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<document id="E9082D7B4B6482EA7A102A86049A12A2" ID-CLB-Dataset="7783" ID-DOI="10.5852/ejt.2021.738.1261" ID-GBIF-Dataset="02cf5931-b18e-4d58-a6ff-9c7b1e37e68d" ID-ISSN="2118-9773" ID-Zenodo-Dep="4594541" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615410255512" checkinUser="plazi" docAuthor="Turner, Ian M. &amp; Cheek, Martin" docDate="2021" docId="355087D8FFE3E47E1877FBDCEA74C0FC" docLanguage="en" docName="ejt-738_1-60.pdf" docOrigin="European Journal of Taxonomy 738" docSource="http://dx.doi.org/10.5852/ejt.2021.738.1261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chassalia kinabaluensis T. Y. Yu 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="C969FFA0FFF9E4631A5DFFA8EF2BC42F" masterDocTitle="Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species" masterLastPageNumber="60" masterPageNumber="1" pageNumber="27" updateTime="1698932934732" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1E6CAFE210961B448C0BC92934E8A796" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="740FCEB8FC638E3F266D214A1845B92B">Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species</mods:title>
</mods:titleInfo>
<mods:name id="8C0A1FF4A3B9C40A59E9FC91AD67B89A" type="personal">
<mods:role id="A1621A8FDD6FEDCD49F6EF4B9CE73599">
<mods:roleTerm id="D6529BDA2D0DEE4ACF98DF1309D14ECB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D23972C72BB60E176B15C0865E1A32A3">Turner, Ian M.</mods:namePart>
<mods:nameIdentifier id="394DC704996B5EED54C85B8FAA1C9154" type="email">i.turner@kew.org</mods:nameIdentifier>
</mods:name>
<mods:name id="F1DAE15BFEB4C693FB2674C72DDCC452" type="personal">
<mods:role id="B309219122D80C67E0826A5F37DC659B">
<mods:roleTerm id="108A745F43E7D4DCB696FD2E8274652E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="959DAA2B8F4BCE81A8E8B0BE9DE54771">Cheek, Martin</mods:namePart>
<mods:affiliation id="894EA599D2A2745C027AE784C2A71B73">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Queen Mary University of London, E 1 4 NS, UK. Singapore Botanical Liaison Officer, Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569, Singapore.</mods:affiliation>
<mods:nameIdentifier id="A4E67720BE09B1B8BE6FA2A79CA9E42E" type="email">m.cheek@kew.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5CE321415A3063A91E9696E73D7BA457">text</mods:typeOfResource>
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<mods:titleInfo id="76E7FCE7675650BB30E64ADFDA8C2E4B">
<mods:title id="AF3ED227C0E85233A702A848FFD618AA">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D238AF78FE1BE4CE6F8755DACFF86B43">
<mods:date id="E059855CA3EE54408293170E23695000">2021</mods:date>
<mods:detail id="B1865B6E84ECE921EE252408EAE20400" type="pubDate">
<mods:number id="38A64FCABC89F89D0858ED74224DBA40">2021-03-09</mods:number>
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<mods:detail id="61193C1BFE173D64D3D3F3983784B563" type="volume">
<mods:number id="BF847C0E34EBD0849E8DE67DE1F9862E">738</mods:number>
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<mods:start id="41428E31392150D591E9A5D7D240069A">1</mods:start>
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<mods:url id="772B540B6CA2949964C98EF8C61F6FE5">http://dx.doi.org/10.5852/ejt.2021.738.1261</mods:url>
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<mods:classification id="860B2F0E1363432D027AAF358427804E">journal article</mods:classification>
<mods:identifier id="080E7FC8F52BEF99CBFA0CFB68BF9DB5" type="CLB-Dataset">7783</mods:identifier>
<mods:identifier id="C6E67EE5B874D2C44C0E2E6D8AA73B92" type="DOI">10.5852/ejt.2021.738.1261</mods:identifier>
<mods:identifier id="8EE5697BE9C7E58EE36B01BCAFBCAF44" type="GBIF-Dataset">02cf5931-b18e-4d58-a6ff-9c7b1e37e68d</mods:identifier>
<mods:identifier id="1119C31A8EDBA0B83303730DF0B4CE52" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="102E670C5317057880FD40B1B9D9608C" type="Zenodo-Dep">4594541</mods:identifier>
</mods:mods>
<treatment id="355087D8FFE3E47E1877FBDCEA74C0FC" ID-DOI="http://doi.org/10.5281/zenodo.4594527" ID-GBIF-Taxon="180327057" ID-Zenodo-Dep="4594527" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC" httpUri="http://treatment.plazi.org/id/355087D8FFE3E47E1877FBDCEA74C0FC" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27">
