smaller than diameter of tubercles. Elytra without color pattern.
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</paragraph>
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<paragraph id="BF1036C29136FFE6188202B4FBF6F9A1" blockId="4.[136,778,1199,2093]" lastBlockId="4.[810,1452,1199,2092]" pageId="4" pageNumber="503">
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Head: transverse, length
|
||
<quantity id="78579B279136FFE619E302B4FDBAF89E" box="[476,567,1777,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="503" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, head length/body length ratio 5.3. Maximum width
|
||
<quantity id="78579B279136FFE61A160350FD07F8A2" box="[553,650,1813,1840]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="503" unit="mm" value="1.2">1.2 mm</quantity>
|
||
. Head not narrowed in front of eyes, gradually narrowing behind eyes, with neck constriction weak. Head length in front of eyes less than behind them, slightly longer than diameter of eyes. Protuberances on head not visible. Eyes large, protruding laterally, approximately
|
||
<quantity id="78579B279136FFE61A20038FFDF4F877" box="[543,633,1994,2021]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="503" unit="mm" value="0.4">0.4 mm</quantity>
|
||
in diameter. Head width/ocular diameter ratio 3.1. Labrum transverse, medially incised, anterior angles not rounded.Praementum small, semioval; mentum, submentum and gular plate fused. Mandibles with at least 2 preapical teeth; weakly curved. Antennae reaching slightly beyond pronotal base,
|
||
<quantity id="78579B279136FFE61B4F015EFC41FAA4" box="[880,972,1307,1334]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="503" unit="mm" value="2.7">2.7 mm</quantity>
|
||
long, weakly moniliform, attached near anterior ocular margins. Antennomeres slightly widening distally; 1–6 elongate-oval, subsequent ones narrower, subtrapezoidal; scapus length
|
||
<quantity id="78579B279136FFE61CB301CDFB67FA30" box="[1164,1258,1416,1442]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="503" unit="mm" value="0.4">0.4 mm</quantity>
|
||
, width
|
||
<quantity id="78579B279136FFE61D7701CDFA2AFA30" box="[1352,1447,1416,1442]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="503" unit="mm" value="0.3">0.3 mm</quantity>
|
||
; pedicellus equal to antennomere 3, 0.7 times as long as scapus, almost 0.8 times as long as 4, and 1.4 times as long as 5; remaining antennomeres gradually decreasing in size towards antennal apex.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29136FFE61B600278FA21F8BD" blockId="4.[810,1452,1199,2092]" pageId="4" pageNumber="503">
|
||
Pronotum:
|
||
<quantity id="78579B279136FFE61BD00278FBD5F9C5" box="[1007,1112,1597,1623]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="503" unit="mm" value="1.8">1.8 mm</quantity>
|
||
long, maximum width—
|
||
<quantity id="78579B279136FFE61B150224FC19F9E9" box="[810,916,1633,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="503" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; posterior margin 1.5 times wider than long. Pronotal maximum width/head width ratio 2.2. Anterior angles strongly protruding forward. Pronotal posterior margin straight, without posterior angles protruding posterolaterally; lateral margin dentate; disk with two median protuberances.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29136FFE61B60037FFC21F7BE" blockId="4.[810,1452,1199,2092]" pageId="4" pageNumber="503">
|
||
Prosternum:
|
||
<quantity id="78579B279136FFE61BCA037FFBDCF8C6" box="[1013,1105,1850,1876]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="503" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long. Body length/prosternal length ratio 7.9. Prosternum longer than wide. Prosternal length/head length ratio 0.7. Notopleural and pleurosternal sutures clearly visible, former thickened, both angled towards midline of body at 1/3 of anterior length, then straight. Prosternal process indistinct, procoxae contiguous.
|
||
</paragraph>
|
||
<caption id="EBD0664A9137FFE718B7010FFEB6FA5C" ID-DOI="http://doi.org/10.5281/zenodo.5530248" ID-Zenodo-Dep="5530248" httpUri="https://zenodo.org/record/5530248/files/figure.png" pageId="5" pageNumber="504" startId="5.[136,235,1354,1378]" targetBox="[430,1149,205,1313]" targetPageId="5">
|
||
<paragraph id="BF1036C29137FFE718B7010FFEB6FA5C" blockId="5.[136,1451,1354,1486]" pageId="5" pageNumber="504">
|
||
<emphasis id="8DDBEAD09137FFE718B7010FFE88FAF0" bold="true" box="[136,261,1354,1378]" pageId="5" pageNumber="504">FIGURE 4.</emphasis>
|
||
Line drawings of elytra of
|
||
<taxonomicName id="78AF4D419137FFE71A1C010EFD7EFAF0" authorityName="Ponomarenko" authorityYear="1966" box="[547,755,1354,1378]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="504" phylum="Arthropoda" rank="species" species="excellens">
|
||
<emphasis id="8DDBEAD09137FFE71A1C010EFD7EFAF0" box="[547,755,1354,1378]" italics="true" pageId="5" pageNumber="504">Notocupes excellens</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="8DDBEAD09137FFE71B3F010FFC9FFAF0" bold="true" box="[768,786,1354,1378]" pageId="5" pageNumber="504">A</emphasis>
|
||
, Mirrored line drawing of left elytra of part.
|
||
<emphasis id="8DDBEAD09137FFE71CD9010FFB7AFAF0" bold="true" box="[1254,1271,1354,1378]" pageId="5" pageNumber="504">B</emphasis>
|
||
, Line drawing of right elytra of part.
|
||
<emphasis id="8DDBEAD09137FFE7196C012BFEE8FA14" bold="true" box="[339,357,1390,1414]" pageId="5" pageNumber="504">C</emphasis>
|
||
, Line drawing of left elytra of elongate-oval cells, counterpart.
|
||
<emphasis id="8DDBEAD09137FFE71BC9012BFB85FA14" bold="true" box="[1014,1032,1390,1414]" pageId="5" pageNumber="504">D</emphasis>
|
||
, Mirrored line drawing of right elytra of counterpart.
|
||
<emphasis id="8DDBEAD09137FFE7193701D7FE94FA38" bold="true" box="[264,281,1426,1450]" pageId="5" pageNumber="504">E</emphasis>
|
||
, General scheme of
|
||
<taxonomicName id="78AF4D419137FFE719D501D6FD34FA38" authorityName="Ponomarenko" authorityYear="1966" box="[490,697,1426,1450]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="504" phylum="Arthropoda" rank="species" species="excellens">
|
||
<emphasis id="8DDBEAD09137FFE719D501D6FD34FA38" box="[490,697,1426,1450]" italics="true" pageId="5" pageNumber="504">Notocupes excellens</emphasis>
|
||
</taxonomicName>
|
||
elytra. Labels here and in the following: Roman numerals—fields; Arabic numerals—veins.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BF1036C29137FFE71882024FFE71F9B6" blockId="5.[136,778,1546,2093]" box="[189,508,1546,1572]" pageId="5" pageNumber="504">Scutellar shield: pentagonal.</paragraph>
|
||
<paragraph id="BF1036C29137FFE718820275FCD9F972" blockId="5.[136,778,1546,2093]" lastBlockId="5.[810,1452,1548,2092]" pageId="5" pageNumber="504">Elytra: Rectangular cells with somewhat rounded angles. Field I lacks cells, other fields with double rows of cells; fields II and IV contain 26 cells, field III 21 cells; field V has 20–21 cells and field VI about 25; field VII lacks cells. Number of cell rows 10. Cells occupy more than half distance between veins, often reaching lateral margins of fields, forming weak zigzag patterns. Elytral length/width ratio 3.1. Eight veins in total, veins 2 and 3 with common stem, starting 2 cells anterior to basal elytral margin; vein 2 curved, bordering epipleural margin; vein 3 gently s-shaped; veins 2 and 3 with four cells merged, veins 4 with five cells merged anterior to elytral apex; veins 2+3 and 4+5 meet margin of elytra. Maximum width of elytra at midlength. Epipleural margin dentate near elytral base; epipleuron narrow, width near elytral base/width near apex ratio 4. Elytral base straight, humeral bulge distinct and convex; elytral apex acute. Elytra extending beyond abdominal apex posteriorly. Elytral maximum width/pronotal maximum width ratio 1.5.</paragraph>
|
||
<paragraph id="BF1036C29137FFE71B6002AFFC15F7BE" blockId="5.[810,1452,1548,2092]" pageId="5" pageNumber="504">
|
||
Metaventrite: long, width at anterior margin
|
||
<quantity id="78579B279137FFE71B150355FC0CF8B8" box="[810,897,1808,1834]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="504" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, at posterior margin
|
||
<quantity id="78579B279137FFE71C5A0355FB24F8B8" box="[1125,1193,1808,1834]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="504" unit="mm" value="2.0">2 mm</quantity>
|
||
. Pronotal length/length of metaventrite ratio 1.8; posterior width of metaventrite/ anterior width ratio 2.5; width at posterior margin/length ratio 2; metaventrite slightly shorter than mesoventrite. Lateral region with oblique depressions, covered with large tubercles, similar to those on abdominal sternites. Median portion of metaventrite with elongate-triangular elevation.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29134FFE4188204DBFDAFFE02" blockId="6.[136,778,158,1387]" pageId="6" pageNumber="505">
