415 lines
52 KiB
XML
415 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.67564" ID-GBIF-Dataset="57d9ade8-67e7-4d65-9581-b85a80e88893" ID-PMC="PMC8292283" ID-Pensoft-Pub="1313-2970-1048-145" ID-Pensoft-UUID="93FE96A4127A5D8FAA3A39BCB3E4B729" ID-PubMed="34316267" ID-ZooBank="578795FD41BF4807B0016F7B5AC4D4B9" ModsDocID="1313-2970-1048-145" checkinTime="1626283267302" checkinUser="pensoft" docAuthor="Lantsov, Vladimir I. & Pilipenko, Valentin E." docDate="2021" docId="35B73B80118E5247AD3A67AD3A54266C" docLanguage="en" docName="ZooKeys 1048: 145-175" docOrigin="ZooKeys 1048" docPubDate="2021-07-13" docSource="http://dx.doi.org/10.3897/zookeys.1048.67564" docTitle="Tipula (Lunatipula) eleniya Lantsov & Pilipenko 2021, sp. nov." docType="treatment" docUuid="2847C4F8-9727-4FC3-BECD-C552B75382C8" docUuidSource="ZooBank" docVersion="4" id="93FE96A4127A5D8FAA3A39BCB3E4B729" lastPageNumber="145" masterDocId="93FE96A4127A5D8FAA3A39BCB3E4B729" masterDocTitle="A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group" masterLastPageNumber="175" masterPageNumber="145" pageNumber="145" updateTime="1668150580932" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lantsov, Vladimir I.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8275-496X</mods:nameIdentifier>
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<mods:affiliation>Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">lantsov@megalog.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pilipenko, Valentin E.</mods:namePart>
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<mods:affiliation>Lomonosov Moscow State University, GSP- 1, Leninskie Gory, Moscow 119991, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1048</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>145</mods:start>
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<mods:end>175</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1048.67564</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.67564</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1048-145</mods:identifier>
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<mods:identifier type="ZooBank">578795FD41BF4807B0016F7B5AC4D4B9</mods:identifier>
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<mods:identifier type="Pensoft-UUID">93FE96A4127A5D8FAA3A39BCB3E4B729</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="183543015" LSID="urn:lsid:zoobank.org:act:2847C4F8-9727-4FC3-BECD-C552B75382C8" httpUri="http://treatment.plazi.org/id/35B73B80118E5247AD3A67AD3A54266C" lastPageNumber="145" pageId="0" pageNumber="145">
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<subSubSection pageId="0" pageNumber="145" type="nomenclature">
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<paragraph pageId="0" pageNumber="145">
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<taxonomicName LSID="http://zoobank.org/2847C4F8-9727-4FC3-BECD-C552B75382C8" authority="Lantsov & Pilipenko, 2021" authorityName="Lantsov & Pilipenko" authorityYear="2021" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) eleniya" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="eleniya" status="sp. nov." subGenus="Lunatipula">Tipula (Lunatipula) eleniya</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="145">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">, 5</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">, 13B</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">, 14B, G</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="materials_examined">
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<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<materialsCitation collectingDate="2008-06-14" collectingDateMax="2008-06-18" collectingDateMin="2008-06-14" collectorName="K. Tomkovich" country="Russia" county="Psekhako Mt." elevation="2000" latitude="43.691113" location="Krasnodarskiy Kray" longLatPrecision="669" longitude="40.366665" municipality="Sochi" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="145">
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<emphasis italics="true" pageId="0" pageNumber="145">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="Russia">Russia</collectingCountry>
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•
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<specimenCount type="male">1 male</specimenCount>
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;
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<location LSID="urn:lsid:plazi:treatment:35B73B80118E5247AD3A67AD3A54266C:A287E8935C212BB00F2EF2259839A17E" country="Russia" county="Psekhako Mt." latitude="43.691113" longLatPrecision="669" longitude="40.366665" municipality="Sochi" name="Krasnodarskiy Kray">Krasnodarskiy Kray</location>
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,
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<collectingMunicipality>Sochi</collectingMunicipality>
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env.
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<collectingCounty>Psekhako Mt.</collectingCounty>
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,
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<geoCoordinate degrees="43" direction="north" minutes="41" orientation="latitude" precision="15" seconds="28" value="43.691113">43°41'28"N</geoCoordinate>
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,
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<geoCoordinate degrees="40" direction="east" minutes="22" orientation="longitude" precision="925" value="40.366665">40°22'E</geoCoordinate>
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; alt. ~
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="m" value="2000.0">2000 m</elevation>
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</quantity>
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;
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<collectingDate value="2008-06-14" valueMax="2008-06-18" valueMin="2008-06-14">14-18 Jun. 2008</collectingDate>
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;
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<collectorName>K. Tomkovich</collectorName>
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leg.; ZISP
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</materialsCitation>
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.