<subSubSection id="F5E36545FFE3E4791877FBDCEB21C0A0" box="[554,1034,1140,1167]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="BD4636CEFFE3E4791877FBDCEB21C0A0" blockId="26.[554,1034,1140,1240]" box="[554,1034,1140,1167]" pageId="26" pageNumber="27">
<heading id="E60E81A2FFE3E4791877FBDCEB21C0A0" box="[554,1034,1140,1167]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="7AF94D4DFFE3E4791877FBDCEC83C0A0" ID-CoL="9V56S" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[554,936,1140,1167]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="kinabaluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFE3E4791877FBDCEC7AC0A0" bold="true" box="[554,849,1140,1167]" italics="true" pageId="26" pageNumber="27">Chassalia kinabaluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFE3E47919F2FBDDEB21C0A0" box="[943,1034,1141,1167]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE3E479181AFB30EC77C0F7" pageId="26" pageNumber="27" type="description">
<paragraph id="BD4636CEFFE3E479181AFB30ECC6C09C" blockId="26.[554,1034,1140,1240]" box="[583,1005,1176,1203]" pageId="26" pageNumber="27">urn:lsid:ipni.org:names:77215407-1</paragraph>
<paragraph id="BD4636CEFFE3E4791885FB15EC77C0F7" blockId="26.[554,1034,1140,1240]" box="[728,860,1213,1240]" pageId="26" pageNumber="27">
<figureCitation id="25C22A4BFFE3E4791885FB15EC77C0F7" box="[728,860,1213,1240]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="27.[189,232,1925,1951]" captionStartId-1="28.[189,232,1999,2025]" captionTargetBox-0="[241,1315,403,1862]" captionTargetBox-1="[265,1322,265,1962]" captionTargetId-0="figure-11@27.[189,1398,285,1865]" captionTargetId-1="figure-14@28.[265,1322,265,1962]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="Fig. 14. Chassalia kinabaluensis T.Y.Yu sp. nov.A. Flowering shoot. B. Flower. C. Part of infructescence. D. Dry fruit. E. Pyrene ventral surface. From John H. Beaman 11299. Drawn by T.Y. Yu." captionText-1="Fig. 15. Holotype of Chassalia kinabaluensis T.Y.Yu sp. nov. (John H. Beaman 11299)." figureDoi-0="http://doi.org/10.5281/zenodo.4594576" figureDoi-1="http://doi.org/10.5281/zenodo.4594578" httpUri-0="https://zenodo.org/record/4594576/files/figure.png" httpUri-1="https://zenodo.org/record/4594578/files/figure.png" pageId="26" pageNumber="27">Figs 14–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE3E4791AE0FB57EA59C1BC" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="BD4636CEFFE3E4791AE0FB57EE1DC136" blockId="26.[189,310,1279,1305]" box="[189,310,1279,1305]" pageId="26" pageNumber="27">
<heading id="E60E81A2FFE3E4791AE0FB57EE1DC136" bold="true" box="[189,310,1279,1305]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="8F8DEADCFFE3E4791AE0FB57EE1DC136" bold="true" box="[189,310,1279,1305]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE3E4791AE0FA87EA59C1BC" blockId="26.[189,1398,1327,1427]" pageId="26" pageNumber="27">
Similar to
<taxonomicName id="7AF94D4DFFE3E4791B1EFA87ED4AC166" authority="Turner &amp; Cheek, 2021" authorityName="Turner &amp; Cheek" authorityYear="2021" box="[323,609,1327,1353]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="atropurpurea" status="sp. nov.">
<emphasis id="8F8DEADCFFE3E4791B1EFA87ED4AC166" box="[323,609,1327,1353]" italics="true" pageId="26" pageNumber="27">Chassalia atropurpurea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFE3E4791833FA98EDFBC165" box="[622,720,1328,1354]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, differs in having green-brown, not dark purple stems, pedicels, midribs and inflorescence axes; in having a shorter peduncle,
<quantity id="7A019B2BFFE3E47919A4FAFCEB63C141" box="[1017,1096,1364,1391]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="cm" value="3.5">3.5 cm</quantity>
long when flowering (not
<quantity id="7A019B2BFFE3E4791AE0FAD0EE01C1BC" box="[189,298,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="26" pageNumber="27" unit="cm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 cm</quantity>
long); differs also in having relatively long pedicels,
<quantity id="7A019B2BFFE3E47919C0FAD0ECD4C1BC" box="[925,1023,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="26" pageNumber="27" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long (not subsessile or sessile).
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE3E4791AE0FA13EB16C22A" box="[189,1085,1467,1541]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="BD4636CEFFE3E4791AE0FA13EE6FC1FA" blockId="26.[189,324,1467,1493]" box="[189,324,1467,1493]" pageId="26" pageNumber="27">
<heading id="E60E81A2FFE3E4791AE0FA13EE6FC1FA" bold="true" box="[189,324,1467,1493]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="8F8DEADCFFE3E4791AE0FA13EE6FC1FA" bold="true" box="[189,324,1467,1493]" pageId="26" pageNumber="27">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE3E4791AE0FA43EB16C22A" blockId="26.[189,1085,1515,1541]" box="[189,1085,1515,1541]" pageId="26" pageNumber="27">The epithet alludes to Mount Kinabalu to which the new species is endemic.</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE3E4791AE0F985EE90C3C6" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="BD4636CEFFE3E4791AE0F985EFD1C268" blockId="26.[189,250,1581,1607]" box="[189,250,1581,1607]" pageId="26" pageNumber="27">
<heading id="E60E81A2FFE3E4791AE0F985EFD1C268" bold="true" box="[189,250,1581,1607]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="8F8DEADCFFE3E4791AE0F985EFD1C268" bold="true" box="[189,250,1581,1607]" pageId="26" pageNumber="27">
<typeStatus id="6242886CFFE3E4791AE0F985EFD1C268" box="[189,250,1581,1607]" pageId="26" pageNumber="27">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE3E4791AE0F9F5EEA1C2EF" blockId="26.[189,1399,1629,1728]" pageId="26" pageNumber="27">
<materialsCitation id="0D913C93FFE3E4791AE0F9F5EEADC2EF" ID-GBIF-Occurrence="3050815334" collectingDate="1995-03-23" collectionCode="H, K, UNIMAS, IBEC" country="Malaysia" county="Mount Kinabalu" elevation="1850" latitude="6.0333333" location="Borneo" longLatPrecision="1305" longitude="116.55" municipality="Liawagu River Trail" pageId="26" pageNumber="27" specimenCode="K001129712" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="C5EE765EFFE3E4791AE0F9F5EE7FC258" box="[189,340,1629,1655]" name="Malaysia" pageId="26" pageNumber="27">MALAYSIA</collectingCountry>
–
<emphasis id="8F8DEADCFFE3E4791B25F9F5ED1FC258" bold="true" box="[376,564,1629,1655]" pageId="26" pageNumber="27">
<location id="B8266015FFE3E4791B25F9F5EEFEC258" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791B25F9F5EEFEC258" box="[376,469,1629,1655]" country="Malaysia" county="Mount Kinabalu" latitude="6.0333333" longLatPrecision="1305" longitude="116.55" municipality="Liawagu River Trail" name="Borneo" pageId="26" pageNumber="27" stateProvince="Sabah">Borneo</location>
,
<collectingRegion id="7F3DF82CFFE3E4791BBBF9F5ED1FC258" box="[486,564,1629,1655]" country="Malaysia" name="Sabah" pageId="26" pageNumber="27">Sabah</collectingRegion>
</emphasis>
•
<collectingCounty id="54274E42FFE3E4791809F9F6EC37C258" box="[596,796,1629,1656]" pageId="26" pageNumber="27">Mount Kinabalu</collectingCounty>
, head of
<collectingMunicipality id="5D22ACB4FFE3E47919CCF9F5EBA1C258" box="[913,1162,1629,1655]" pageId="26" pageNumber="27">Liawagu River Trail</collectingMunicipality>
; alt.
<quantity id="7A019B2BFFE3E4791E94F9F5EA5AC257" box="[1225,1393,1629,1656]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="26" pageNumber="27" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<elevation id="36D4D1FDFFE3E4791E94F9F5EA5AC257" box="[1225,1393,1629,1656]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="26" pageNumber="27" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800–1900 m</elevation>
</quantity>
;
<geoCoordinate id="D8CD5009FFE3E4791AE0F928EE0BC2B3" box="[189,288,1664,1692]" degrees="6" direction="north" minutes="02" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="6.0333333">6°02′ N</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFE3E4791B6CF928EE86C2B3" box="[305,429,1664,1692]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="116.55">116°33′ E</geoCoordinate>
;
<date id="C947100EFFE3E4791BE2F92AED41C2B3" box="[447,618,1665,1692]" pageId="26" pageNumber="27" value="1995-03-23">
<collectingDate id="D903E9E6FFE3E4791BE2F92AED41C2B3" box="[447,618,1665,1692]" pageId="26" pageNumber="27" value="1995-03-23">23 Mar. 1995</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E4791821F929ECB5C2B3" box="[636,926,1665,1692]" italics="true" pageId="26" pageNumber="27">
John
<collectionCode id="DBE8AE0BFFE3E479189CF92AEDF1C2B4" box="[705,730,1666,1691]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="26" pageNumber="27" type="Herbarium">H</collectionCode>
. Beaman 11299
</emphasis>
;
<typeStatus id="6242886CFFE3E47919EEF929EB35C2B4" box="[947,1054,1665,1691]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
:
<collectionCode id="DBE8AE0BFFE3E4791E72F929EB6EC2B3" box="[1071,1093,1665,1692]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
[
<specimenCode id="ED5F9EB5FFE3E4791E11F929EBD9C2B3" box="[1100,1266,1665,1692]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K001129712</specimenCode>
]!; isotypes:
<collectionCode id="DBE8AE0BFFE3E4791AE0F90EEE1FC2EF" box="[189,308,1702,1728]" country="Malaysia" httpUri="http://grbio.org/cool/z1cp-qx8g" name="Universiti Malaysia Sarawak" pageId="26" pageNumber="27">UNIMAS</collectionCode>
,
<collectionCode id="DBE8AE0BFFE3E4791B62F90EEEADC2EF" box="[319,390,1702,1728]" pageId="26" pageNumber="27">IBEC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="BD4636CEFFE3E4791AE0F940EE9FC32D" blockId="26.[189,436,1768,1794]" box="[189,436,1768,1794]" pageId="26" pageNumber="27">
<heading id="E60E81A2FFE3E4791AE0F940EE9FC32D" bold="true" box="[189,436,1768,1794]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="8F8DEADCFFE3E4791AE0F940EE9FC32D" bold="true" box="[189,436,1768,1794]" pageId="26" pageNumber="27">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE3E4791AE0F8B0EE90C3C6" blockId="26.[187,1399,1816,2025]" pageId="26" pageNumber="27">
<materialsCitation id="0D913C93FFE3E4791AE0F8B0EE1AC378" ID-GBIF-Occurrence="3050815339" collectingDate="1932-06" collectionCode="J, M, S, K" collectorName="Clemens" country="Malaysia" elevation="1500" location="Mount Kinabalu" municipality="Tenompok" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Sabah">
<collectingCountry id="C5EE765EFFE3E4791AE0F8B0EE7FC31D" box="[189,340,1816,1842]" name="Malaysia" pageId="26" pageNumber="27">MALAYSIA</collectingCountry>
–
<emphasis id="8F8DEADCFFE3E4791B31F8B0ED09C31D" bold="true" box="[364,546,1816,1842]" pageId="26" pageNumber="27">
Borneo,
<collectingRegion id="7F3DF82CFFE3E4791B89F8B0ED09C31D" box="[468,546,1816,1842]" country="Malaysia" name="Sabah" pageId="26" pageNumber="27">Sabah</collectingRegion>
</emphasis>
•
<collectingMunicipality id="5D22ACB4FFE3E4791865F8B0ED93C31D" box="[568,696,1816,1842]" pageId="26" pageNumber="27">Tenompok</collectingMunicipality>
,
<location id="B8266015FFE3E479189FF8B1ECAEC31D" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E479189FF8B1ECAEC31D" box="[706,901,1816,1843]" country="Malaysia" municipality="Tenompok" name="Mount Kinabalu" pageId="26" pageNumber="27" stateProvince="Sabah">Mount Kinabalu</location>
; alt.
<quantity id="7A019B2BFFE3E47919E7F8B0EB3FC31C" box="[954,1044,1816,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">
<elevation id="36D4D1FDFFE3E47919E7F8B0EB3FC31C" box="[954,1044,1816,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">1500 m</elevation>
</quantity>
;
<date id="C947100EFFE3E4791E42F8B1EBB8C31D" box="[1055,1171,1816,1843]" pageId="26" pageNumber="27" value="1932-06">
<collectingDate id="D903E9E6FFE3E4791E42F8B1EBB8C31D" box="[1055,1171,1816,1843]" pageId="26" pageNumber="27" value="1932-06">Jun. 1932</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E4791EC3F8B1EE2FC379" italics="true" pageId="26" pageNumber="27">
<collectionCode id="DBE8AE0BFFE3E4791EC3F8B1EB84C31D" box="[1182,1199,1817,1842]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="26" pageNumber="27" type="Herbarium">J</collectionCode>
. &amp;
<collectionCode id="DBE8AE0BFFE3E4791E8BF8B1EBDFC31C" box="[1238,1268,1817,1843]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="26" pageNumber="27" type="Herbarium">M</collectionCode>
.
<collectionCode id="DBE8AE0BFFE3E4791EAAF8B1EA2CC31C" box="[1271,1287,1817,1843]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="26" pageNumber="27" type="Herbarium">S</collectionCode>
.
<collectorName id="100C5318FFE3E4791F4CF8B0EA5DC31D" box="[1297,1398,1816,1842]" pageId="26" pageNumber="27">Clemens</collectorName>
28169
</emphasis>
;
<collectionCode id="DBE8AE0BFFE3E4791B4BF895EE1AC378" box="[278,305,1853,1879]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
</materialsCitation>
! •
<materialsCitation id="0D913C93FFE3E4791B12F895ECC5C378" ID-GBIF-Occurrence="3050815326" box="[335,1006,1852,1879]" collectingDate="1964-01-27" collectionCode="K" country="Malaysia" elevation="1500" location="Mesilau River" pageId="26" pageNumber="27" specimenCode="RSNB 4157" specimenCount="1" stateProvince="Borneo">
<location id="B8266015FFE3E4791B12F895EED0C378" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791B12F895EED0C378" box="[335,507,1853,1879]" country="Malaysia" name="Mesilau River" pageId="26" pageNumber="27" stateProvince="Borneo">Mesilau River</location>
; alt.
<quantity id="7A019B2BFFE3E479186EF894EDBBC378" box="[563,656,1852,1879]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">
<elevation id="36D4D1FDFFE3E479186EF894EDBBC378" box="[563,656,1852,1879]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="1500.0">1500 m</elevation>
</quantity>
;
<date id="C947100EFFE3E47918C1F895EC05C378" box="[668,814,1853,1879]" pageId="26" pageNumber="27" value="1964-01-27">
<collectingDate id="D903E9E6FFE3E47918C1F895EC05C378" box="[668,814,1853,1879]" pageId="26" pageNumber="27" value="1964-01-27">27 Jan.1964</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E4791966F895ECEFC378" box="[827,964,1853,1879]" italics="true" pageId="26" pageNumber="27">
<specimenCode id="ED5F9EB5FFE3E4791966F895ECEFC378" box="[827,964,1853,1879]" collectionCode="RSNB" pageId="26" pageNumber="27">RSNB 4157</specimenCode>
</emphasis>
;
<collectionCode id="DBE8AE0BFFE3E479198EF895ECC5C378" box="[979,1006,1853,1879]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
</materialsCitation>
! •
<materialsCitation id="0D913C93FFE3E4791E51F895EDEFC354" ID-GBIF-Occurrence="3050815378" collectingDate="2001-08-30" collectionCode="F, R, K" country="Malaysia" county="Ranau" location="Lower Montane Forest" municipality="Tenompok" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Borneo">
<collectingCounty id="54274E42FFE3E4791E51F895EB71C378" box="[1036,1114,1853,1879]" pageId="26" pageNumber="27">Ranau</collectingCounty>
,
<collectingMunicipality id="5D22ACB4FFE3E4791E3BF895EBC8C378" box="[1126,1251,1853,1879]" pageId="26" pageNumber="27">Tenompok</collectingMunicipality>
<collectionCode id="DBE8AE0BFFE3E4791EB6F895EBD5C378" box="[1259,1278,1853,1879]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="26" pageNumber="27" type="Herbarium">F</collectionCode>
.
<collectionCode id="DBE8AE0BFFE3E4791F5FF895EA32C378" box="[1282,1305,1853,1879]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="26" pageNumber="27">R</collectionCode>
.,
<location id="B8266015FFE3E4791F77F895EE51C353" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791F77F895EE51C353" country="Malaysia" county="Ranau" municipality="Tenompok" name="Lower Montane Forest" pageId="26" pageNumber="27" stateProvince="Borneo">Lower Montane Forest</location>
;
<date id="C947100EFFE3E4791BDBF8C9ED01C354" box="[390,554,1889,1915]" pageId="26" pageNumber="27" value="2001-08-30">
<collectingDate id="D903E9E6FFE3E4791BDBF8C9ED01C354" box="[390,554,1889,1915]" pageId="26" pageNumber="27" value="2001-08-30">30 Aug. 2001</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E479186BF8CAEDB2C354" box="[566,665,1889,1915]" italics="true" pageId="26" pageNumber="27">JTP 825</emphasis>
;
<collectionCode id="DBE8AE0BFFE3E47918F4F8C9EDEFC354" box="[681,708,1889,1915]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
</materialsCitation>
! •
<materialsCitation id="0D913C93FFE3E47918BFF8C9EEF2C3EB" ID-GBIF-Occurrence="3050815323" collectingDate="1983-12-20" collectionCode="H, K, MSC, UKMS" country="Malaysia" county="East Mesilau River" elevation="1800" latitude="6.05" location="Mesilau Cave" longLatPrecision="1305" longitude="-116.6" municipality="Mt Kinabalu" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Borneo">
along the
<collectingCounty id="54274E42FFE3E479190BF8CAEB13C354" box="[854,1080,1889,1916]" pageId="26" pageNumber="27">East Mesilau River</collectingCounty>
between
<collectingMunicipality id="5D22ACB4FFE3E4791EF7F8CAEA6BC354" box="[1194,1344,1889,1916]" pageId="26" pageNumber="27">Mt Kinabalu</collectingMunicipality>
golf course site and
<location id="B8266015FFE3E4791B28F82DED37C38F" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791B28F82DED37C38F" box="[373,540,1925,1952]" country="Malaysia" county="East Mesilau River" latitude="6.05" longLatPrecision="1305" longitude="-116.6" municipality="Mt Kinabalu" name="Mesilau Cave" pageId="26" pageNumber="27" stateProvince="Borneo">Mesilau Cave</location>
;
<geoCoordinate id="D8CD5009FFE3E4791874F82CEDACC38F" box="[553,647,1924,1952]" degrees="6" direction="north" minutes="03" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="6.05">6°03′ N</geoCoordinate>
,
<geoCoordinate id="D8CD5009FFE3E47918C9F82CEC3FC38F" box="[660,788,1924,1952]" degrees="116" direction="west" minutes="36" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="-116.6">116°36′ W</geoCoordinate>
; alt.
<quantity id="7A019B2BFFE3E4791913F82EECDFC38F" box="[846,1012,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="26" pageNumber="27" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">
<elevation id="36D4D1FDFFE3E4791913F82EECDFC38F" box="[846,1012,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="26" pageNumber="27" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700–1900 m</elevation>
</quantity>
;
<date id="C947100EFFE3E4791E5CF82EEB88C38F" box="[1025,1187,1926,1952]" pageId="26" pageNumber="27" value="1983-12-20">
<collectingDate id="D903E9E6FFE3E4791E5CF82EEB88C38F" box="[1025,1187,1926,1952]" pageId="26" pageNumber="27" value="1983-12-20">20 Dec. 1983</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E4791EEDF82DEFD3C3EB" italics="true" pageId="26" pageNumber="27">
<location id="B8266015FFE3E4791EEDF82DEBC2C3B0" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791EEDF82DEBC2C3B0" box="[1200,1257,1925,1951]" country="Malaysia" county="East Mesilau River" latitude="6.05" longLatPrecision="1305" longitude="-116.6" municipality="Mt Kinabalu" name="John" pageId="26" pageNumber="27" stateProvince="Borneo">John</location>
<collectionCode id="DBE8AE0BFFE3E4791EAFF82EEA20C3B0" box="[1266,1291,1926,1951]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="26" pageNumber="27" type="Herbarium">H</collectionCode>
.
<location id="B8266015FFE3E4791F4BF82EEA5DC3B0" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791F4BF82EEA5DC3B0" box="[1302,1398,1926,1951]" country="Malaysia" county="East Mesilau River" latitude="6.05" longLatPrecision="1305" longitude="-116.6" municipality="Mt Kinabalu" name="Beaman" pageId="26" pageNumber="27" stateProvince="Borneo">Beaman</location>
7977
</emphasis>
;
<collectionCode id="DBE8AE0BFFE3E4791B55F802EE0FC3EB" box="[264,292,1962,1988]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFE3E4791B6AF802EE53C3EB" box="[311,376,1962,1988]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15587" lsid="urn:lsid:biocol.org:col:15587" name="Michigan State University" pageId="26" pageNumber="27" type="Herbarium">MSC</collectionCode>
,
<collectionCode id="DBE8AE0BFFE3E4791BD9F802EEF2C3EB" box="[388,473,1962,1988]" collectionName="Sudan, Khartoum, University of Khartoum, Sudan Natural History Museum" country="Sudan" httpUri="http://grbio.org/cool/vy3j-vjek" name="Sudan Natural History Museum" pageId="26" pageNumber="27">UKMS</collectionCode>
</materialsCitation>
•
<materialsCitation id="0D913C93FFE3E4791BA9F802EE9CC3C6" ID-GBIF-Occurrence="3050815352" collectingDate="1966-05-27" collectionCode="K, L" country="Malaysia" elevation="1630" location="Ding Hou" municipality="Mt Kinabalu National Park" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Borneo">
<collectingMunicipality id="5D22ACB4FFE3E4791BA9F802EC12C3EB" box="[500,825,1962,1988]" pageId="26" pageNumber="27">Mt Kinabalu National Park</collectingMunicipality>
, montane oak forest; alt.
<quantity id="7A019B2BFFE3E4791E3BF802EBE8C3EB" box="[1126,1219,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="26" pageNumber="27" unit="m" value="1630.0">
<elevation id="36D4D1FDFFE3E4791E3BF802EBE8C3EB" box="[1126,1219,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="26" pageNumber="27" unit="m" value="1630.0">1630 m</elevation>
</quantity>
;
<date id="C947100EFFE3E4791E8DF802EA59C3EB" box="[1232,1394,1962,1988]" pageId="26" pageNumber="27" value="1966-05-27">
<collectingDate id="D903E9E6FFE3E4791E8DF802EA59C3EB" box="[1232,1394,1962,1988]" pageId="26" pageNumber="27" value="1966-05-27">27 May 1966</collectingDate>
</date>
;
<emphasis id="8F8DEADCFFE3E4791AE0F867EE4EC3C7" box="[189,357,1998,2024]" italics="true" pageId="26" pageNumber="27">
<location id="B8266015FFE3E4791AE0F867EE19C3C7" LSID="urn:lsid:plazi:treatment:355087D8FFE3E47E1877FBDCEA74C0FC:B8266015FFE3E4791AE0F867EE19C3C7" box="[189,306,1999,2024]" country="Malaysia" municipality="Mt Kinabalu National Park" name="Ding Hou" pageId="26" pageNumber="27" stateProvince="Borneo">Ding Hou</location>
227
</emphasis>
;
<collectionCode id="DBE8AE0BFFE3E4791B29F866EEBBC3C7" box="[372,400,1998,2024]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
!,
<collectionCode id="DBE8AE0BFFE3E4791BFFF867EE9CC3C6" box="[418,439,1999,2025]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="26" pageNumber="27" type="Herbarium">L</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="E9866646FFE2E4781AE0F82DEBFBC3EC" ID-DOI="http://doi.org/10.5281/zenodo.4594576" ID-Zenodo-Dep="4594576" httpUri="https://zenodo.org/record/4594576/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1925,1951]" targetBox="[241,1315,403,1862]" targetPageId="27">
<paragraph id="BD4636CEFFE2E4781AE0F82DEBFBC3EC" blockId="27.[186,1399,1925,1987]" pageId="27" pageNumber="28">
<emphasis id="8F8DEADCFFE2E4781AE0F82DEE3CC3B0" bold="true" box="[189,279,1925,1951]" pageId="27" pageNumber="28">Fig. 14.</emphasis>
<taxonomicName id="7AF94D4DFFE2E4781B46F82DEDA7C3B0" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[283,652,1925,1951]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="kinabaluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFE2E4781B46F82DED1DC3B0" box="[283,566,1925,1951]" italics="true" pageId="27" pageNumber="28">Chassalia kinabaluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFE2E47818CDF82DEDC3C3B0" box="[656,744,1925,1951]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="8F8DEADCFFE2E47818B7F82DEDD4C3B0" bold="true" box="[746,767,1925,1951]" pageId="27" pageNumber="28">A</emphasis>
. Flowering shoot.
<emphasis id="8F8DEADCFFE2E478198BF82DECC1C3B0" bold="true" box="[982,1002,1925,1951]" pageId="27" pageNumber="28">B</emphasis>
. Flower.
<emphasis id="8F8DEADCFFE2E4781E0EF82DEB43C3B0" bold="true" box="[1107,1128,1925,1951]" pageId="27" pageNumber="28">C</emphasis>
. Part of infructescence.
<emphasis id="8F8DEADCFFE2E4781AE7F800EFE4C3ED" bold="true" box="[186,207,1960,1986]" pageId="27" pageNumber="28">D</emphasis>
. Dry fruit.
<emphasis id="8F8DEADCFFE2E4781B0AF800EE40C3ED" bold="true" box="[343,363,1960,1986]" pageId="27" pageNumber="28">E</emphasis>
. Pyrene ventral surface. From
<emphasis id="8F8DEADCFFE2E4781888F800ECC3C3ED" box="[725,1000,1960,1986]" italics="true" pageId="27" pageNumber="28">John H. Beaman 11299</emphasis>
. Drawn by T.Y. Yu.
</paragraph>
</caption>
<caption id="E9866646FFE5E47F1AE0F867EBEAC3C5" ID-DOI="http://doi.org/10.5281/zenodo.4594578" ID-Zenodo-Dep="4594578" box="[189,1217,1999,2026]" httpUri="https://zenodo.org/record/4594578/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1999,2025]" targetBox="[265,1322,265,1962]" targetPageId="28">
<paragraph id="BD4636CEFFE5E47F1AE0F867EBEAC3C5" blockId="28.[189,1217,1999,2026]" box="[189,1217,1999,2026]" pageId="28" pageNumber="29">
<emphasis id="8F8DEADCFFE5E47F1AE0F867EE32C3C5" bold="true" box="[189,281,1999,2026]" pageId="28" pageNumber="29">Fig. 15.</emphasis>
Holotype of
<taxonomicName id="7AF94D4DFFE5E47F1BE8F878EC00C3C5" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[437,811,2000,2026]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="kinabaluensis" status="sp. nov.">
<emphasis id="8F8DEADCFFE5E47F1BE8F878EDF9C3C5" box="[437,722,2000,2026]" italics="true" pageId="28" pageNumber="29">Chassalia kinabaluensis</emphasis>
T.Y.Yu
</taxonomicName>
<taxonomicNameLabel id="94BE57A7FFE5E47F196FF878ECA7C3C5" box="[818,908,2000,2026]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="8F8DEADCFFE5E47F19C0F878EB9BC3C5" box="[925,1200,2000,2026]" italics="true" pageId="28" pageNumber="29">John H. Beaman 11299</emphasis>
).
</paragraph>
</caption>
<subSubSection id="F5E36545FFE4E47E1AE0FEA0EBDEC04D" pageId="29" pageNumber="30" type="description">
<paragraph id="BD4636CEFFE4E47E1AE0FEA0EE65C50D" blockId="29.[189,334,264,290]" box="[189,334,264,290]" pageId="29" pageNumber="30">
<heading id="E60E81A2FFE4E47E1AE0FEA0EE65C50D" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="8F8DEADCFFE4E47E1AE0FEA0EE65C50D" bold="true" box="[189,334,264,290]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE4E47E1AE0FE9FEBDEC04D" blockId="29.[188,1400,311,1122]" pageId="29" pageNumber="30">
Shrub to treelet,
<quantity id="7A019B2BFFE4E47E1BDEFE9FEED0C57D" box="[387,507,311,338]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="29" pageNumber="30" unit="m" value="0.7" valueMax="0.9" valueMin="0.5">0.5–0.9 m</quantity>
tall, glabrous, stems green-brown (also pedicels, midribs and inflorescence axes). Internodes from flowering branch (1–)2–5(–9.5) cm ×
<quantity id="7A019B2BFFE4E47E19C0FEF3EB2FC559" box="[925,1028,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
, lower internodes terete, upper internodes usually flattened, canaliculate, broadening at apex, base
<quantity id="7A019B2BFFE4E47E1987FED7EB17C5B6" box="[986,1084,383,409]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
wide, apex
<quantity id="7A019B2BFFE4E47E1E91FED7EA05C5B6" box="[1228,1326,383,410]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
wide. Leaf-blades narrowly elliptic to narrowly oblanceolate, 13–21.5 ×
<quantity id="7A019B2BFFE4E47E1996FE0AEB10C592" box="[971,1083,418,445]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="29" pageNumber="30" unit="cm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 cm</quantity>
, margin entire, apex acute, base cuneate; midrib slightly raised above, domatia absent; secondary nerves 8–12 on each side of the midrib, prominent above; tertiary nerves not visible. Petiole terete, sometimes canaliculate, usually slightly winged, 10–25 ×
<quantity id="7A019B2BFFE4E47E1BABFDA5ED76C607" box="[502,605,525,552]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
. Stipule not sheathing, broadly flabellate, 5–11 ×
<quantity id="7A019B2BFFE4E47E1EE6FDA5EA1AC607" box="[1211,1329,525,552]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="29" pageNumber="30" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, apex usually broken. Inflorescence a compound cyme,
<quantity id="7A019B2BFFE4E47E195FFD99EC43C663" box="[770,872,561,588]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="29" pageNumber="30" unit="cm" value="9.0" valueMax="12.0" valueMin="6.0">6–12 cm</quantity>
long; peduncle
<quantity id="7A019B2BFFE4E47E1E79FD99EBBAC663" box="[1060,1169,561,588]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="29" pageNumber="30" unit="cm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 cm</quantity>
long; bracts of first internode of rachis and first branches 2, triangular, flabellate or lanceolate, 3–12 ×
<quantity id="7A019B2BFFE4E47E1ED4FDFDEBC6C640" box="[1161,1261,597,623]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
, sometimes with 2–3 teeth on the margin; first internode of rachis and first branches
<quantity id="7A019B2BFFE4E47E1E14FDD0EBFAC6BC" box="[1097,1233,632,659]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.5" metricValueMin="4.5" pageId="29" pageNumber="30" unit="cm" value="6.5" valueMax="8.5" valueMin="4.5">4.5–8.5 cm</quantity>
long; second internode of rachis and second branches
<quantity id="7A019B2BFFE4E47E18F8FD34EC01C698" box="[677,810,668,695]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="29" pageNumber="30" unit="cm" value="3.5" valueMax="4.5" valueMin="2.5">2.5–4.5 cm</quantity>
long; bracts of capitulum oblong-obovate, 3–5 ×
<quantity id="7A019B2BFFE4E47E1AE2FD68EE2CC6F5" box="[191,263,704,730]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="2.0">2 mm</quantity>
, apex rounded. Pedicel short, 2–4 ×
<quantity id="7A019B2BFFE4E47E1898FD17EC0FC6F5" box="[709,804,703,730]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="mm" value="0.5">0.5 mm</quantity>
. Flowers subsessile, usually
<quantity id="7A019B2BFFE4E47E1EDDFD17EB9BC6F5" box="[1152,1200,703,730]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="30" unit="in" value="5.0">5 in</quantity>
each capitulum,
<quantity id="7A019B2BFFE4E47E1AE0FD4BEE10C6D1" box="[189,315,739,766]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long, hypanthium 1–3 ×
<quantity id="7A019B2BFFE4E47E1802FD4BEDC4C6D1" box="[607,751,739,766]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="29" pageNumber="30" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
, calyx tube 0.3–1 ×
<quantity id="7A019B2BFFE4E47E1985FD4BEB79C6D1" box="[984,1106,739,766]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 mm</quantity>
, lobes 0.3 ×
<quantity id="7A019B2BFFE4E47E1EBEFD4BEA58C6D1" box="[1251,1395,739,766]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="29" pageNumber="30" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
. Corolla cream white or purplish white, tube base narrow,
<quantity id="7A019B2BFFE4E47E193BFCAFECDEC70E" box="[870,1013,775,801]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="29" pageNumber="30" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.6–1.2 mm</quantity>
wide, broadening from midway, apex
<quantity id="7A019B2BFFE4E47E1AA2FC82EEA4C76A" box="[255,399,810,837]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="29" pageNumber="30" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
wide, corolla lobes triangular, 2.5 ×
<quantity id="7A019B2BFFE4E47E190FFC83ECB7C76A" box="[850,924,811,837]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="2.0">2 mm</quantity>
. Fruits iridescent blue, ellipsoid, 7–8 ×
<quantity id="7A019B2BFFE4E47E1AE0FCE6EE1DC746" box="[189,310,846,873]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="29" pageNumber="30" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">4–5.5 mm</quantity>
, with pedicels
<quantity id="7A019B2BFFE4E47E1BBFFCE6ED68C746" box="[482,579,846,873]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long. Pyrenes plano-convex, dorsal surface convex, with two adjacent parallel longitudinal crests, each
<quantity id="7A019B2BFFE4E47E1812FCDAED86C7A3" box="[591,685,882,908]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="mm" value="0.3">0.3 mm</quantity>
wide, curved,
<quantity id="7A019B2BFFE4E47E193BFCDAECEFC7A3" box="[870,964,882,908]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="mm" value="0.3">0.3 mm</quantity>
apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, central part slightly elevated, margin with an endocarpal outgrowth
<quantity id="7A019B2BFFE4E47E190EFC11EC85C7FB" box="[851,942,953,980]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="mm" value="0.6">0.6 mm</quantity>
wide, with a groove between the edge of endocarp and endocarpal outgrowth
<quantity id="7A019B2BFFE4E47E18C7FC75EC00C7D8" box="[666,811,989,1015]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
wide. Raphal opening at base of the endocarpal outgrowth, narrowly ovoid, 0.5 ×
<quantity id="7A019B2BFFE4E47E180FFBA9ED9AC034" box="[594,689,1025,1051]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="0.2">0.2 mm</quantity>
,
<quantity id="7A019B2BFFE4E47E18E0FBA9EC67C034" box="[701,844,1025,1051]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
from the edge of the pyrene wall, apex with a bony spine, spine triangular, erect, 0.5–0.7 ×
<quantity id="7A019B2BFFE4E47E1884FB8CEC40C010" box="[729,875,1060,1087]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
. PGS not detected. Seed light brown, hemiellipsoid, slightly crested at central part of dorsal side, 0.3 ×
<quantity id="7A019B2BFFE4E47E19D9FBE0ECE0C04D" box="[900,971,1096,1122]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="30" unit="mm" value="4.0">4 mm</quantity>
, endosperm not detected.
</paragraph>
</subSubSection>
<subSubSection id="F5E36545FFE4E47E1AE0FB21EA74C0FC" pageId="29" pageNumber="30" type="distribution">
<paragraph id="BD4636CEFFE4E47E1AE0FB21EE94C08C" blockId="29.[189,447,1161,1187]" box="[189,447,1161,1187]" pageId="29" pageNumber="30">
<heading id="E60E81A2FFE4E47E1AE0FB21EE94C08C" bold="true" box="[189,447,1161,1187]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="8F8DEADCFFE4E47E1AE0FB21EE94C08C" bold="true" box="[189,447,1161,1187]" pageId="29" pageNumber="30">Distribution, habitat</emphasis>
</heading>
</paragraph>
<paragraph id="BD4636CEFFE4E47E1AE0FB11EA74C0FC" blockId="29.[189,1375,1208,1236]" box="[189,1375,1208,1236]" pageId="29" pageNumber="30">
Borneo:
<collectingRegion id="7F3DF82CFFE4E47E1B7EFB11EE45C0FC" box="[291,366,1209,1235]" country="Malaysia" name="Sabah" pageId="29" pageNumber="30">Sabah</collectingRegion>
, Mount Kinabalu (Kinabalu National Park), Lower montane forest, alt.
<quantity id="7A019B2BFFE4E47E1EEAFB10EA70C0FC" box="[1207,1371,1208,1235]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="29" pageNumber="30" unit="m" value="1300.0" valueMax="1500.0" valueMin="1100.0">1100–1500 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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