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<document id="FEAF5964C13EEA0C3A61CF6397619380" ID-DOI="10.11646/zootaxa.3999.2.3" ID-GBIF-Dataset="8f47b303-98a7-43ce-b5bb-d1b4bf02555b" ID-ISSN="1175-5326" ID-Zenodo-Dep="237045" ID-ZooBank="E4B0B754-1C94-4B61-B612-848804B059A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280765185" checkinUser="plazi" docAuthor="Mcmahan, Caleb D., Matamoros, Wilfredo A., Piller, Kyle R. &amp; Chakrabarty, Prosanta" docDate="2015" docId="36162773FFB1FFB1AB8EFC32FCE80290" docLanguage="en" docName="zt03999p234.pdf" docOrigin="Zootaxa 3999 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomocichla Regan 1908" docType="treatment" docVersion="9" lastPageNumber="230" masterDocId="CA2F5F0BFFA3FFA2AB19FF8EFF850318" masterDocTitle="Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera" masterLastPageNumber="234" masterPageNumber="211" pageNumber="229" updateTime="1698613983299" updateUser="plazi">
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<mods:title id="D662A9FB2578977C82B97D055AD3C6A2">Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera</mods:title>
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<treatment id="36162773FFB1FFB1AB8EFC32FCE80290" ID-DOI="http://doi.org/10.5281/zenodo.5672274" ID-GBIF-Taxon="119640588" ID-Zenodo-Dep="5672274" LSID="urn:lsid:plazi:treatment:36162773FFB1FFB1AB8EFC32FCE80290" httpUri="http://treatment.plazi.org/id/36162773FFB1FFB1AB8EFC32FCE80290" lastPageId="19" lastPageNumber="230" pageId="18" pageNumber="229">
<subSubSection id="F6A5C5EEFFB1FFB0AB8EFC32FD550726" pageId="18" pageNumber="229" type="nomenclature">
<paragraph id="BE009665FFB1FFB0AB8EFC32FD9100CE" blockId="18.[151,532,956,1016]" box="[151,532,956,982]" pageId="18" pageNumber="229">
<heading id="E5482109FFB1FFB0AB8EFC32FD9100CE" bold="true" box="[151,532,956,982]" fontSize="11" level="1" pageId="18" pageNumber="229" reason="1">
<emphasis id="8CCB4A77FFB1FFB0AB8EFC32FD9100CE" bold="true" box="[151,532,956,982]" pageId="18" pageNumber="229">
Genus
<taxonomicName id="79BFEDE6FFB1FFB0ABF7FC32FD9100CE" ID-CoL="7XM5" ID-ENA="63182" authority="Regan 1908" authorityName="Regan" authorityYear="1908" box="[238,532,956,982]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0ABF7FC32FEFE00CE" bold="true" box="[238,379,956,982]" italics="true" pageId="18" pageNumber="229">Tomocichla</emphasis>
<bibRefCitation id="DA2EEB94FFB1FFB0AA98FC32FD9100CE" author="Regan" box="[385,532,956,982]" pageId="18" pageNumber="229" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annals and Magazine of Natural History, Series 8, 2 (11), 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="BE009665FFB1FFB0AB8EFC50FF7800E0" blockId="18.[151,532,956,1016]" box="[151,253,990,1016]" pageId="18" pageNumber="229">
(
<figureCitation id="26848AE0FFB1FFB0AB86FC50FF7100E0" box="[159,244,990,1016]" captionStart="FIGURE 18" captionStartId="18.[151,250,1943,1965]" captionTargetBox="[159,1421,1468,1887]" captionTargetId="figure@18.[151,1435,1433,1921]" captionTargetPageId="18" captionText="FIGURE 18. Tomocichla tuba, FMNH 7734, 135 mm SL. Right side photographed as left side has been dissected away in this specimen." httpUri="https://zenodo.org/record/237065/files/figure.png" pageId="18" pageNumber="229">Fig. 18</figureCitation>
)
</paragraph>
<paragraph id="BE009665FFB1FFB0AB8EFBABFD550726" blockId="18.[151,1436,1061,1411]" box="[151,720,1061,1086]" pageId="18" pageNumber="229">
<emphasis id="8CCB4A77FFB1FFB0AB8EFBABFEE60726" bold="true" box="[151,355,1061,1086]" pageId="18" pageNumber="229">Inclusive species.</emphasis>
<taxonomicName id="79BFEDE6FFB1FFB0AA73FBA9FE320726" box="[362,439,1061,1086]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="species" species="tuba">
<emphasis id="8CCB4A77FFB1FFB0AA73FBA9FEF20726" box="[362,375,1063,1086]" italics="true" pageId="18" pageNumber="229">T</emphasis>
.
<emphasis id="8CCB4A77FFB1FFB0AA9CFBABFE320726" box="[389,439,1061,1086]" italics="true" pageId="18" pageNumber="229">tuba</emphasis>
</taxonomicName>
(
<typeStatus id="610428C7FFB1FFB0AADCFBA8FE720726" box="[453,503,1062,1086]" pageId="18" pageNumber="229">type</typeStatus>
species),
<taxonomicName id="79BFEDE6FFB1FFB0A97FFBA9FD550726" box="[614,720,1062,1086]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="species" species="asfraci">
<emphasis id="8CCB4A77FFB1FFB0A97FFBA9FDF60726" box="[614,627,1063,1086]" italics="true" pageId="18" pageNumber="229">T</emphasis>
.
<emphasis id="8CCB4A77FFB1FFB0A99BFBA8FD550726" box="[642,720,1062,1086]" italics="true" pageId="18" pageNumber="229">asfraci</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F6A5C5EEFFB1FFB0ABDEFBC4FECE0622" pageId="18" pageNumber="229" type="diagnosis">
<paragraph id="BE009665FFB1FFB0ABDEFBC4FECE0622" blockId="18.[151,1436,1061,1411]" pageId="18" pageNumber="229">
<emphasis id="8CCB4A77FFB1FFB0ABDEFBC4FEC4077B" bold="true" box="[199,321,1098,1123]" pageId="18" pageNumber="229">Diagnosis.</emphasis>
<taxonomicName id="79BFEDE6FFB1FFB0AA52FBC4FE4E077B" box="[331,459,1098,1123]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0AA52FBC4FE4E077B" box="[331,459,1098,1123]" italics="true" pageId="18" pageNumber="229">Tomocichla</emphasis>
</taxonomicName>
is diagnosed by the presence of conical but compressed teeth in the anterior of the upper jaw, and strongly pronounced rounded posterior edges of the dorsal and anal fins. These species possess an overall elongate body, but the body and head depth are higher than that of most members of
<taxonomicName id="79BFEDE6FFB1FFB0AF84FB1CFB7C07B3" box="[1181,1273,1170,1195]" class="Actinopterygii" family="Cichlidae" genus="Theraps" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0AF84FB1CFB7C07B3" box="[1181,1273,1170,1195]" italics="true" pageId="18" pageNumber="229">Theraps</emphasis>
</taxonomicName>
or
<taxonomicName id="79BFEDE6FFB1FFB0AE07FB1CFA1007B3" box="[1310,1429,1170,1195]" class="Actinopterygii" family="Cichlidae" genus="Rheoheros" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="8CCB4A77FFB1FFB0AE07FB1CFA1007B3" box="[1310,1429,1170,1195]" italics="true" pageId="18" pageNumber="229">Rheoheros</emphasis>
</taxonomicName>
. The number of anal-fin spines is four or five, and the mouth is subterminal with large lips. Pigmentation is largely dark in dorsal portions of the body and white, gray, or cream colored ventrally.
<taxonomicName id="79BFEDE6FFB1FFB0AF5AFB54FB4007EB" box="[1091,1221,1242,1267]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0AF5AFB54FB4007EB" box="[1091,1221,1242,1267]" italics="true" pageId="18" pageNumber="229">Tomocichla</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="79BFEDE6FFB1FFB0AB8EFB73FEC1060E" box="[151,324,1277,1302]" class="Actinopterygii" family="Cichlidae" genus="Paraneetroplus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0AB8EFB73FEC1060E" box="[151,324,1277,1302]" italics="true" pageId="18" pageNumber="229">Paraneetroplus</emphasis>
</taxonomicName>
but does not possess the anterior-facing (horizontally-positioned anterior) dentary teeth present in
<taxonomicName id="79BFEDE6FFB1FFB0AB8EFAACFEC10623" box="[151,324,1314,1339]" class="Actinopterygii" family="Cichlidae" genus="Paraneetroplus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB1FFB0AB8EFAACFEC10623" box="[151,324,1314,1339]" italics="true" pageId="18" pageNumber="229">Paraneetroplus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F6A5C5EEFFB1FFB0ABDEFACBFE2A069A" pageId="18" pageNumber="229" type="distribution">
<paragraph id="BE009665FFB1FFB0ABDEFACBFE2A069A" blockId="18.[151,1436,1061,1411]" pageId="18" pageNumber="229">
<emphasis id="8CCB4A77FFB1FFB0ABDEFACBFEE50646" bold="true" box="[199,352,1349,1374]" pageId="18" pageNumber="229">Distribution.</emphasis>
Atlantic slope of Middle
<collectingCountry id="C6A8D6F5FFB1FFB0A999FACBFD650646" box="[640,736,1349,1374]" name="United States of America" pageId="18" pageNumber="229">America</collectingCountry>
from the Río Escondido drainage in
<collectingCountry id="C6A8D6F5FFB1FFB0AF67FACBFB740646" box="[1150,1265,1349,1374]" name="Nicaragua" pageId="18" pageNumber="229">Nicaragua</collectingCountry>
to the rivers of Bocas del Toro,
<collectingCountry id="C6A8D6F5FFB1FFB0AA57FAE5FE2E069A" box="[334,427,1387,1410]" name="Panama" pageId="18" pageNumber="229">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="EAC0C6EDFFB1FFB0AB8EF819FF7B04D4" httpUri="https://zenodo.org/record/237065/files/figure.png" pageId="18" pageNumber="229" targetBox="[159,1421,1468,1887]" targetPageId="18">
<paragraph id="BE009665FFB1FFB0AB8EF819FF7B04D4" blockId="18.[151,1436,1943,1996]" pageId="18" pageNumber="229">
<emphasis id="8CCB4A77FFB1FFB0AB8EF819FE9C04B5" bold="true" box="[151,281,1943,1965]" pageId="18" pageNumber="229">FIGURE 18</emphasis>
.
<taxonomicName id="79BFEDE6FFB1FFB0AA3FF819FE4A04B5" box="[294,463,1943,1965]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="229" phylum="Chordata" rank="species" species="tuba">
<emphasis id="8CCB4A77FFB1FFB0AA3FF819FE4A04B5" box="[294,463,1943,1965]" italics="true" pageId="18" pageNumber="229">Tomocichla tuba</emphasis>
</taxonomicName>
, FMNH 7734, 135mm SL. Right side photographed as left side has been dissected away in this specimen.
</paragraph>
</caption>
<subSubSection id="F6A5C5EEFFB0FFB1ABDEFF19FCD30258" pageId="19" pageNumber="230" type="discussion">
<paragraph id="BE009665FFB0FFB1ABDEFF19FCD30258" blockId="19.[151,1437,151,392]" pageId="19" pageNumber="230">
<emphasis id="8CCB4A77FFB0FFB1ABDEFF19FECA03A8" bold="true" box="[199,335,151,176]" pageId="19" pageNumber="230">Comments.</emphasis>
<taxonomicName id="79BFEDE6FFB0FFB1AA4EFF19FE5C03A8" box="[343,473,151,176]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1AA4EFF19FE5C03A8" box="[343,473,151,176]" italics="true" pageId="19" pageNumber="230">Tomocichla</emphasis>
</taxonomicName>
was initially considered part of the Regans
<taxonomicName id="79BFEDE6FFB0FFB1A8CFFF19FBB703A8" box="[982,1074,151,176]" class="Actinopterygii" family="Cichlidae" genus="Theraps" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1A8CFFF19FBB703A8" box="[982,1074,151,176]" italics="true" pageId="19" pageNumber="230">Theraps</emphasis>
</taxonomicName>
section of
<taxonomicName id="79BFEDE6FFB0FFB1AFA8FF19FAB203A8" box="[1201,1335,151,176]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1AFA8FF19FAB203A8" box="[1201,1335,151,176]" italics="true" pageId="19" pageNumber="230">Cichlasoma</emphasis>
</taxonomicName>
. Species of
<taxonomicName id="79BFEDE6FFB0FFB1ABAEFF32FEBC03CD" box="[183,313,188,213]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1ABAEFF32FEBC03CD" box="[183,313,188,213]" italics="true" pageId="19" pageNumber="230">Tomocichla</emphasis>
</taxonomicName>
occur in fast-flowing rivers and is one of the few genera of herichthyin cichlids found outside of northern Middle
<collectingCountry id="C6A8D6F5FFB0FFB1AA42FF51FE3A03E0" box="[347,447,223,248]" name="United States of America" pageId="19" pageNumber="230">America</collectingCountry>
.
<bibRefCitation id="DA2EEB94FFB0FFB1AAD4FF6EFDEC03E0" author="Regan" box="[461,617,224,248]" pageId="19" pageNumber="230" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annals and Magazine of Natural History, Series 8, 2 (11), 455 - 464." type="journal article" year="1908">Regan (1908)</bibRefCitation>
initially proposed
<taxonomicName id="79BFEDE6FFB0FFB1A85CFF51FC4203E0" box="[837,967,223,248]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1A85CFF51FC4203E0" box="[837,967,223,248]" italics="true" pageId="19" pageNumber="230">Tomocichla</emphasis>
</taxonomicName>
based on the posterior placement of the pelvic fins.
<bibRefCitation id="DA2EEB94FFB0FFB1AA3BFE8AFE540205" author="Bussing" box="[290,465,260,285]" pageId="19" pageNumber="230" refString="Bussing, W. A. (1975) Taxonomy and biological aspects of the Central American cichlid fishes Cichlasoma sieboldii and C. tuba. Revista de Biologia Tropical, 23 (2), 189 - 211." type="journal article" year="1975">Bussing (1975)</bibRefCitation>
offers information on aspects of the taxonomy and biology of
<taxonomicName id="79BFEDE6FFB0FFB1AFA3FE8AFAB90205" box="[1210,1340,260,285]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="8CCB4A77FFB0FFB1AFA3FE8AFAB90205" box="[1210,1340,260,285]" italics="true" pageId="19" pageNumber="230">Tomocichla</emphasis>
</taxonomicName>
such as comparisons with similar species and morphological variation.
</paragraph>
</subSubSection>
<subSubSection id="F6A5C5EEFFB0FFB1ABDEFEC2FCE80290" pageId="19" pageNumber="230" type="materials_examined">
<paragraph id="BE009665FFB0FFB1ABDEFEC2FCE80290" blockId="19.[151,1437,151,392]" pageId="19" pageNumber="230">
<emphasis id="8CCB4A77FFB0FFB1ABDEFEC2FE37027D" bold="true" box="[199,434,332,357]" pageId="19" pageNumber="230">Material examined.</emphasis>
<taxonomicName id="79BFEDE6FFB0FFB1AAA3FEC3FD8C027D" box="[442,521,332,357]" class="Actinopterygii" family="Cichlidae" genus="Tomocichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="230" phylum="Chordata" rank="species" species="tuba">
<emphasis id="8CCB4A77FFB0FFB1AAA3FEC3FE42027C" box="[442,455,333,356]" italics="true" pageId="19" pageNumber="230">T</emphasis>
.
<emphasis id="8CCB4A77FFB0FFB1AACEFEC2FD8C027D" box="[471,521,332,357]" italics="true" pageId="19" pageNumber="230">tuba</emphasis>
</taxonomicName>
,
<collectionCode id="D8AE0EA0FFB0FFB1A900FEC3FDEF027C" LSID="urn:lsid:biocol.org:col:34795" box="[537,618,333,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="19" pageNumber="230">FMNH</collectionCode>
7734 [n=2,
<collectingCountry id="C6A8D6F5FFB0FFB1A9E1FEC2FCFD027D" box="[760,888,332,357]" name="Costa Rica" pageId="19" pageNumber="230">Costa Rica</collectingCountry>
: Río Matina],
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14684 [n=1,
<collectingCountry id="C6A8D6F5FFB0FFB1AE01FEC2FA1C027D" box="[1304,1433,332,357]" name="Costa Rica" pageId="19" pageNumber="230">Costa Rica</collectingCountry>
: Río Barranca],
<collectionCode id="D8AE0EA0FFB0FFB1AA5AFEFEFE270290" box="[323,418,368,392]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="19" pageNumber="230">LSUMZ</collectionCode>
14729 [n=1,
<collectingCountry id="C6A8D6F5FFB0FFB1A920FEFEFD320290" box="[569,695,367,392]" name="Costa Rica" pageId="19" pageNumber="230">Costa Rica</collectingCountry>
: Río Sarapiquí].
</paragraph>
</subSubSection>
</treatment>
</document>