|
||
Abdomen: pattern of sternites forming distinct relief. Sides of apical sternite VII and sternites III–VI (ventrites 1–4) with paired roundish or triangular depressions (
|
||
<figureCitation id="27942A479134FFE41AE304A3FF1BFEB7" captionStart="FIGURE 2" captionStartId="3.[136,235,1884,1908]" captionTargetBox="[160,1426,1092,1852]" captionTargetId="figure-381@3.[151,1436,1085,1861]" captionTargetPageId="3" captionText="FIGURE 2. Notocupes excellens (PIN, 1989/3007). A, Wetted with 95% ethanol. B, Dry. C, Interpretative line drawing (here and on the further figures square “a” indicate area with small tubercles and square “b” indicate area with larger tubercles).Abbreviations here and in the following figures: dep, depressions; epl, epipleura; gen, genae; hypo, hypomeron; lab, labrum; me+sme+gu, fused mentum, submentum and gular plate; mnd, mandibles; pl, pleura; pra, praementum; prst, prosternum; prtr, protrochantin; s.gu, sutura gularis; s.n.pl., sutura notopleuralis; s.pl.st., sutura pleurasternalis." figureDoi="http://doi.org/10.5281/zenodo.5530244" httpUri="https://zenodo.org/record/5530244/files/figure.png" pageId="6" pageNumber="505">Fig. 2</figureCitation>
|
||
—dep). Abdomen slightly narrowing towards pointed apex, starting from the fourth ventrite. Length ratio of visible ventrites (sternites III–VII) 2:1:1:1:2.6. Length/ width ratio of apical sternite VII 1.6.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29134FFE4188205DEFC87FD6A" blockId="6.[136,778,158,1387]" pageId="6" pageNumber="505">
|
||
Legs: procoxae round, contiguous, located at posterior prosternal margin,
|
||
<quantity id="78579B279134FFE419D705FBFDC4FE4B" box="[488,585,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="505" unit="mm" value="0.4">0.4 mm</quantity>
|
||
wide. Pronotal width/procoxal width ratio 6.7. Profemora thickest at midlength,
|
||
<quantity id="78579B279134FFE419310643FEE6FDB3" box="[270,363,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="505" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long; profemoral length/maximum width of pronotum ratio 1.8. Tibiae parallel-sided, with longitudinal keel and one or two rows of transverse ridges; length
|
||
<quantity id="78579B279134FFE419050637FE41FD1F" box="[314,460,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="6" pageNumber="505" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
|
||
. Tarsomeres sizes from 1 to 5 are: 0.2:0.2:0.1:0.1:0.2. Middle coxae rounded. Metacoxae cover more than half of first abdominal ventrite (sternite III), width 1.3 times exceeding length.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29134FFE418820747FD2BFCD3" blockId="6.[136,778,158,1387]" pageId="6" pageNumber="505">
|
||
<emphasis id="8DDBEAD09134FFE418820747FEF9FC8E" bold="true" box="[189,372,770,796]" pageId="6" pageNumber="505">Measurements.</emphasis>
|
||
Length—
|
||
<quantity id="78579B279134FFE419D60747FDCFFC8F" box="[489,578,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="505" unit="mm" value="9.5">9.5 mm</quantity>
|
||
, width—
|
||
<quantity id="78579B279134FFE41A920747FC88FC8F" box="[685,773,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="505" unit="mm" value="3.3">3.3 mm</quantity>
|
||
, elytral length—
|
||
<quantity id="78579B279134FFE419040763FE1EFCD3" box="[315,403,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="505" unit="mm" value="6.5">6.5 mm</quantity>
|
||
, elytral width—
|
||
<quantity id="78579B279134FFE41A730763FD2FFCD3" box="[588,674,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="505" unit="mm" value="2.1">2.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F7B565499134FFE41882070CFC87FAF9" pageId="6" pageNumber="505" type="discussion">
|
||
<paragraph id="BF1036C29134FFE41882070CFF52FB09" blockId="6.[136,778,158,1387]" pageId="6" pageNumber="505">
|
||
<emphasis id="8DDBEAD09134FFE41882070CFEBCFCF1" bold="true" box="[189,305,841,867]" pageId="6" pageNumber="505">Remarks.</emphasis>
|
||
In the original descriptions it was stated that the cephalic area between the eyes has longitudinal keels, that pointed protuberances are present above the eyes, and that deep antennal furrows are present on the ventral surface of the head. Our investigation reveals that these observations were likely incorrect: the head is mostly visible in ventral view, which makes it impossible to verify the characters mentioned above. Structures addressed as “antennal furrows” are most likely gular sutures.
|
||
</paragraph>
|
||
<paragraph id="BF1036C29134FFE4188200E6FC87FAF9" blockId="6.[136,778,158,1387]" pageId="6" pageNumber="505">
|
||
The species under consideration is similar to Triassic
|
||
<taxonomicName id="78AF4D419134FFE418D70082FEEAFB72" authorityName="Ponomarenko" authorityYear="1969" box="[232,359,1222,1249]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="505" phylum="Arthropoda" rank="species" species="laticella">
|
||
<emphasis id="8DDBEAD09134FFE418D70082FF76FB72" box="[232,251,1223,1248]" italics="true" pageId="6" pageNumber="505">N</emphasis>
|
||
.
|
||
<emphasis id="8DDBEAD09134FFE419340083FEEAFB72" box="[267,359,1222,1248]" italics="true" pageId="6" pageNumber="505">laticella</emphasis>
|
||
</taxonomicName>
|
||
in the length of the antennae, which also reach slightly beyond the base of the pronotum; with
|
||
<taxonomicName id="78AF4D419134FFE418F00149FD04FAB4" authority="Strelnikova, 2019" authorityName="Strelnikova" authorityYear="2019" box="[207,649,1291,1318]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="505" phylum="Arthropoda" rank="species" species="khasurtyiensis">
|
||
<emphasis id="8DDBEAD09134FFE418F00149FF6FFAB7" box="[207,226,1292,1317]" italics="true" pageId="6" pageNumber="505">N</emphasis>
|
||
.
|
||
<emphasis id="8DDBEAD09134FFE418C1014EFE13FAB7" box="[254,414,1291,1317]" italics="true" pageId="6" pageNumber="505">khasurtyiensis</emphasis>
|
||
<bibRefCitation id="DB3E4B339134FFE4198B014EFD04FAB4" author="Strelnikova, O. D." box="[436,649,1291,1318]" pageId="6" pageNumber="505" pagination="292 - 299" refId="ref11443" refString="Strelnikova, O. D. (2019) New cupedids (Insecta: Coleoptera, Cupedidae) from the Lower Cretaceous of Buryatia. Paleontological Journal, 53 (3), 292 - 299. https: // doi. org / 10.1134 / S 0031030119030146" type="journal article" year="2019">Strelnikova, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
it shares moniliform antenna, with
|
||
<taxonomicName id="78AF4D419134FFE419F7016AFF65FAF9" authority="Ponomarenko, 1966 a" authorityName="Ponomarenko" authorityYear="1966" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="505" phylum="Arthropoda" rank="species" species="caudatus">
|
||
<emphasis id="8DDBEAD09134FFE419F7016AFE56FADA" box="[456,475,1327,1352]" italics="true" pageId="6" pageNumber="505">N</emphasis>
|
||
.
|
||
<emphasis id="8DDBEAD09134FFE419CD016BFDDAFADA" box="[498,599,1326,1352]" italics="true" pageId="6" pageNumber="505">caudatus</emphasis>
|
||
<bibRefCitation id="DB3E4B339134FFE41A58016BFF65FAF9" author="Ponomarenko, A. G." pageId="6" pageNumber="505" pagination="138 - 143" refId="ref10350" refString="Ponomarenko, A. G. (1966 a) New beetles of the family Cupedidae (Coleoptera) from Mesozoic deposits of Transbaikalia. Revue d'Entomologie de l'URSS, 45, 1, 138 - 143 [In Russian]." type="journal article" year="1966">Ponomarenko, 1966 a</bibRefCitation>
|
||
</taxonomicName>
|
||
dentate lateral edge of the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |