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<document id="09644BAF3DD35A2FFF5988E5994B8EB8" ID-CLB-Dataset="24800" ID-DOI="10.11646/zootaxa.4700.4.9" ID-GBIF-Dataset="14dfc815-148a-4ed7-a4c4-34af7019a1eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557635" ID-ZooBank="B62D48AE-1FC0-4126-B8D3-55EE34CA8F4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575035501164" checkinUser="plazi" docAuthor="Moravec, Jiří" docDate="2019" docId="366387F9FFC02A1C98DCF9E6BF5EF859" docLanguage="en" docName="zootaxa.4700.4.9.pdf.imf" docOrigin="Zootaxa 4700 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheilonycha Lacordaire 1842" docType="treatment" docVersion="9" lastPageNumber="504" masterDocId="CA5AFF81FFC22A1F984BFFBABA1AFFE5" masterDocTitle="Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense- 22. Genus Cheilonycha Lacordaire, 1842 (Coleoptera: Cicindelidae)" masterLastPageNumber="534" masterPageNumber="501" pageNumber="503" updateTime="1698769564174" updateUser="ExternalLinkService">
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<mods:title id="BD74385BEDBDA89FA28EA6381CF12D1A">Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense- 22. Genus Cheilonycha Lacordaire, 1842 (Coleoptera: Cicindelidae)</mods:title>
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<treatment id="366387F9FFC02A1C98DCF9E6BF5EF859" ID-GBIF-Taxon="160599128" LSID="urn:lsid:plazi:treatment:366387F9FFC02A1C98DCF9E6BF5EF859" httpUri="http://treatment.plazi.org/id/366387F9FFC02A1C98DCF9E6BF5EF859" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<subSubSection id="F6D06564FFC02A1D98DCF9E6B87EF993" box="[151,612,1628,1655]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="BE7536EFFFC02A1D98DCF9E6B87EF993" blockId="2.[151,612,1628,1655]" box="[151,612,1628,1655]" pageId="2" pageNumber="503">
<heading id="E53D8183FFC02A1D98DCF9E6B87EF993" bold="true" box="[151,612,1628,1655]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="8CBEEAFDFFC02A1D98DCF9E6B87EF993" bold="true" box="[151,612,1628,1655]" pageId="2" pageNumber="503">
Genus
<taxonomicName id="79CA4D6CFFC02A1D98A5F9E6B87EF993" ID-CoL="8KW4C" authority="Lacordaire, 1842" authorityName="Lacordaire" authorityYear="1842" box="[238,612,1628,1655]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC02A1D98A5F9E6BB9DF992" bold="true" box="[238,391,1628,1655]" italics="true" pageId="2" pageNumber="503">Cheilonycha</emphasis>
<bibRefCitation id="DA5B4B1EFFC02A1D99C4F9E6B87EF993" author="Lacordaire, J. T." box="[399,612,1628,1654]" pageId="2" pageNumber="503" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire, 1842</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="F6D06564FFC02A1D98DCF91BBE6CF95C" box="[151,1142,1696,1721]" pageId="2" pageNumber="503" type="reference_group">
<paragraph id="BE7536EFFFC02A1D98DCF91BBE6CF95C" blockId="2.[151,1151,1696,1905]" box="[151,1142,1696,1721]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF91BBE6CF95C" box="[151,1142,1696,1721]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF91BBBEBF95C" ID-CoL="8KW4C" authority="Lacordaire, 1842: 28" authorityName="Lacordaire" authorityPageNumber="28" authorityYear="1842" box="[151,497,1696,1721]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC02A1D98DCF91BBB0DF95C" box="[151,279,1697,1721]" italics="true" pageId="2" pageNumber="503">Cheilonycha</emphasis>
<treatmentCitation id="3F6B10FEFFC02A1D9955F91ABBEBF95C" author="Lacordaire, J. T." box="[286,497,1696,1721]" page="28" pageId="2" pageNumber="503" year="1842">
<bibRefCitation id="DA5B4B1EFFC02A1D9955F91ABBEBF95C" author="Lacordaire, J. T." box="[286,497,1696,1721]" pageId="2" pageNumber="503" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire, 1842: 28</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(reprint Quadart, the paper (Liége) was published in 1843: 92).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F6D06564FFC02A1D98DCF97ABE65F93D" box="[151,1151,1727,1752]" pageId="2" pageNumber="503" type="type_taxon">
<paragraph id="BE7536EFFFC02A1D98DCF97ABE65F93D" blockId="2.[151,1151,1696,1905]" box="[151,1151,1727,1752]" pageId="2" pageNumber="503">
<emphasis id="8CBEEAFDFFC02A1D98DCF97ABB04F93D" bold="true" box="[151,286,1727,1752]" pageId="2" pageNumber="503">
<typeStatus id="6171884DFFC02A1D98DCF97ABAD6F93D" box="[151,204,1728,1752]" pageId="2" pageNumber="503">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="79CA4D6CFFC02A1D9961F905B866F93D" ID-CoL="5Z55V" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[298,636,1727,1752]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC02A1D9961F905BBF4F932" box="[298,494,1727,1751]" italics="true" pageId="2" pageNumber="503">Cicindela chalybea</emphasis>
<bibRefCitation id="DA5B4B1EFFC02A1D99BFF905B866F93D" author="Dejean, P. M. F. A." box="[500,636,1727,1752]" pageId="2" pageNumber="503" refId="ref18541" refString="Dejean, P. M. F. A. (1825) Species general des Coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp. https: // doi. org / 10.5962 / bhl. title. 2086" type="book" year="1825">Dejean, 1825</bibRefCitation>
</taxonomicName>
(by original monotypy—see “Discussion” below).
</paragraph>
</subSubSection>
<subSubSection id="F6D06564FFC02A1D98DCF964BBE1F894" pageId="2" pageNumber="503" type="reference_group">
<paragraph id="BE7536EFFFC02A1D98DCF964B8EFF913" blockId="2.[151,1151,1696,1905]" box="[151,757,1757,1782]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF964B8EFF913" box="[151,757,1757,1782]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF964BB16F913" authority="Agassiz 1846" authorityName="Agassiz" authorityYear="1846" box="[151,268,1758,1782]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC02A1D98DCF964BB16F913" box="[151,268,1758,1782]" italics="true" pageId="2" pageNumber="503">Chilonycha</emphasis>
</taxonomicName>
:
<treatmentCitation id="3F6B10FEFFC02A1D9953F967BBD1F913" author="Agassiz, L." box="[280,459,1757,1782]" page="78" pageId="2" pageNumber="503" year="1846">
<bibRefCitation id="DA5B4B1EFFC02A1D9953F967BBD1F913" author="Agassiz, L." box="[280,459,1757,1782]" pageId="2" pageNumber="503" refId="ref18141" refString="Agassiz, L. (1846) Nomenclatoris Zoologici, Index Universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium mnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Jent et Gassmann, Soloduri, viii + 393 pp." type="book" year="1846">Agassiz 1846: 78</bibRefCitation>
</treatmentCitation>
, 80 (unjustified emendation).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE7536EFFFC02A1D98DCF947B89DF8F0" blockId="2.[151,1151,1696,1905]" box="[151,647,1789,1813]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF947B89DF8F0" box="[151,647,1789,1813]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF947BB0CF8F0" ID-CoL="8LSDX" authorityName="Agassiz" authorityYear="1846" box="[151,278,1789,1813]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC02A1D98DCF947BB0CF8F0" box="[151,278,1789,1813]" italics="true" pageId="2" pageNumber="503">Odontochila</emphasis>
</taxonomicName>
group
<taxonomicName id="79CA4D6CFFC02A1D9915F947BBC9F8F0" authority="Horn 1910" authorityName="Horn" authorityYear="1910" box="[350,467,1789,1813]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC02A1D9915F947BBC9F8F0" box="[350,467,1789,1813]" italics="true" pageId="2" pageNumber="503">Chilonycha</emphasis>
</taxonomicName>
:
<treatmentCitation id="3F6B10FEFFC02A1D99ABF947B899F8F0" author="Horn, W." box="[480,643,1789,1813]" page="203" pageId="2" pageNumber="503" year="1910">
<bibRefCitation id="DA5B4B1EFFC02A1D99ABF947B899F8F0" author="Horn, W." box="[480,643,1789,1813]" pageId="2" pageNumber="503" pagination="103 - 208" refId="ref18907" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P. Genera Insectorum, 82 B, 103 - 208, pls. 6 - 15." type="book chapter" year="1910">Horn 1910: 203</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE7536EFFFC02A1D98DCF8A1BBDBF8D6" blockId="2.[151,1151,1696,1905]" box="[151,449,1819,1843]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF8A1BBDBF8D6" box="[151,449,1819,1843]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF8A1BB16F8D6" authority="Horn 1910" authorityName="Horn" authorityYear="1910" box="[151,268,1819,1843]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC02A1D98DCF8A1BB16F8D6" box="[151,268,1819,1843]" italics="true" pageId="2" pageNumber="503">Chilonycha</emphasis>
</taxonomicName>
:
<treatmentCitation id="3F6B10FEFFC02A1D9952F8A6BBA7F8D6" author="Horn, W." box="[281,445,1819,1843]" page="203" pageId="2" pageNumber="503" year="1910">
<bibRefCitation id="DA5B4B1EFFC02A1D9952F8A6BBA7F8D6" author="Horn, W." box="[281,445,1819,1843]" pageId="2" pageNumber="503" pagination="103 - 208" refId="ref18907" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P. Genera Insectorum, 82 B, 103 - 208, pls. 6 - 15." type="book chapter" year="1910">Horn 1910: 203</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE7536EFFFC02A1D98DCF880B84DF8B7" blockId="2.[151,1151,1696,1905]" box="[151,599,1850,1874]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF880B84DF8B7" box="[151,599,1850,1874]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF880BBBEF8B7" authority="" box="[151,420,1850,1874]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC02A1D98DCF880BB0CF8B7" box="[151,278,1850,1874]" italics="true" pageId="2" pageNumber="503">Odontochila</emphasis>
(
<emphasis id="8CBEEAFDFFC02A1D996EF880BB80F8B7" box="[293,410,1850,1874]" italics="true" pageId="2" pageNumber="503">Chilonycha</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="3F6B10FEFFC02A1D99E4F881B849F8B7" author="Horn, W." box="[431,595,1850,1874]" page="124" pageId="2" pageNumber="503" year="1934">
<bibRefCitation id="DA5B4B1EFFC02A1D99E4F881B849F8B7" author="Horn, W." box="[431,595,1850,1874]" pageId="2" pageNumber="503" pagination="124 - 125" refId="ref19173" refString="Horn, W. (1934) On the three forms of Odontochila subgenus Chilonycha (Cicind., Col). Extracto de la Revista Chilena de Historia Natural, 38, 124 - 125." type="journal article" year="1934">Horn 1934: 124</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE7536EFFFC02A1D98DCF8E3BBE1F894" blockId="2.[151,1151,1696,1905]" box="[151,507,1880,1905]" pageId="2" pageNumber="503">
<treatmentCitationGroup id="9EDA11C1FFC02A1D98DCF8E3BBE1F894" box="[151,507,1880,1905]" pageId="2" pageNumber="503">
<taxonomicName id="79CA4D6CFFC02A1D98DCF8E3BBECF894" ID-CoL="8KW4C" authority=": Reichardt 1977: 374" authorityName=": Reichardt" authorityPageNumber="374" authorityYear="1977" box="[151,502,1880,1905]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC02A1D98DCF8E3BB0DF894" box="[151,279,1881,1905]" italics="true" pageId="2" pageNumber="503">Cheilonycha</emphasis>
:
<treatmentCitation id="3F6B10FEFFC02A1D996EF8E2BBECF894" author="Reichardt, H." box="[293,502,1880,1905]" page="374" pageId="2" pageNumber="503" year="1977">
<bibRefCitation id="DA5B4B1EFFC02A1D996EF8E2BBECF894" author="Reichardt, H." box="[293,502,1880,1905]" pageId="2" pageNumber="503" pagination="346 - 493" refId="ref22249" refString="Reichardt, H. (1977) A synopsis of the genera of Neotropical Carabidae (Insecta: Coleoptera). Questiones Entomologicae, 13, 346 - 493." type="journal article" year="1977">Reichardt 1977: 374</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F6D06564FFC02A1C98DCF824BAD0FD62" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="diagnosis">
<paragraph id="BE7536EFFFC02A1C98DCF824BAD0FD62" blockId="2.[151,1437,1950,2014]" lastBlockId="3.[151,1437,153,1980]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<emphasis id="8CBEEAFDFFC02A1D98DCF824BB8DF85C" bold="true" box="[151,407,1950,1977]" pageId="2" pageNumber="503">Differential diagnosis</emphasis>
. Protarsi sexually dimorphic, first three protarsomeres in male dilated; body medium-sized with markedly wide elytra in comparison to head and pronotum, glabrous, either uniformly black-blue to cyaneous-blue, usually with violaceous lustre, or with chatoyant green, black-blue, or deep violaceous-blue head and pronotum while elytra shiny red or gold-bronze, usually with shiny green lateral areas and shiny red elytral disc, completely lacking white maculation; elytral disc mostly distinctly convex with distinct posterior declivity; elytral apices rounded towards small sutural spine, microserrulation extremely fine, indistinct, irregular or partly absent; elytral punctation either almost regular and dense, or punctures rather spaced and partly irregular; pronotum glabrous, slightly but notably wider than long, discal surface and surface of head finely or extremely finely and densely vermicular-wavy to zigzag-wavy striate-rugulose; palpi and antennomeres 14 metallic-black with various iridescence; labrum primarily 4-setose, metallic blue or blue-green throughout, with various violet or green iridescence, shape sexually dimorphic: male labrum either rather short with effaced or rounded anterolateral and outer anterior teeth and with only median tooth, or with notably anteriad-prolonged tridentate anteromedian lobe; female labrum with prolonged tridentate median lobe and more or less distinctly prolonged median tooth; mandibles normally shaped with arcuate lateral margins, subsymmetrical, each mandible in both sexes with four teeth (and basal molar); aedeagus elongate, widest in middle, apical half attenuated towards rounded apex; internal sac lacking any flagellum.
</paragraph>
</subSubSection>
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<paragraph id="BE7536EFFFC12A1C988CFD2ABE2BFD4E" blockId="3.[151,1437,153,1980]" box="[199,1073,656,683]" pageId="3" pageNumber="504">
<emphasis id="8CBEEAFDFFC12A1C988CFD2ABB43FD4E" bold="true" box="[199,345,656,683]" pageId="3" pageNumber="504">Distribution</emphasis>
.
<collectingCountry id="C6DD767FFFC12A1C992CFD2BBBB4FD4E" box="[359,430,657,683]" name="Brazil" pageId="3" pageNumber="504">Brazil</collectingCountry>
,
<collectingCountry id="C6DD767FFFC12A1C99F1FD2BB839FD4E" box="[442,547,657,683]" name="Paraguay" pageId="3" pageNumber="504">Paraguay</collectingCountry>
,
<collectingCountry id="C6DD767FFFC12A1C9A66FD2BB886FD4E" box="[557,668,657,683]" name="Argentina" pageId="3" pageNumber="504">Argentina</collectingCountry>
and
<collectingCountry id="C6DD767FFFC12A1C9A98FD2BB932FD4E" box="[723,808,657,683]" name="Bolivia" pageId="3" pageNumber="504">Bolivia</collectingCountry>
, habitat termitophilous.
</paragraph>
</subSubSection>
<subSubSection id="F6D06564FFC12A1C988CFD0EBF5EF859" pageId="3" pageNumber="504" type="discussion">
<paragraph id="BE7536EFFFC12A1C988CFD0EBE5DFCD9" blockId="3.[151,1437,153,1980]" pageId="3" pageNumber="504">
<emphasis id="8CBEEAFDFFC12A1C988CFD0EBB5FFD2A" bold="true" box="[199,325,692,719]" pageId="3" pageNumber="504">Discussion</emphasis>
. The original description of the genus
<taxonomicName id="79CA4D6CFFC12A1C9B49FD0FB98AFD2A" authorityName=": Reichardt" authorityYear="1977" box="[770,912,693,719]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9B49FD0FB98AFD2A" box="[770,912,693,719]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
was first published in a reprint and spread by
<bibRefCitation id="DA5B4B1EFFC12A1C98DCFD63BB7CFD16" author="Lacordaire, J. T." box="[151,358,729,755]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire (1842)</bibRefCitation>
. Although the main publication came out one year later (
<bibRefCitation id="DA5B4B1EFFC12A1C9BA1FD63BEB2FD16" author="Lacordaire, J. T." box="[1002,1192,729,755]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19368" refString="Lacordaire, J. T. (1843) Revision de la famille des Cicindelides (Cicindelidae) de l'ordre des Coleopteres, accompagnee de la creation de quelques genres nouveaux Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118." type="journal article" year="1843">Lacordaire 1843</bibRefCitation>
), the publication date should be 1842. It is also in accordance with the genus-group name
<taxonomicName id="79CA4D6CFFC12A1C9BC6FD47BEF9FCF2" authority="Lacordaire, 1842" authorityName="Lacordaire" authorityYear="1842" box="[909,1251,765,791]" class="Insecta" family="Carabidae" genus="Phyllodroma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9BC6FD47BE04FCF2" box="[909,1054,765,791]" italics="true" pageId="3" pageNumber="504">Phyllodroma</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C9C6FFD47BEF9FCF2" author="Lacordaire, J. T." box="[1060,1251,765,791]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire, 1842</bibRefCitation>
</taxonomicName>
published in the same reprint (see also
<bibRefCitation id="DA5B4B1EFFC12A1C99D9FC9BB82BFCDE" author="Lorenz, W." box="[402,561,800,827]" pageId="3" pageNumber="504" refId="ref19835" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. Wolfgang Lorenz, Tutzing, iv + 530 pp." type="book" year="2005">Lorenz 2005a</bibRefCitation>
,b,
<bibRefCitation id="DA5B4B1EFFC12A1C9A04FC9BB955FCDE" author="Erwin, T. L. &amp; Pearson, D. L." box="[591,847,801,827]" pageId="3" pageNumber="504" refId="ref18698" refString="Erwin, T. L. &amp; Pearson, D. L. (2008) A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of the life. Vol. II. Carabidae-Nebriformes 2 - Cicindelitae. Pensoft Series Faunistica 84. Pensoft Publishers, Sofia, 366 pp., 33 pls." type="book" year="2008">Erwin &amp; Pearson 2008</bibRefCitation>
and
<bibRefCitation id="DA5B4B1EFFC12A1C9BCDFC9BBE21FCD9" author="Moravec, J." box="[902,1083,801,828]" pageId="3" pageNumber="504" refId="ref20947" refString="Moravec, J. (2018 b) Taxonomic revision of the Neotropical tiger beetle genera of the subtribe Odontocheilina. Volume 1. Odontocheila Laporte de Castelnau, Cenothyla Rivalier and Phyllodroma Lacordaire. (Coleoptera: Cicindelidae). Biosfericka rezervace Dolni Morava, o. p. s., Lednice na Morave, 623 + 2 pp." type="book" year="2018">Moravec 2018b</bibRefCitation>
).
</paragraph>
<paragraph id="BE7536EFFFC12A1C988CFCFFBE15F9B6" blockId="3.[151,1437,153,1980]" pageId="3" pageNumber="504">
<bibRefCitation id="DA5B4B1EFFC12A1C988CFCFFBB89FCBA" author="Lacordaire, J. T." box="[199,403,837,863]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire (1842)</bibRefCitation>
based his original description of the genus
<taxonomicName id="79CA4D6CFFC12A1C9BC9FCFFBE0AFCBA" authorityName=": Reichardt" authorityYear="1977" box="[898,1040,837,863]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9BC9FCFFBE0AFCBA" box="[898,1040,837,863]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
on
<taxonomicName id="79CA4D6CFFC12A1C9C70FCFFBECAFCBA" box="[1083,1232,837,863]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9C70FCFFBECAFCBA" box="[1083,1232,837,863]" italics="true" pageId="3" pageNumber="504">Ch. chalybea</emphasis>
</taxonomicName>
(originally
<taxonomicName id="79CA4D6CFFC12A1C9D1DFCFFBBCAFC66" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9D1DFCFFBB28FC66" italics="true" pageId="3" pageNumber="504">Cicindela chalybea</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C9973FCD3BBCAFC66" author="Dejean, P. M. F. A." box="[312,464,872,899]" pageId="3" pageNumber="504" refId="ref18541" refString="Dejean, P. M. F. A. (1825) Species general des Coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp. https: // doi. org / 10.5962 / bhl. title. 2086" type="book" year="1825">Dejean, 1825</bibRefCitation>
</taxonomicName>
). He mentioned under the description:
<emphasis id="8CBEEAFDFFC12A1C9B36FCD3BE0DFC42" italics="true" pageId="3" pageNumber="504">“je ne connais que lespece brésilienne ci-dessus qui puisse rentrer dans ce genre. Me exemplaires viennent de la province Goyaz”</emphasis>
(freely translated: “
<emphasis id="8CBEEAFDFFC12A1C9CBFFC34BEBBFC2E" italics="true" pageId="3" pageNumber="504">I only know the Brazilian species above that can fit into this genus. My specimens come from Goyaz province</emphasis>
”). The last sentence is somewhat puzzling, because
<taxonomicName id="79CA4D6CFFC12A1C9994FC6FB869FC0A" box="[479,627,981,1007]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9994FC6FB869FC0A" box="[479,627,981,1007]" italics="true" pageId="3" pageNumber="504">Ch. chalybea</emphasis>
</taxonomicName>
has not been recorded from the state of
<collectingRegion id="7C0EF80DFFC12A1C9C71FC6FBE61FC0A" box="[1082,1147,981,1007]" country="Brazil" name="Goias" pageId="3" pageNumber="504">Goiás</collectingRegion>
(= Goyaz). Nevertheless, the state is very large and has its long boundary with the state of
<collectingRegion id="7C0EF80DFFC12A1C9B30FC43BE08FBF6" box="[891,1042,1017,1043]" country="Brazil" name="Minas Gerais" pageId="3" pageNumber="504">Minas Gerais</collectingRegion>
where
<taxonomicName id="79CA4D6CFFC12A1C9C2CFC43BEEEFBF6" box="[1127,1268,1017,1043]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9C2CFC43BEEEFBF6" box="[1127,1268,1017,1043]" italics="true" pageId="3" pageNumber="504">Ch chalybea</emphasis>
</taxonomicName>
occurs. In any event, a possibility that Lacordaire misidentified
<taxonomicName id="79CA4D6CFFC12A1C9A8DFBA7B94EFBD2" box="[710,852,1053,1079]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9A8DFBA7B94EFBD2" box="[710,852,1053,1079]" italics="true" pageId="3" pageNumber="504">Ch chalybea</emphasis>
</taxonomicName>
with
<taxonomicName id="79CA4D6CFFC12A1C9BDCFBA7BE25FBD2" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[919,1087,1053,1079]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="8CBEEAFDFFC12A1C9BDCFBA7BE25FBD2" box="[919,1087,1053,1079]" italics="true" pageId="3" pageNumber="504">Ch. auripennis</emphasis>
</taxonomicName>
which was described 14 years later and was therefore unknown to him, would be a mere speculation, because there is no evidence of such confusion. Unfortunately, Lacordaire did not describe the dorsal body coloration which immediately differentiates the species, while the other characters are basically shared with all species of the genus. It must be noted here that it is impossible to trace the specimens which Lacordaire had in his hand, and, moreover, they might have been mislabelled. Furthermore, the confusion was almost impossible, because it was improbable that Lacordaire overlooked the fundamental distinguishing character of
<taxonomicName id="79CA4D6CFFC12A1C9ACDFB4FB903FAEA" box="[646,793,1269,1295]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9ACDFB4FB903FAEA" box="[646,793,1269,1295]" italics="true" pageId="3" pageNumber="504">Ch. chalybea</emphasis>
</taxonomicName>
, which is the uniformly blue coloration clearly described by
<bibRefCitation id="DA5B4B1EFFC12A1C98F1FAA3BB41FAD6" author="Dejean, P. M. F. A." box="[186,347,1304,1331]" pageId="3" pageNumber="504" refId="ref18541" refString="Dejean, P. M. F. A. (1825) Species general des Coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp. https: // doi. org / 10.5962 / bhl. title. 2086" type="book" year="1825">Dejean (1825)</bibRefCitation>
as “
<emphasis id="8CBEEAFDFFC12A1C99C7FAA3BF92FAD6" box="[396,1416,1305,1331]" italics="true" pageId="3" pageNumber="504">Elle est entiérement tant en-dessus quen dessous, dune belle couleur bleue assez brillante</emphasis>
”. Nevertheless,
<bibRefCitation id="DA5B4B1EFFC12A1C997DFA87BBADFAB2" author="Horn, W." box="[310,439,1340,1367]" pageId="3" pageNumber="504" pagination="209 - 487" refId="ref18947" refString="Horn, W. (1915) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, 82 C, 209 - 487, pls. 16 - 23." type="book chapter" year="1915">Horn (1915</bibRefCitation>
and repeatedly 1926a), giving no reasons for his decision but probably influenced by the occurrence in
<collectingRegion id="7C0EF80DFFC12A1C9973FADBBB63FA9E" box="[312,377,1377,1403]" country="Brazil" name="Goias" pageId="3" pageNumber="504">Goiás</collectingRegion>
(Goyaz) mentioned by
<bibRefCitation id="DA5B4B1EFFC12A1C9ACCFADBB956FA9E" author="Lacordaire, J. T." box="[647,844,1377,1403]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire (1842</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9B13FADBB989FA9E" author="Lacordaire, J. T." box="[856,915,1377,1403]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19368" refString="Lacordaire, J. T. (1843) Revision de la famille des Cicindelides (Cicindelidae) de l'ordre des Coleopteres, accompagnee de la creation de quelques genres nouveaux Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118." type="journal article" year="1843">1843</bibRefCitation>
), gave the
<typeStatus id="6171884DFFC12A1C9C44FADBBE25FA9E" box="[1039,1087,1377,1403]" pageId="3" pageNumber="504">type</typeStatus>
species of
<taxonomicName id="79CA4D6CFFC12A1C9C8BFADBBF54FA9E" authorityName=": Reichardt" authorityYear="1977" box="[1216,1358,1377,1403]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9C8BFADBBF54FA9E" box="[1216,1358,1377,1403]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
as
<taxonomicName id="79CA4D6CFFC12A1C9D3EFADBBBBCFA7A" authority="Lucas, 1857" authorityName="Lucas" authorityYear="1857" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="auripennis">
<emphasis id="8CBEEAFDFFC12A1C9D3EFADBBF8CFA9E" box="[1397,1430,1377,1403]" italics="true" pageId="3" pageNumber="504">Ch</emphasis>
.
<emphasis id="8CBEEAFDFFC12A1C98DCFA3CBB15FA7A" box="[151,271,1414,1439]" italics="true" pageId="3" pageNumber="504">auripennis</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C9953FA3FBBBCFA7A" author="Lucas, P. H." box="[280,422,1412,1439]" pageId="3" pageNumber="504" refId="ref19939" refString="Lucas, P. H. (1857 - 1859) Animaux nouveaux ou rares recueillis pendant l'expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para. Entomologie. P. Bertrand, Paris, 204 pp., 20 pls." type="book" year="1857">Lucas, 1857</bibRefCitation>
</taxonomicName>
. From the nomenclatorial point of view, this treatment by
<bibRefCitation id="DA5B4B1EFFC12A1C9C0FFA3FBED0FA7A" author="Horn, W." box="[1092,1226,1412,1439]" pageId="3" pageNumber="504" pagination="209 - 487" refId="ref18947" refString="Horn, W. (1915) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, 82 C, 209 - 487, pls. 16 - 23." type="book chapter" year="1915">Horn (1915</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9C93FA3FBF04FA45" author="Horn, W." box="[1240,1310,1413,1440]" pageId="3" pageNumber="504" refId="ref19064" refString="Horn, W. (1926 a) Carabidae, Cicindelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, 345 pp." type="book" year="1926">1926a</bibRefCitation>
) cannot be accepted as a justified subsequent
<typeStatus id="6171884DFFC12A1C9A5CFA13B85DFA26" box="[535,583,1449,1475]" pageId="3" pageNumber="504">type</typeStatus>
designation, because
<bibRefCitation id="DA5B4B1EFFC12A1C9B77FA13BE1DFA26" author="Lacordaire, J. T." box="[828,1031,1449,1475]" pageId="3" pageNumber="504" pagination="85 - 118" refId="ref19310" refString="Lacordaire, J. T. (1842) Revision de la famille des Cicindelides de l'ordre des Coleopteres. Quadart, Liege, 40 pp. [published in 1843 in Memoires de la Societe Royale des Sciences de Liege, 1, 85 - 118, however, reprints Quadart, Liege appeared in 1842]" type="book chapter" year="1842">Lacordaire (1842)</bibRefCitation>
mentioned only the name
<taxonomicName id="79CA4D6CFFC12A1C9D64FA13BB86FA02" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9D64FA13BAE6FA02" italics="true" pageId="3" pageNumber="504">Cicindela chalybea</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C994EFA77BB86FA02" author="Dejean, P. M. F. A." box="[261,412,1484,1511]" pageId="3" pageNumber="504" refId="ref18541" refString="Dejean, P. M. F. A. (1825) Species general des Coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp. https: // doi. org / 10.5962 / bhl. title. 2086" type="book" year="1825">Dejean, 1825</bibRefCitation>
</taxonomicName>
on which he based his description of his new genus
<taxonomicName id="79CA4D6CFFC12A1C9BB1FA77BE92FA02" authorityName=": Reichardt" authorityYear="1977" box="[1018,1160,1485,1511]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9BB1FA77BE92FA02" box="[1018,1160,1485,1511]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
—see Art. 67.2.1 (
<bibRefCitation id="DA5B4B1EFFC12A1C9D12FA77BAC9F9EE" author="ICZN" pageId="3" pageNumber="504" refId="ref19236" refString="ICZN (1999) International Code of Zoological Nomenclature, fourth edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
). Not only that Lacordaire clearly referred to
<taxonomicName id="79CA4D6CFFC12A1C9A99FA4BB97FF9EE" box="[722,869,1521,1547]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9A99FA4BB97FF9EE" box="[722,869,1521,1547]" italics="true" pageId="3" pageNumber="504">Ch. chalybea</emphasis>
</taxonomicName>
and no other species-name was mentioned by him either in the private reprint spread in 1842 or in the main paper published in 1843, but as discussed above, there is no evidence that he confused
<taxonomicName id="79CA4D6CFFC12A1C99A8F983B86DF9B6" box="[483,631,1593,1619]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C99A8F983B86DF9B6" box="[483,631,1593,1619]" italics="true" pageId="3" pageNumber="504">Ch. chalybea</emphasis>
</taxonomicName>
with any other species of the genus.
</paragraph>
<paragraph id="BE7536EFFFC12A1C988CF9E7B9E5F97E" blockId="3.[151,1437,153,1980]" pageId="3" pageNumber="504">
Recently also
<bibRefCitation id="DA5B4B1EFFC12A1C992FF9E7B813F992" author="Lorenz, W." box="[356,521,1629,1655]" pageId="3" pageNumber="504" refId="ref19739" refString="Lorenz, W. (1998 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 502 pp." type="book" year="1998">Lorenz (1998a</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9A58F9E6B843F992" author="Lorenz, W." box="[531,601,1628,1655]" pageId="3" pageNumber="504" refId="ref19835" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. Wolfgang Lorenz, Tutzing, iv + 530 pp." type="book" year="2005">2005a</bibRefCitation>
) and
<bibRefCitation id="DA5B4B1EFFC12A1C9ADEF9E7B9BFF992" author="Erwin, T. L. &amp; Pearson, D. L." box="[661,933,1629,1655]" pageId="3" pageNumber="504" refId="ref18698" refString="Erwin, T. L. &amp; Pearson, D. L. (2008) A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of the life. Vol. II. Carabidae-Nebriformes 2 - Cicindelitae. Pensoft Series Faunistica 84. Pensoft Publishers, Sofia, 366 pp., 33 pls." type="book" year="2008">Erwin &amp; Pearson (2008)</bibRefCitation>
have
<taxonomicName id="79CA4D6CFFC12A1C9BAEF9E7BF4DF992" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[997,1367,1628,1655]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="chalybea">
<emphasis id="8CBEEAFDFFC12A1C9BAEF9E7BEA7F992" box="[997,1213,1629,1655]" italics="true" pageId="3" pageNumber="504">Cicindela chalybea</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C9C88F9E7BF4DF992" author="Dejean, P. M. F. A." box="[1219,1367,1628,1655]" pageId="3" pageNumber="504" refId="ref18541" refString="Dejean, P. M. F. A. (1825) Species general des Coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp. https: // doi. org / 10.5962 / bhl. title. 2086" type="book" year="1825">Dejean, 1825</bibRefCitation>
</taxonomicName>
as the
<typeStatus id="6171884DFFC12A1C98DCF93BBADDF97E" box="[151,199,1665,1691]" pageId="3" pageNumber="504">type</typeStatus>
species of
<taxonomicName id="79CA4D6CFFC12A1C990EF93BBBC9F97E" authorityName=": Reichardt" authorityYear="1977" box="[325,467,1665,1691]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C990EF93BBBC9F97E" box="[325,467,1665,1691]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
in accordance with the reasons mentioned above.
</paragraph>
<paragraph id="BE7536EFFFC12A1C988CF91FBF5EF859" blockId="3.[151,1437,153,1980]" pageId="3" pageNumber="504">
<bibRefCitation id="DA5B4B1EFFC12A1C988CF91FBBBFF95A" author="Blackwelder, R. E." box="[199,421,1701,1727]" pageId="3" pageNumber="504" pagination="1 - 188" refId="ref18300" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, The west Indies, and South Amer- ica Part 1. Smithsonian Institution United States National Museum, Bulletin, 185, 1 - 188. https: // doi. org / 10.5479 / si. 03629236.185. i" type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
inadequately treated the genus-group name (as “subg.
<taxonomicName id="79CA4D6CFFC12A1C9C4BF91FBE9BF95A" box="[1024,1153,1701,1727]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC12A1C9C4BF91FBE9BF95A" box="[1024,1153,1701,1727]" italics="true" pageId="3" pageNumber="504">Chilonycha</emphasis>
</taxonomicName>
”) as a synonym of
<emphasis id="8CBEEAFDFFC12A1C9D1EF91FBAEBF906" italics="true" pageId="3" pageNumber="504">Odon- tocheila</emphasis>
<bibRefCitation id="DA5B4B1EFFC12A1C98B1F973B82BF906" author="Laporte de Castelnau, F. L." box="[250,561,1737,1763]" pageId="3" pageNumber="504" pagination="27 - 38" refId="ref19415" refString="Laporte de Castelnau, F. L. (1834) Observations sur la tribu des Cicindeletes. Revue Entomologique, 2, 27 - 38." type="journal article" year="1834">Laporte de Castelnau, 1834</bibRefCitation>
(spelled by Blackwelder as “
<taxonomicName id="79CA4D6CFFC12A1C9B34F973BE11F906" authorityName="Agassiz" authorityYear="1846" box="[895,1035,1737,1763]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9B34F973BE11F906" box="[895,1035,1737,1763]" italics="true" pageId="3" pageNumber="504">Odontochila</emphasis>
</taxonomicName>
”).
<bibRefCitation id="DA5B4B1EFFC12A1C9C7BF973BEACF906" author="Horn, W." box="[1072,1206,1737,1763]" pageId="3" pageNumber="504" pagination="103 - 208" refId="ref18907" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P. Genera Insectorum, 82 B, 103 - 208, pls. 6 - 15." type="book chapter" year="1910">Horn (1910</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9C88F972BEE4F906" author="Horn, W." box="[1219,1278,1736,1763]" pageId="3" pageNumber="504" pagination="209 - 487" refId="ref18947" refString="Horn, W. (1915) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, 82 C, 209 - 487, pls. 16 - 23." type="book chapter" year="1915">1915</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9D41F973BF4BF906" author="Horn, W." box="[1290,1361,1737,1763]" pageId="3" pageNumber="504" refId="ref19064" refString="Horn, W. (1926 a) Carabidae, Cicindelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, 345 pp." type="book" year="1926">1926a</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9D16F973BF82F906" author="Horn, W." box="[1373,1432,1737,1763]" pageId="3" pageNumber="504" pagination="124 - 125" refId="ref19173" refString="Horn, W. (1934) On the three forms of Odontochila subgenus Chilonycha (Cicind., Col). Extracto de la Revista Chilena de Historia Natural, 38, 124 - 125." type="journal article" year="1934">1934</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C98DCF957BACAF8E2" author="Horn, W." box="[151,208,1773,1799]" pageId="3" pageNumber="504" pagination="1 - 71" refId="ref19210" refString="Horn, W. (1938) 2000 Zeichnungen von Cicindelinae. Entomologische Beihefte aus Berlin-Dahlem, 5, 1 - 71, 90 Tafeln." type="book chapter" year="1938">1938</bibRefCitation>
) used the spelling “
<taxonomicName id="79CA4D6CFFC12A1C99FFF957B82FF8E2" box="[436,565,1773,1799]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC12A1C99FFF957B82FF8E2" box="[436,565,1773,1799]" italics="true" pageId="3" pageNumber="504">Chilonycha</emphasis>
</taxonomicName>
” and
<taxonomicName id="79CA4D6CFFC12A1C9A31F957B91CF8E2" authorityName="Agassiz" authorityYear="1846" box="[634,774,1773,1799]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9A31F957B91CF8E2" box="[634,774,1773,1799]" italics="true" pageId="3" pageNumber="504">Odontochila</emphasis>
</taxonomicName>
, both forms of spelling arising from the unjustified emendations by
<bibRefCitation id="DA5B4B1EFFC12A1C9959F8ABBBDAF8C9" author="Agassiz, L." box="[274,448,1809,1836]" pageId="3" pageNumber="504" refId="ref18141" refString="Agassiz, L. (1846) Nomenclatoris Zoologici, Index Universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium mnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Jent et Gassmann, Soloduri, viii + 393 pp." type="book" year="1846">Agassiz (1846)</bibRefCitation>
. Rivalier (1970) still maintained the spelling of the genus-group name as “
<taxonomicName id="79CA4D6CFFC12A1C9D44F8ABBF8AF8CE" box="[1295,1424,1809,1835]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Chilonycha">
<emphasis id="8CBEEAFDFFC12A1C9D44F8ABBF8AF8CE" box="[1295,1424,1809,1835]" italics="true" pageId="3" pageNumber="504">Chilonycha</emphasis>
</taxonomicName>
” (equally with “
<taxonomicName id="79CA4D6CFFC12A1C9909F88FBBD4F8AA" authorityName="Agassiz" authorityYear="1846" box="[322,462,1845,1871]" class="Insecta" family="Carabidae" genus="Odontochila" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C9909F88FBBD4F8AA" box="[322,462,1845,1871]" italics="true" pageId="3" pageNumber="504">Odontochila</emphasis>
</taxonomicName>
” by
<bibRefCitation id="DA5B4B1EFFC12A1C9A4CF88FB8BFF8B5" author="Rivalier, E." box="[519,677,1845,1872]" pageId="3" pageNumber="504" pagination="195 - 237" refId="ref22279" refString="Rivalier, E. (1969) Demembrement du genre Odontochila (Col. Cicindelidae) et revision des principales especes. Annales de la Societe entomologique de France, Nouvelle Serie, 5 (1), 195 - 237." type="journal article" year="1969">Rivalier 1969</bibRefCitation>
).
<bibRefCitation id="DA5B4B1EFFC12A1C9AF1F88FB963F8AA" author="Reichardt, H." box="[698,889,1845,1871]" pageId="3" pageNumber="504" pagination="346 - 493" refId="ref22249" refString="Reichardt, H. (1977) A synopsis of the genera of Neotropical Carabidae (Insecta: Coleoptera). Questiones Entomologicae, 13, 346 - 493." type="journal article" year="1977">Reichardt (1977)</bibRefCitation>
was probably the first author who restituted the original name spelling as
<taxonomicName id="79CA4D6CFFC12A1C99FFF8E3B858F896" authorityName=": Reichardt" authorityYear="1977" box="[436,578,1881,1907]" class="Insecta" family="Carabidae" genus="Cheilonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="8CBEEAFDFFC12A1C99FFF8E3B858F896" box="[436,578,1881,1907]" italics="true" pageId="3" pageNumber="504">Cheilonycha</emphasis>
</taxonomicName>
; moreover, he treated the genus in the subtribe Odontocheilina in accordance with the present revision. The restituted name spelling has been used by all subsequent authors such as
<bibRefCitation id="DA5B4B1EFFC12A1C9D0BF8C7BAFEF85E" author="Wiesner, J." pageId="3" pageNumber="504" refId="ref22668" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt. Checklist of the tiger beetles of the world (Coleoptera, Cicindelidae). Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner (1992)</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C98BBF81BB86DF85E" author="Pearson, D. L. &amp; Guerra, J. F. &amp; Brzoska, D. W." box="[240,631,1953,1979]" pageId="3" pageNumber="504" pagination="379 - 524" refId="ref22071" refString="Pearson, D. L. Guerra, J. F. &amp; Brzoska, D. W. (1999) The Tiger beetles of Bolivia: their Identification, Distribution and Natural History (Coleoptera: Cicindelidae). Contributions on Entomology, International, 3 (4), 379 - 524." type="journal article" year="1999">Pearson, Guerra &amp; Brzoska (1999)</bibRefCitation>
, Bosquet (2002),
<bibRefCitation id="DA5B4B1EFFC12A1C9B74F81BBE4EF85E" author="Erwin, T. L. &amp; Pearson, D. L." box="[831,1108,1953,1979]" pageId="3" pageNumber="504" refId="ref18698" refString="Erwin, T. L. &amp; Pearson, D. L. (2008) A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of the life. Vol. II. Carabidae-Nebriformes 2 - Cicindelitae. Pensoft Series Faunistica 84. Pensoft Publishers, Sofia, 366 pp., 33 pls." type="book" year="2008">Erwin &amp; Pearson (2008)</bibRefCitation>
,
<bibRefCitation id="DA5B4B1EFFC12A1C9C14F81BBF01F85E" author="Moravec, J." box="[1119,1307,1953,1979]" pageId="3" pageNumber="504" pagination="127 - 206" refId="ref20875" refString="Moravec, J. (2018 a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense- 20. Beckerium W. Horn stat. restit., Mesochila Rivalier stat. nov. and Poecilochila Rivalier stat. nov. (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 103 (2), 127 - 206" type="journal article" year="2018">Moravec (2018a</bibRefCitation>
, b).
</paragraph>
</subSubSection>
</treatment>
</document>