treatments-xml/data/36/6E/3B/366E3BE326515983827B262999391B0E.xml

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<mods:title id="DDF3DD2F9434D149C6A5233C3050DA4B">Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae)</mods:title>
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<mods:namePart id="24EBDA8053622F8D762B17C93DA757CB">Patitucci, Luciano Damian</mods:namePart>
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<mods:affiliation id="ACB35A1EB24F30E19DF2F560B73B8698">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="01E135FC8ADBD1E1B2391BE6393B5A27">Mulieri, Pablo Ricardo</mods:namePart>
<mods:affiliation id="B0B1F3C2002A0D2D16D489C539016EB4">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="BADBDBE450D2B65F40F4870C20A8C055">Couri, Marcia Souto</mods:namePart>
<mods:affiliation id="79F7A4BB80F8369D1EEF000B291382CF">Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brazil &amp; Conselho Nacional de Desenvolvimento Cientifico e Tecnologico (CNPq), Brazil</mods:affiliation>
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<mods:namePart id="370FE787914F25C8EF1FA2600AB452EF">Dominguez, Martha Cecilia</mods:namePart>
<mods:affiliation id="9F72055DAED7F4D84A412A67D29EC217">Laboratorio de Entomologia, Instituto Argentino de Investigaciones de Zonas Aridas (IADIZA-CONICET) Mendoza, Mendoza, Argentina</mods:affiliation>
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<mods:date id="2847494C6EE76214F12938522240EA44">2023</mods:date>
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<paragraph id="2D6CBA22444B7FC3C651A74119F66281" pageId="0" pageNumber="611">
<taxonomicName id="3269969BC1BFDB1853BD6B4A86DBE7B9" ID-CoL="WV6V" LSID="366E3BE3-2651-5983-827B-262999391B0E" authority="(Malloch, 1934) comb. rest." authorityName="comb. rest." baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">Coenosia nigerrima (Malloch, 1934) comb. rest.</taxonomicName>
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<subSubSection id="69D84FDB0FD65085DE8229878C2A65B3" pageId="0" pageNumber="611" type="male (fig. 21a)">
<paragraph id="009690EC9BEDAA2C02D3580DF6D4926F" pageId="0" pageNumber="611">Male (Fig. 21A).</paragraph>
<paragraph id="DDEA8832B5E7D4B09A6B05E1E2004141" pageId="0" pageNumber="611">
Length. Body: 3.40-3.94 mm, wing: 2.80-3.49 mm. 
<emphasis id="8159934DA1A7E983ACB126E822B2C2B3" bold="true" italics="true" pageId="0" pageNumber="611">Head</emphasis>
: (Fig. 
<figureCitation id="7CA9D676FD57082E563747CA22B6D5D0" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21B</figureCitation>
). Frons, fronto-orbital plate, parafacial and gena black. 5-6 pairs of frontal setae. Frons at vertex about one third of the head width. Frons with similar length and width, with frontal triangle short, black, not reaching lunule. Gena with a similar width to the postpedicel. Fronto-orbital plate with 6-8 short setulae, close to parafacialia. Parafacial in lateral view narrow. Antenna black; in lateral view inserted over the mid-level of the eye; arista with its longest microtrichia shorter than its basal diameter. Palpus black. 
<emphasis id="22FD70ED95C2B6DF6EA74B411E7BEE2A" bold="true" italics="true" pageId="0" pageNumber="611">Thorax</emphasis>
: (Fig. 
<figureCitation id="0F5662B582771F0B443C31A961BD49EB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21C</figureCitation>
). Black with brown pollinosity, without vitta; anterior and posterior spiracles black. Chaetotaxy: acr s strong and biseriate; anterior presutural pair longer than the extra pair of presutural dorsocentral (eaDC); dorsocentrals 1+3, eaDC is less than one fifth of the aDC. Katepisternum with 2-3 setulae. 
<emphasis id="0AFA9A281FD5942FBD580B44AB3F3A66" bold="true" italics="true" pageId="0" pageNumber="611">Wing</emphasis>
: Tegula black. Both calypters hyaline with black margins; halter black. 
<emphasis id="19D19F550C49EB12BC7DC7150CFEACE2" bold="true" italics="true" pageId="0" pageNumber="611">Legs</emphasis>
: Coxae, trochanters, femora, tibiae, and tarsus black. Fore femur with a row of strong pd to pv setae, and a row of strong av setae on basal third. Mid femur with 1-2 long setae on ventral surface, a row of setae on anterior surface, and 2 preapical setae on pd to posterior surface; mid tibia with one ad and one pd median setae, with the same length and ad seta positioned below pd seta. Hind femur with a row of ad and a row of av setae, with one preapical ad seta and one pd seta. Hind tibia similar size of claws and pulvilli of the three legs. 
<emphasis id="FBA596C1D9ABDBCD059BED82010B07E4" bold="true" italics="true" pageId="0" pageNumber="611">Abdomen</emphasis>
: (Fig. 
<figureCitation id="4CB9DC20EBA1623127BB82A8CFC7DB5E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21D</figureCitation>
). Black without spots. Tergite 6 not visible in dorsal or lateral view. Sternite 5 broader than long, apical margin concave with a thin 
<normalizedToken id="CBDC50962E72EE2FE982CB35303E75FA" originalValue="“U”">&quot;U&quot;</normalizedToken>
shape reduced towards the base, and without membrane; few setae concentrated on the lobes and some long and strong on apical margin (Fig. 
<figureCitation id="C967486DC829222740D91F04EF15BC15" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21E</figureCitation>
). 
<emphasis id="67C8B790E8879C506DC0E1522EDEA492" bold="true" italics="true" pageId="0" pageNumber="611">Terminalia</emphasis>
: Cercus curved in lateral view, and two longitudinal keels only on basal half, apical margin straight. Little spines on inner surface. Surstylus broad at base and at apex, curved, with a pointed process at apical third in inner surface (Fig. 
<figureCitation id="C40B455D8E5F9C8C64068E96779AD626" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21F-H</figureCitation>
). Hypandrium tubular longer than wide, with several little spicules close to pregt. Aedeagus with pgt developed and poor sclerotized; epiphalus slightly sclerotized, and distiph tubular, strongly sclerotized at base (Fig. 
<figureCitation id="BDEA42CD96E87D7988A271284C618FDA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Coenosia nigerrima, male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, dorsal view (SEM), G cercus and surstylus, dorsal view, H cercus, surstylus and phallic complex, lateral view, I phallic complex, lateral view, J phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows." figureDoi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611">21I, J</figureCitation>
).
</paragraph>
<caption id="AFEC861ABCDDEBE2B82F3281C595A62C" doi="10.3897/asp.81.e104969.figure21" httpUri="https://binary.pensoft.net/fig/882089" pageId="0" pageNumber="611" start="Figure 21" startId="F21">
<paragraph id="22971C8ED928220B4E9F747D51E948D4" pageId="0" pageNumber="611">
<emphasis id="0805ED2EBFBA9891DD174CA7691085CB" bold="true" pageId="0" pageNumber="611">Figure 21.</emphasis>
<taxonomicName id="3F7B8F680C0600BEA8DE43F7FF1C9BA9" authorityName="comb. rest." baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="2449A8F18DE7E53805DDFB23E901EA06" italics="true" pageId="0" pageNumber="611">Coenosia nigerrima</emphasis>
</taxonomicName>
, male. 
<emphasis id="FD23655137E0496E18434ED39AA73A99" bold="true" pageId="0" pageNumber="611">A</emphasis>
Lateral view, 
<emphasis id="AD29E9975701A4F817B071141567B82A" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view, 
<emphasis id="1913A6182C8622DD59030DE9CC4B3B58" bold="true" pageId="0" pageNumber="611">C</emphasis>
thorax, dorsal view, 
<emphasis id="7C1300C147FDFE28C866943BCC461D20" bold="true" pageId="0" pageNumber="611">D</emphasis>
abdomen, dorsal view, 
<emphasis id="2F84E1E136365B69AB6A4804BF4CF3BE" bold="true" pageId="0" pageNumber="611">E</emphasis>
fifth sternite, 
<emphasis id="65BDEB6EC4073E79A88A97B37D0D1C15" bold="true" pageId="0" pageNumber="611">F</emphasis>
cercus, dorsal view (SEM), 
<emphasis id="40D7CA2F3A88503CBA1F2517A4C7601A" bold="true" pageId="0" pageNumber="611">G</emphasis>
cercus and surstylus, dorsal view, 
<emphasis id="B7A82D2DFFAB36120ABE414E2F9BB425" bold="true" pageId="0" pageNumber="611">H</emphasis>
cercus, surstylus and phallic complex, lateral view, 
<emphasis id="ECB867E7B9640ED8BE3E575F9236BB06" bold="true" pageId="0" pageNumber="611">I</emphasis>
phallic complex, lateral view, 
<emphasis id="B3D6042DECBF839617E1C57A53E5FF44" bold="true" pageId="0" pageNumber="611">J</emphasis>
phallic complex, detail of distal tip. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E-H), 0.05 mm (I-J). Hypandrium, spicules on ventral surface - red arrows.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="049E90B61598BC3D4355F109F84A95EB" pageId="0" pageNumber="611" type="female (fig. 22a, b)">
<paragraph id="0117C9A85ED8731D4384F609B84C2DB9" pageId="0" pageNumber="611">Female (Fig. 22A, B).</paragraph>
<paragraph id="0928D0CB766653F02F7E2E5CCE0363AE" pageId="0" pageNumber="611">
Length. Body: 3.75-4.10 mm, wing: 3.60-4.30 mm. Differs from male as follows: 
<emphasis id="0CAD8B177333CD96F2E2F1FF06AB9439" bold="true" italics="true" pageId="0" pageNumber="611">Wing</emphasis>
: Both calypters hyaline with white margins. 
<emphasis id="5A409528010E798123779F60DD14BDAE" bold="true" italics="true" pageId="0" pageNumber="611">Ovipositor</emphasis>
: Tergites 6, 7 and 8 with 2 long and wide scletotized plates; epiproct triangular, with hair-like setae, cercus longer than epiproct (Fig. 
<figureCitation id="FF2DE97C177A0E191E2A7801D747B198" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coenosia nigerrima, female. A: Lateral view, B head, frontal view, C ovipositor, dorsal view, distal tip, D ovipositor, ventral view, distal tip, E spermathecae. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E)." figureDoi="10.3897/asp.81.e104969.figure22" httpUri="https://binary.pensoft.net/fig/882090" pageId="0" pageNumber="611">22C</figureCitation>
). Sternites 6 and 7 with long plates; sternite 8 with a central long plate and 2 small plates each with 4-5 setae on distal margin; hypoproct triangular, with several short setae on distal margin (Fig. 
<figureCitation id="3766EEB26F2DAB07598BE67763A1B190" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coenosia nigerrima, female. A: Lateral view, B head, frontal view, C ovipositor, dorsal view, distal tip, D ovipositor, ventral view, distal tip, E spermathecae. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E)." figureDoi="10.3897/asp.81.e104969.figure22" httpUri="https://binary.pensoft.net/fig/882090" pageId="0" pageNumber="611">22D</figureCitation>
). Spermathecae (Fig. 
<figureCitation id="C624D81FCD9CB9DC840572641DDCE01C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Coenosia nigerrima, female. A: Lateral view, B head, frontal view, C ovipositor, dorsal view, distal tip, D ovipositor, ventral view, distal tip, E spermathecae. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E)." figureDoi="10.3897/asp.81.e104969.figure22" httpUri="https://binary.pensoft.net/fig/882090" pageId="0" pageNumber="611">22E</figureCitation>
).
</paragraph>
<caption id="B81635D1EA641D7841870F93501864AE" doi="10.3897/asp.81.e104969.figure22" httpUri="https://binary.pensoft.net/fig/882090" pageId="0" pageNumber="611" start="Figure 22" startId="F22">
<paragraph id="A911DD6F33D42E4CB537EBD1CEB1E6F4" pageId="0" pageNumber="611">
<emphasis id="ECBE5D6623443C60230CE7881A8E8D66" bold="true" pageId="0" pageNumber="611">Figure 22.</emphasis>
<taxonomicName id="8A95AA11AB753BC2DC90D2C86DE016B5" authorityName="comb. rest." baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="0782E80A28BCFC1B3585D6FADD889065" italics="true" pageId="0" pageNumber="611">Coenosia nigerrima</emphasis>
</taxonomicName>
, female. 
<emphasis id="B46EDC0701BE235272729389E8C52934" bold="true" pageId="0" pageNumber="611">A</emphasis>
: Lateral view, 
<emphasis id="EC2143FCA127CE88040D814089A1459D" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view, 
<emphasis id="70B90D44EDECD118A8E18AA0FCDEA082" bold="true" pageId="0" pageNumber="611">C</emphasis>
ovipositor, dorsal view, distal tip, 
<emphasis id="9EAAB97BCB495C5C1F9C57DDF6CBC5D2" bold="true" pageId="0" pageNumber="611">D</emphasis>
ovipositor, ventral view, distal tip, 
<emphasis id="1EECF2623E338AA677A3B9AB29CD6D82" bold="true" pageId="0" pageNumber="611">E</emphasis>
spermathecae. Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8574C496D8A07865D3AB42EC5EE0501F" pageId="0" pageNumber="611" type="type material">
<paragraph id="A9E2C89F31B5471A5BDD8ABDB37939B1" pageId="0" pageNumber="611">Type material.</paragraph>
<paragraph id="7672AD5B957BF09A83456902E7F5055B" pageId="0" pageNumber="611">
<emphasis id="928B272A02157B2DC188D3E209333A93" bold="true" pageId="0" pageNumber="611">
<emphasis id="DF849F539B92D1754718A0F9C71D9AE3" italics="true" pageId="0" pageNumber="611">Paratypes</emphasis>
.
</emphasis>
Two females; MNRJ. [pinned, without abdomen] Original labels: &quot;Casa Pangue / Llanquihue / Chile Dec 1926 / R&amp; E. Shannon&quot; print. on white paper, 
<normalizedToken id="9D775A2AEF00694047FAAA8996EBB35C" originalValue="“Paratypo”">&quot;Paratypo&quot;</normalizedToken>
print. on green paper, black frame; 
<normalizedToken id="3116CE2BF2F836AA0C99478E9684BDB8" originalValue="“♀”">&quot;♀&quot;</normalizedToken>
handwr. on white paper. &quot;M.N.R.J [print.] / 4401 [handwr.]&quot; on white paper, black frame. 
<emphasis id="37851F8CFACC3B3FFF5E5116792C0D7B" italics="true" pageId="0" pageNumber="611">Paratype</emphasis>
: male; MNRJ. [glued on card, without abdomen]. Original labels: &quot;Casa Pangue / llanquihue / Chile Dec 1926 / R&amp; E. Shannon&quot; print. on white paper, &quot; 
<taxonomicName id="3F8BDDA71EFC3697699739CE2924FC19" authorityName="Malloch" authorityYear="1920" class="Insecta" family="Muscidae" genus="Neodexiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neodexiopsis" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="genus">Neodexiopsis</taxonomicName>
/ 
<taxonomicName id="605C898A1A15C1455D26B501F5758C0A" baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Neodexiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neodexiopsis nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">Neodexiopsis nigerrima</taxonomicName>
(Mall.) [handwr.] / S.M. Lopes det. [print.]&quot; on wihte paper. 
<normalizedToken id="2237938E06506AB3E3175EE439AE0D6A" originalValue="“Paratypo”">&quot;Paratypo&quot;</normalizedToken>
print. on green paper, black frame; 
<normalizedToken id="6618037CF860A631FDE6454A60CE3AED" originalValue="“♀”">&quot;♀&quot;</normalizedToken>
handwr. on white paper. &quot;M.N.R.J [print.] / 4401 [handwr.]&quot; on white paper, black frame.
</paragraph>
</subSubSection>
<subSubSection id="67B36F7CA41C341F45794445BCE232B0" pageId="0" pageNumber="611" type="materials_examined">
<paragraph id="4BF1B344481200F40C4BDCFF80AECD6F" pageId="0" pageNumber="611">Additional material examined.</paragraph>
<paragraph id="B8DC39814BF91ACA84AF5AE0131A6454" pageId="0" pageNumber="611">
<materialsCitation id="08258C1B24E544BB6D9CC4EF7AAB643D" collectingDate="2014-10" collectorName="Mulieri Patitucci, Torretta, Malaise" country="ARGENTINA" elevation="538" latitude="-42.718803" location="Seccional Lago Verde" longLatPrecision="1" longitude="-71.72747" specimenCount="12" specimenCount-female="2" specimenCount-male="10" stateProvince="Chubut province">
<collectingCountry id="664A8A95D752681E6A6A60C19329BCE5" name="Argentina">ARGENTINA</collectingCountry>
- 
<emphasis id="BA397713AF62A5DB0D0C4E0460A4C009" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="F9A1E4489F85B0129D2A9D2B908BACC8" country="Argentina" name="Chubut">Chubut province</collectingRegion>
</emphasis>
• 
<specimenCount id="36FEA5D64D6D7D372B8CB6727FC6A20A" type="female">2 females</specimenCount>
, 
<specimenCount id="DCA784CE1B8045FD37CDDEC65BA6264A" type="male">10 males</specimenCount>
; PNLA, 
<location id="CE67F78A95A8FFEE25FE627DB409FE9D" LSID="urn:lsid:plazi:treatment:366E3BE326515983827B262999391B0E:CE67F78A95A8FFEE25FE627DB409FE9D" country="ARGENTINA" latitude="-42.718803" longLatPrecision="1" longitude="-71.72747" name="Seccional Lago Verde" stateProvince="Chubut province">Seccional Lago Verde</location>
; 
<geoCoordinate id="700D70B186AEEE5CC808C5C07345BB74" degrees="42.718803" direction="south" orientation="latitude" precision="1" value="-42.718803">-42.718803</geoCoordinate>
, 
<geoCoordinate id="606D76CB08757FB67FDBBA878428CE77" degrees="71.727470" direction="west" orientation="longitude" precision="1" value="-71.72747">-71.727470</geoCoordinate>
; 
<elevation id="FA44E129D7CAC3D82DE96FBC538C6223" metricMagnitude="2" metricUnit="m" metricValue="5.38" unit="m" value="538.0">
<quantity id="E815A48036071C4D49DBE44844F9422C" metricMagnitude="2" metricUnit="m" metricValue="5.38" unit="m" value="538.0">538 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate id="ABDC6DC29C055008094740E8ABB9EDF1" value="2014-10">Oct 2014</collectingDate>
, 
<collectorName id="2D097A67412681479DE6D06512B9B729">Mulieri Patitucci</collectorName>
&amp; 
<collectorName id="BE04069AED91B23FF2E7E582EE438576">Torretta</collectorName>
leg.; MACN; 
<collectorName id="FE26B2B52A8E002F9C2403607AAFFF30">Malaise</collectorName>
</materialsCitation>
• 
<materialsCitation id="10EE3121E175E78E9BAD4171B2670585" collectingDate="2011-01-11" collectorName="Patitucci" country="Argentina" elevation="200" latitude="-42.0963" location="Pitranto Grande" longLatPrecision="41" longitude="-71.612" specimenCount="1" specimenCount-female="1" stateProvince="Chubut">
<specimenCount id="473D6A87A3601A6BC33FDB28A34CF9E3" type="female">1 female</specimenCount>
; PNLP, 
<location id="202B7857824483100F849CB81EB3D4CD" LSID="urn:lsid:plazi:treatment:366E3BE326515983827B262999391B0E:202B7857824483100F849CB81EB3D4CD" country="Argentina" latitude="-42.0963" longLatPrecision="41" longitude="-71.612" name="Pitranto Grande" stateProvince="Chubut">Pitranto Grande</location>
; 
<geoCoordinate id="1C4B824238194E3323B89E41BB16C050" degrees="42.0963" direction="south" orientation="latitude" precision="5" value="-42.0963">-42.0963</geoCoordinate>
, 
<geoCoordinate id="620328158DA57ACAEBF1F552EF0664D9" degrees="71.612" direction="west" orientation="longitude" precision="55" value="-71.612">-71.612</geoCoordinate>
9; 
<elevation id="FC25D71859B88B7EAF76C2668776980D" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<quantity id="F0FBDCD0FF34FDB9C0012E221B32FD09" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate id="AF088C034EDC73DD6F72FA9E36FE4E06" value="2011-01-11">11 Jan 2011</collectingDate>
; 
<collectorName id="B7DC3CD9699FCCB3AE126A6D2C2B1719">Patitucci</collectorName>
leg.; MACN
</materialsCitation>
. - 
<materialsCitation id="29A16164F1F90AE0CF24DBFCF68743D9" collectingDate="1952-01" collectorName="Schajovskoi" country="Argentina" latitude="-40.155155" location="Pucara" longLatPrecision="1" longitude="-71.631615" specimenCount="2" specimenCount-female="2" stateProvince="Neuquen province">
<emphasis id="CE876EB39D851B0979F03B8357403F5E" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="0274794C45866F636F9C431CC9036EDE" country="Argentina" name="Neuquen">
<normalizedToken id="6D9502712849218A59C08EB628F63D09" originalValue="Neuquén">Neuquen</normalizedToken>
province
</collectingRegion>
</emphasis>
• 
<specimenCount id="86A3F712D22C0535CB80D3E29DF45227" type="female">2 females</specimenCount>
; PNL, 
<location id="0306510161EC2D267CDEF29F38EBBDFB" LSID="urn:lsid:plazi:treatment:366E3BE326515983827B262999391B0E:0306510161EC2D267CDEF29F38EBBDFB" country="Argentina" latitude="-40.155155" longLatPrecision="1" longitude="-71.631615" name="Pucara" stateProvince="Neuquen province">Pucara</location>
; 
<geoCoordinate id="62BA22570AE9065749D1A01C88438664" degrees="40.1551565" direction="south" orientation="latitude" precision="1" value="-40.155155">-40.1551565</geoCoordinate>
, 
<geoCoordinate id="DA12B1192938816EE00844B1E2AD40F3" degrees="71.63161488" direction="west" orientation="longitude" precision="1" value="-71.631615">-71.63161488</geoCoordinate>
; 
<collectingDate id="AAE87974349772142DAB8A97AF7FFF45" value="1952-01">Jan 1952</collectingDate>
; 
<collectorName id="BBAE237BBA8D7DDB70FCB6AE7E234F90">Schajovskoi</collectorName>
leg.; MACN
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1EFDD1570132D41B8B2F7235186C0D2A" pageId="0" pageNumber="611" type="distribution">
<paragraph id="80E776CEF0B0082C83B8154B525C4798" pageId="0" pageNumber="611">Distribution (Fig. 23).</paragraph>
<paragraph id="3FD0F8F11F9D96E5C90CA02653E6D0FE" pageId="0" pageNumber="611">
ARGENTINA: Chubut (new record), 
<normalizedToken id="124858ABD6C169832D572877FCB0666D" originalValue="Neuquén">Neuquen</normalizedToken>
(new record), 
<normalizedToken id="EBB416DE4152B7CACD8ED7FD6827D20B" originalValue="Río">Rio</normalizedToken>
Negro. CHILE: R. de Bio Bio, R. de los Lagos.
</paragraph>
<caption id="F9949EB38FEACC711CA772AB13C7F073" doi="10.3897/asp.81.e104969.figure23" httpUri="https://binary.pensoft.net/fig/882091" pageId="0" pageNumber="611" start="Figure 23" startId="F23">
<paragraph id="C52992A38BD84D4E79F508B18D246D2F" pageId="0" pageNumber="611">
<emphasis id="157D4393C4F20B116D5ED8EC48FE790C" bold="true" pageId="0" pageNumber="611">Figure 23.</emphasis>
Geographical distribution of 
<taxonomicName id="A94EF65684460599AFB8A75B9EB06340" authorityName="comb. rest." baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="874D9C1AD2642772FF0D28BEA497803D" italics="true" pageId="0" pageNumber="611">Coenosia nigerrima</emphasis>
</taxonomicName>
. 
<emphasis id="E9671EE43C557B52D4724F8B9937B2E0" bold="true" pageId="0" pageNumber="611">A</emphasis>
Subantarctic subregion records (Argentina, Chile), 
<emphasis id="3DA7F7402F571CAF537B4FC8608EB315" bold="true" pageId="0" pageNumber="611">B</emphasis>
Atlantic forest region records (Brazil). Biogeographic subantarctic subregion - green area, new records - black circle; bibliographic records - yellow triangle: bibliographic records in Brazil - black triangle.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="315CE2D3167233ED53BAF0DD85AD3680" pageId="0" pageNumber="611" type="remarks">
<paragraph id="8596C418DE535A73C6CA7EACA1D204C0" pageId="0" pageNumber="611">Remarks.</paragraph>
<paragraph id="5F74C0F1DBF278186FDFEE4DD85062E9" pageId="0" pageNumber="611">
<bibRefCitation id="4D67669DA4A7A0B33A238A3539032C9C" author="Malloch, JR" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="611" refId="B38" refString="Malloch, JR, 1934. Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, 171-346." title="Muscidae. In: Diptera of Patagonia and South Chile. Part 7, Fascicle 2. British Museum (Natural History), London, 171 - 346." year="1934">Malloch (1934)</bibRefCitation>
described 
<taxonomicName id="A7D8E6240725AA5EAF97D5DF422E6007" lsidName="C. nigerrima" pageId="0" pageNumber="611" rank="species" species="nigerrima">
<emphasis id="E743209D24F08ACE7692D1A2431D0AE8" italics="true" pageId="0" pageNumber="611">C. nigerrima</emphasis>
</taxonomicName>
with females and male specimens from Argentina and Chile, and proposed it as the type species of 
<taxonomicName id="97FA1A6F266EBB7CD66EEAB1BA78B8A9" authorityName="Malloch" authorityYear="1934" class="Insecta" family="Muscidae" genus="Austrocoenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrocoenosia" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="BAD3FFAC4D69A17A6D026858E708D19F" italics="true" pageId="0" pageNumber="611">Austrocoenosia</emphasis>
</taxonomicName>
. Also, this author highlighted the dark black coloration and the absence of marks on the thorax and abdomen, characters that differentiate this species from the remaining 
<taxonomicName id="7F15DB0711055BF7A2D925C2C0E66968" authorityName="Malloch" authorityYear="1934" class="Insecta" family="Muscidae" genus="Austrocoenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrocoenosia" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="06F431F5FCC50DC62EB8D563CB372712" italics="true" pageId="0" pageNumber="611">Austrocoenosia</emphasis>
</taxonomicName>
species. Later, 
<bibRefCitation id="25BF0009F52D0C8062A4BD32258496BE" author="Pont, AC" journalOrPublisher="In: A Catalogue of the Diptera of the Americas South of the United States, Museu de Zoologia, Universidade de Sao Paulo" pageId="0" pageNumber="611" pagination="1 - 111" refId="B51" refString="Pont, AC, 1972. Family Muscidae. In: A Catalogue of the Diptera of the Americas South of the United States, Museu de Zoologia, Universidade de Sao Paulo 97: 1 - 111" title="Family Muscidae." volume="97" year="1972">Pont (1972)</bibRefCitation>
transferred the species to 
<taxonomicName id="903BE5BD43220EEDEEFBAC306138036C" authorityName="Meigen" authorityYear="1826" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="923D36B820DC4ABB745B895955935CC8" italics="true" pageId="0" pageNumber="611">Coenosia</emphasis>
</taxonomicName>
. After that, 
<bibRefCitation id="8813EA129F38F79B6B7AA0DCDCD5FE04" author="Couri, MS" journalOrPublisher="Revista Brasileira de Biologia" pageId="0" pageNumber="611" pagination="499 - 517" refId="B14" refString="Couri, MS, Albuquerque, DDO, 1979. Estudos sobre Neodexiopsis Malloch, 1920 com notas nomenclaturais sobre Coenosiinae (Diptera, Muscidae). Revista Brasileira de Biologia 39: 499 - 517" title="Estudos sobre Neodexiopsis Malloch, 1920 com notas nomenclaturais sobre Coenosiinae (Diptera, Muscidae)." volume="39" year="1979">Couri and Albuquerque (1979)</bibRefCitation>
presented drawing of the female and male terminalia, and transferred the species to 
<taxonomicName id="0D390D78AD82218F8C9A0933164EC13B" authorityName="Malloch" authorityYear="1920" class="Insecta" family="Muscidae" genus="Neodexiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neodexiopsis" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="379A3F20B697C2ACCF466F9CB5D3EC41" italics="true" pageId="0" pageNumber="611">Neodexiopsis</emphasis>
</taxonomicName>
. The general black coloration, and the strongly sclerotized male terminalia make it difficult to correctly identify the species. In the specimens studied in the present work, we observed some differences (presence of keels on the cercus, shape of the surstylus) from the drawings presented by 
<bibRefCitation id="597F51A3F80725DC2D84516831109B18" author="Couri, MS" journalOrPublisher="Revista Brasileira de Biologia" pageId="0" pageNumber="611" pagination="499 - 517" refId="B14" refString="Couri, MS, Albuquerque, DDO, 1979. Estudos sobre Neodexiopsis Malloch, 1920 com notas nomenclaturais sobre Coenosiinae (Diptera, Muscidae). Revista Brasileira de Biologia 39: 499 - 517" title="Estudos sobre Neodexiopsis Malloch, 1920 com notas nomenclaturais sobre Coenosiinae (Diptera, Muscidae)." volume="39" year="1979">Couri and Albuquerque (1979)</bibRefCitation>
. We found morphological and phylogenetic evidence to consider this species as part of the 
<taxonomicName id="E65C8E0F1171487A3973B04896157BCA" lsidName="C. argentifrons" pageId="0" pageNumber="611" rank="species" species="argentifrons">
<emphasis id="A06EEDA3561D2E28CB28B0D0B08D0DE2" italics="true" pageId="0" pageNumber="611">C. argentifrons</emphasis>
</taxonomicName>
group, and thus restored the previous combination proposed by 
<bibRefCitation id="7F3562BCBE0C88B8BE2C433977CFFEA5" author="Pont, AC" journalOrPublisher="In: A Catalogue of the Diptera of the Americas South of the United States, Museu de Zoologia, Universidade de Sao Paulo" pageId="0" pageNumber="611" pagination="1 - 111" refId="B51" refString="Pont, AC, 1972. Family Muscidae. In: A Catalogue of the Diptera of the Americas South of the United States, Museu de Zoologia, Universidade de Sao Paulo 97: 1 - 111" title="Family Muscidae." volume="97" year="1972">Pont (1972)</bibRefCitation>
<taxonomicName id="8C9FE249A7610877D218AA2ED5EA97B0" authorityName="comb. rest." baseAuthorityName="Malloch" baseAuthorityYear="1934" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia nigerrima" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="40ECB61505F4D996F906B69554DF337D" italics="true" pageId="0" pageNumber="611">Coenosia nigerrima</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AFBD5C7032EEB8FAFCD0CF8E092DA920" pageId="0" pageNumber="611">
Regarding its distribution, several authors (
<bibRefCitation id="A5A04E00140A9A43F341BB476B6247D8" DOI="https://doi.org/10.1590/S0085-56262003000300005" author="Costacurta, NDC" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="611" pagination="389 - 397" refId="B10" refString="Costacurta, NDC, Marinoni, RC, Carvalho, CJD, 2003. Fauna de Muscidae (Diptera) em tres localidades do Estado do Parana, Brasil, capturada por armadilha Malaise. Revista Brasileira de Entomologia 47 (3): 389 - 397, DOI: https://doi.org/10.1590/S0085-56262003000300005" title="Fauna de Muscidae (Diptera) em tres localidades do Estado do Parana, Brasil, capturada por armadilha Malaise." url="https://doi.org/10.1590/S0085-56262003000300005" volume="47" year="2003">Costacurta et al. 2003</bibRefCitation>
; 
<bibRefCitation id="14E0A25872C6660EB8E3B2B6C36D2BC2" DOI="https://doi.org/10.1590/S0085-56262006000100014" author="Rodriguez-Fernandez, JI" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="611" pagination="93 - 100" refId="B58" refString="Rodriguez-Fernandez, JI, Carvalho, CJBD, Moura, MO, 2006. Estrutura de assembleias de Muscidae (Diptera) no Parana: uma analise por modelos nulos. Revista Brasileira de Entomologia 50 (1): 93 - 100, DOI: https://doi.org/10.1590/S0085-56262006000100014" title="Estrutura de assembleias de Muscidae (Diptera) no Parana: uma analise por modelos nulos." url="https://doi.org/10.1590/S0085-56262006000100014" volume="50" year="2006">
<normalizedToken id="0878638A28CB925FFECD0AF5CF40947D" originalValue="Rodríguez-Fernández">Rodriguez-Fernandez</normalizedToken>
et al. 2006
</bibRefCitation>
; 
<bibRefCitation id="B71BD9894E5020F86D961C7ACFD071DB" DOI="https://doi.org/10.1590/s1519-566x2010000300006" author="Krueger, RF" journalOrPublisher="Neotropical Entomology" pageId="0" pageNumber="611" pagination="345 - 353" refId="B35" refString="Krueger, RF, Carvalho, CJBD, Ribeiro, PB, 2010. Assembly rules in muscid fly assemblages in the grasslands biome of southern Brazil. Neotropical Entomology 39 (3): 345 - 353, DOI: https://doi.org/10.1590/s1519-566x2010000300006" title="Assembly rules in muscid fly assemblages in the grasslands biome of southern Brazil." url="https://doi.org/10.1590/s1519-566x2010000300006" volume="39" year="2010">
<normalizedToken id="CCE4DF9846B287968D63A6E295339257" originalValue="Krüger">Krueger</normalizedToken>
et al. 2010
</bibRefCitation>
; 
<bibRefCitation id="C0CF86E9694D5ECD31858AB4DC7B9FF2" DOI="https://doi.org/10.1016/j.jcz.2020.09.001" author="Fogaca, JM" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="611" pagination="1 - 7" refId="B22" refString="Fogaca, JM, Gomes, LRP, Carvalho, CJBD, 2020. Ultrastructural morphology and molecular analysis of a remarkable new species of Neodexiopsis (Muscidae, Diptera) from Southern Brazil. Zoologischer Anzeiger 289: 1 - 7, DOI: https://doi.org/10.1016/j.jcz.2020.09.001" title="Ultrastructural morphology and molecular analysis of a remarkable new species of Neodexiopsis (Muscidae, Diptera) from Southern Brazil." url="https://doi.org/10.1016/j.jcz.2020.09.001" volume="289" year="2020">
<normalizedToken id="5517610026329C94A50C3BA2FA3FD20B" originalValue="Fogaça">Fogaca</normalizedToken>
et al. 2020
</bibRefCitation>
) recorded this species in the South of Brazil (Fig. 
<figureCitation id="7FBC44854B69281700317C734E4DF64E" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Geographical distribution of Coenosia nigerrima. A Subantarctic subregion records (Argentina, Chile), B Atlantic forest region records (Brazil). Biogeographic subantarctic subregion - green area, new records - black circle; bibliographic records - yellow triangle: bibliographic records in Brazil - black triangle." figureDoi="10.3897/asp.81.e104969.figure23" httpUri="https://binary.pensoft.net/fig/882091" pageId="0" pageNumber="611">23B</figureCitation>
). Since species of the 
<taxonomicName id="4DD12F1C8C5AD158BE8C9CF2C9EF8AEA" lsidName="C. argentifrons" pageId="0" pageNumber="611" rank="species" species="argentifrons">
<emphasis id="59D3DF2CA063110F5FEC6E40070137A7" italics="true" pageId="0" pageNumber="611">C. argentifrons</emphasis>
</taxonomicName>
group presents restricted distribution in the Central Chilean and Subantarctic subregions; it would be interesting to examine the specimens collected in southern Brazil to establish similarities and differences.
</paragraph>
</subSubSection>
</treatment>
</document>