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<typeStatus>Holotype</typeStatus>
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in good condition; however, left antenna, front right and left hind legs missing.
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="K. Tomkovich" country="Russia" county="Psekhako Mt." elevation="2000" latitude="43.691113" location="Russia" longLatPrecision="669" longitude="40.366665" municipality="Sochi" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry name="Russia">Russia</collectingCountry>
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•
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<specimenCount type="male">1 male</specimenCount>
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; same data as for holotype
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</materialsCitation>
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•
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<materialsCitation collectingDate="2018-07-26" collectorName="Pilipenko" country="Russia" elevation="1200" location="Kamyshanova Polyana" longitude="40.079445" municipality="Krasnodarskiy Kray" specimenCount="2" specimenCount-male="2" stateProvince="Krasnodar State" typeStatus="paratype">
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<specimenCount type="male">2 males</specimenCount>
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;
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<collectingMunicipality>Krasnodarskiy Kray</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:35B73B80118E5247AD3A67AD3A54266C:10D9D16A6140989365F091270B3E370B" country="Russia" longitude="40.079445" municipality="Krasnodarskiy Kray" name="Kamyshanova Polyana" stateProvince="Krasnodar State">Kamyshanova Polyana</location>
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env. [Biological station of
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<collectingRegion country="Russia" name="Krasnodar">Krasnodar State</collectingRegion>
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University], 44°16'91"N,
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<geoCoordinate degrees="40" direction="east" minutes="04" orientation="longitude" precision="15" seconds="46" value="40.079445">40°04'46"E</geoCoordinate>
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; alt.
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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;
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<collectingDate value="2018-07-26">26 Jul. 2018</collectingDate>
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; V..
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<collectorName>Pilipenko</collectorName>
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leg.; ZISP
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="diagnosis">
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<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="145">Male. General coloration grey with silver pruinescence. Tergite 9 with deep rounded notches either side of slightly grooved cone-shaped projection. Sternite 8 with a pair of small appendages, each bearing a long medially curved spine, broad base of appendages with fringe of long whitish hairs. Outer gonostylus small, triangular, slightly thickened distally, covered with long setae. Inner gonostylus with small rod-like outgrowth in middle of outer edge.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">General description.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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Adult (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3A</figureCitation>
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). Male body length 15.7 mm, wing length 16.2 mm, haltere 2.5 mm, nasus 0.13 mm, rostrum 0.8 mm, scape 0.6 mm, pedicel 0.15 mm, 1st flagellomere 0.5 mm. Length (mm) of leg segments, fore (1), mid (2), and hind (3); successively femur, tibia, 1st and 2nd tarsomeres: 1 (9.7; 10.4; 8.5; 3.8), 2 (9.8; 10.5; 9.0; 4.5) and 3 (10.0; 13.0; 9.0; 4.5). Length (mm) of 3rd, 4th, and 5th tarsomeres of all three pairs of legs, approximately the same: 1.0; 0.6; 0.5 mm, respectively.
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</paragraph>
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<caption doi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" pageId="0" pageNumber="145">Figure 3.</emphasis>
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<taxonomicName lsidName="T. (Lunatipula) eleniya" pageId="0" pageNumber="145" rank="species" species="eleniya" subGenus="Lunatipula">T. (Lunatipula) eleniya</taxonomicName>
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sp. nov., male habitus. Holotype
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<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
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general lateral view
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<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
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head and thorax, dorsal view. Scale bars: 10 mm (
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<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
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), 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="145" type="description">
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<paragraph pageId="0" pageNumber="145">Description.</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Head</emphasis>
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(Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3</figureCitation>
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13B</figureCitation>
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) grey with silvery pubescence. Rostrum grey dorsally, ventrally yellow-brown with sparse light rusty setae in black sockets. Nasus distinct, appearing triangular in dorsal aspect, covered with long, sparse procumbent setae. Frons and gula bare. Vertex (Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13B</figureCitation>
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) framed by rows of sparse setae alongside eye, glabrous in center with indistinct narrow longitudinal dark grey line. Genae, including lateral part of rostrum, with very sparse fine pale bristles. Tempora and postgenae with sparse black setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Antennae</emphasis>
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, bent backwards, reaches base of abdomen. Scape grey with silvery pruinescence, pedicel dirty yellow brown. Flagellomeres dark brown with verticils on slightly thickened bases, longest verticils subequal to length of respective flagellomere.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3A, B</figureCitation>
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) grey with silvery pruinescence, and short sparse whitish hairs. Pronotum dark brown with a weak dark stripe medially, and short sparse whitish bristles. Scutum (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3B</figureCitation>
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) with four brownish grey longitudinal stripes, lateral stripe shorter, medial stripes narrowly separated by dirty yellow area. Katepisternum, anepisternum, katepimeron, anepimeron and meron grey, bare, with silvery pruinescence; scutellum and mediotergite light grey with silvery pruinescence; scutal lobes with wide dark grey stripe, bordered by sparse whitish setae and with long yellow setae anterolaterally at wing base. Scutellum framed anteriorly with brown, with sparse yellow setae laterally, and with vague narrow dark grey medial line. Mediotergite light grey with sparse yellow setae and with silvery pruinescence. Laterotergite grey with silvery pruinescence and sparse erect short brown setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="145">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Wings</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3A</figureCitation>
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) with typical venation for the subgenus, transparent with brownish tin, (viewed in transmitted light). Stigma dirty yellow. Oblique lunule proximal to stigma. Setae on costal and subcostal veins; group of short black macrotrichia distally on
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<emphasis italics="true" pageId="0" pageNumber="145">R</emphasis>
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and base of
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<emphasis italics="true" pageId="0" pageNumber="145">R1</emphasis>
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. Distal section of
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<emphasis italics="true" pageId="0" pageNumber="145">Rs</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="145">R4 + 5</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="145">r-m</emphasis>
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, base of pentagonal discal cell lighter and weakened. Stem of cell
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<emphasis italics="true" pageId="0" pageNumber="145">m1</emphasis>
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approximately half length of cell. Apex of vein
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Cu</emphasis>
|
||
1 sharply curved at wing margin.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<taxonomicName genus="Halteres" lsidName="Halteres" pageId="0" pageNumber="145" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Halteres</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Stem grey with base covered with short white setae, knob dark grey.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Coxae light grey with silvery pruinescence; laterally with long sparse whitish yellow setae; medially with very short yellowish setae. Trochanters yellow with long yellow setae. Femora brown, narrowly yellow at base, covered with adpressed short brown setae; tibiae brownish; tarsal segments dark brown with short, adpressed brown setae. Tibial spur formula 1-2-2. Tarsal claws with a small tubercle at the base (magnification 10
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
4.5); with short but clearly visible arolium between ones.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Abdomen</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. T. (Lunatipula) eleniya sp. nov., male habitus. Holotype A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure3" httpUri="https://binary.pensoft.net/fig/566412" pageId="0" pageNumber="145">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4A</figureCitation>
|
||
). Tergites and sternites grey with silvery pruinescence, and sparse fine short golden setae. Tergites with lateral dark grey intermittent stripe, no distinct dorsal stripe. Tergites laterally and distally edged with whitish-yellow, proximal segments with thinner distal edge, becoming wider on caudal segments.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 4.</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) eleniya" pageId="0" pageNumber="145" rank="species" species="eleniya" subGenus="Lunatipula">T. (Lunatipula) eleniya</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
|
||
terminal segments of male abdomen, lateral view (dry)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B-L</emphasis>
|
||
male terminalia (after KOH 10% treatment)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
|
||
hypopygium (without sternite 8), caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
|
||
hypopygium, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
|
||
sternite 8, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
|
||
tergite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
|
||
tergite IX, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
|
||
inner and outer gonostylus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
|
||
inner and outer gonostylus, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
|
||
gonocoxal fragment, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
|
||
semen pump and aedeagus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis>
|
||
adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Terminalia</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14B, G</figureCitation>
|
||
). Hypopygium grey to dark grey, moderately thickened, with silvery pruinescence (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4A-C</figureCitation>
|
||
). Tergite 9 with two deep rounded notches flanking a median cone-shaped projection, the apex of which has a small groove (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4E, F</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5G-L</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14B</figureCitation>
|
||
). The dorsolateral edges of the tergite 9 convex, curved inward. Sternite 8 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4D</figureCitation>
|
||
) with a pair of small appendages bearing long medially curved spine, base of appendages with fringe of long whitish hairs not obscuring gap between. Appendages of sternite 9 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4K, L</figureCitation>
|
||
) abundantly pubescent. Outer gonostylus (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5A-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14G</figureCitation>
|
||
) small, triangular, slightly thickened distally, covered with long bristles. Inner gonostylus (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4G, H</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5A-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14G</figureCitation>
|
||
) with small rod-shaped outgrowth in middle of outer edge (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14G</figureCitation>
|
||
), twice as long as wide at base, slightly narrower medially, tip obliquely truncate. Posterior part of inner gonostylus a wide triangular plate with short sparse brown bristles in inner surface, long whitish setae in outer one. Gonocoxite (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5A-C</figureCitation>
|
||
) with two wide dentate outgrowths, ventral one longer, pointed, dorsal one rounded; long yellowish setae laterally on gonocoxite. Gonocoxal fragment (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4I</figureCitation>
|
||
) with short forked extension at base. Semen pump and aedeagus as in Figure
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4J</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Figure 5.</emphasis>
|
||
<taxonomicName lsidName="T. (Lunatipula) eleniya" pageId="0" pageNumber="145" rank="species" species="eleniya" subGenus="Lunatipula">T. (Lunatipula) eleniya</taxonomicName>
|
||
sp. nov., variation in the structure of the male terminalia
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A, D, G, J</emphasis>
|
||
variant I (holotype)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">B, E, H, K</emphasis>
|
||
variant II (paratype)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">C, F, I, L</emphasis>
|
||
variant III (paratype)
|
||
<emphasis bold="true" pageId="0" pageNumber="145">A-C</emphasis>
|
||
inner and outer gonostylus, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">D-F</emphasis>
|
||
inner and outer gonostylus, latero-caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">G-I</emphasis>
|
||
tergite 9, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="145">J-L</emphasis>
|
||
tergite 9, dorsal view. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<emphasis bold="true" pageId="0" pageNumber="145">Female</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="variation">
|
||
<paragraph pageId="0" pageNumber="145">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Three variants exist of the small rod-like outgrowth of the inner gonostylus: variant I (holotype - Sochi env., Psekhako Mt, alt. 2000 m) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5A, D</figureCitation>
|
||
), variant II (paratype) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5B, E</figureCitation>
|
||
) and variant III (paratype) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5C, F</figureCitation>
|
||
). (variants II and III from Kamyshanova polyna env., alt. 1200 m). There are also some differences in the shape of tergite 9 (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5G-L</figureCitation>
|
||
). Given the paucity of material, and the minor differences observed, the authors consider this to be intraspecific variability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="145">Comparisons to similar species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
The new species is readily separable from all other species of the subgenus by the presence of an outgrowth medially, on the outer edge of the inner gonostylus (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. T. (Lunatipula) eleniya sp. nov. A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium (without sternite 8), caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite IX, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure4" httpUri="https://binary.pensoft.net/fig/566413" pageId="0" pageNumber="145">4G</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. T. (Lunatipula) eleniya sp. nov., variation in the structure of the male terminalia A, D, G, J variant I (holotype) B, E, H, K variant II (paratype) C, F, I, L variant III (paratype) A-C inner and outer gonostylus, lateral view D-F inner and outer gonostylus, latero-caudal view G-I tergite 9, caudal view J-L tergite 9, dorsal view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure5" httpUri="https://binary.pensoft.net/fig/566414" pageId="0" pageNumber="145">5A-F</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="elevation">
|
||
<paragraph pageId="0" pageNumber="145">Elevation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults were collected at altitudes ranging from 1200-2000 m.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="flight period">
|
||
<paragraph pageId="0" pageNumber="145">Flight period.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Adults are active from middle of June through the end of July.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="habitat">
|
||
<paragraph pageId="0" pageNumber="145">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
Mixed moderately moist mesophytic plants, shady communities that include common hornbeam (
|
||
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
|
||
</taxonomicName>
|
||
), oriental beech (
|
||
<emphasis italics="true" pageId="0" pageNumber="145">
|
||
<normalizedToken originalValue="Fāgus">Fagus</normalizedToken>
|
||
<normalizedToken originalValue="orientālis">orientalis</normalizedToken>
|
||
</emphasis>
|
||
), Nordman fir (
|
||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Abies nordmanniana" order="Pinales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="nordmanniana">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Abies nordmanniana</emphasis>
|
||
</taxonomicName>
|
||
), ash vulgaris (
|
||
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
|
||
</taxonomicName>
|
||
), field maple (
|
||
<taxonomicName class="Dicotyledoneae" family="Aceraceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acer campestre" order="Sapindales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="campestre">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Acer campestre</emphasis>
|
||
</taxonomicName>
|
||
), colchis holly (
|
||
<taxonomicName authorityName="Poljark" authorityYear="1947" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex colchica" order="Aquifoliales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="colchica">
|
||
<emphasis italics="true" pageId="0" pageNumber="145">Ilex colchica</emphasis>
|
||
</taxonomicName>
|
||
), etc.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="distribution">
|
||
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">Endemic to the Caucasus: currently known from the West Caucasus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="145" type="etymology">
|
||
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="145">
|
||
<taxonomicName authorityName="Lantsov & Pilipenko" authorityYear="2021" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) eleniya" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="eleniya" subGenus="Lunatipula">Tipula (Lunatipula) eleniya</taxonomicName>
|
||
sp. nov. is named after the mother of the first author, Elena Nikolaevna Lantsova.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |