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<document id="4BC4F2D8256568513EFF5CA6FD2A802A" ID-DOI="10.15298/rusentj.29.2.06" ID-ISSN="0132-8069" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722581546990" checkinUser="felipe" docAuthor="Sokolov, Alexander V. &amp; Perkovsky, Evgeny E." docDate="2020" docId="353D87BFFFB5FFF0FF29C13BFB18650C" docLanguage="en" docName="RussEntomolJ.29.2.157-160.pdf" docOrigin="Russian Entomological Journal 29 (2)" docSource="https://kmkjournals.com/journals/REJ/REJ_Index_Volumes/REJ_29/REJ_29_2_157_160" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Bacanius kirejtshuki Sokolov et Perkovsky 2020, sp.n." docType="treatment" docVersion="3" lastPageNumber="159" masterDocId="C904FFC7FFB7FFF2FFF1C33FFFBF610B" masterDocTitle="The first Eocene species of Bacanius (Coleoptera: Histeridae: Dendrophilinae) from Rovno amber" masterLastPageNumber="160" masterPageNumber="157" pageNumber="159" updateTime="1724436768022" updateUser="carolina">
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<mods:title id="BC58E35A9F1327AA3AA3998C9EBF6EA6">The first Eocene species of Bacanius (Coleoptera: Histeridae: Dendrophilinae) from Rovno amber</mods:title>
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<mods:namePart id="4EDE3710AF3C1C93F7267FCE3E3EA360">Sokolov, Alexander V.</mods:namePart>
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<subSubSection id="F58E6522FFB5FFF0FF29C13BFD546310" box="[216,747,515,539]" pageId="2" pageNumber="159" type="nomenclature">
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<taxonomicName id="7A944D2AFFB5FFF0FF29C13BFD156310" ID-CoL="9THLN" authority="Sokolov et Perkovsky" authorityName="Sokolov et Perkovsky" authorityYear="2020" box="[216,682,515,539]" class="Insecta" family="Histeridae" genus="Bacanius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="species" species="kirejtshuki" status="sp. nov.">
<emphasis id="8FE0EABBFFB5FFF0FF29C13BFE0B6310" box="[216,436,515,539]" italics="true" pageId="2" pageNumber="159">Bacanius kirejtshuki</emphasis>
Sokolov et Perkovsky
</taxonomicName>
,
<emphasis id="8FE0EABBFFB5FFF0FD44C13CFD546310" bold="true" box="[693,747,515,539]" pageId="2" pageNumber="159">
<taxonomicNameLabel id="94D357C0FFB5FFF0FD44C13CFD546310" box="[693,747,515,539]" pageId="2" pageNumber="159" rank="species">sp.n.</taxonomicNameLabel>
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<subSubSection id="F58E6522FFB5FFF0FE5EC11FFDAB6333" box="[431,532,544,568]" pageId="2" pageNumber="159" type="description">
<paragraph id="BD2B36A9FFB5FFF0FE5EC11FFDAB6333" blockId="2.[189,776,515,1187]" box="[431,532,544,568]" pageId="2" pageNumber="159">
<figureCitation id="25AF2A2CFFB5FFF0FE5EC11FFDAF6333" box="[431,528,544,568]" captionStart-0="Figs 13" captionStart-1="Figs 45" captionStartId-0="1.[226,263,2017,2036]" captionStartId-1="2.[226,263,2017,2036]" captionTargetBox-0="[188,1398,558,1980]" captionTargetBox-1="[189,1378,1219,1977]" captionTargetId-0="figure-165@1.[188,1398,558,1980]" captionTargetId-1="figure-502@2.[189,1378,1219,1968]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 13. Bacanius kirejtshuki sp.n.: 1, 3 — habitus, ventral and dorsal view; 2 — head, lateral view. Рис. 13. Bacanius kirejtshuki sp.n.: 1, 3 — габитус, сниЗу и сверху; 2 — голова, сбоку." captionText-1="Figs 45. Bacanius kirejtshuki sp.n., habitus: 4 — dorsal view; 5 — ventral view. Рис. 45. Bacanius kirejtshuki sp.n., габитус: 4 — сверху; 5 — сниЗу." figureDoi-0="http://doi.org/10.5281/zenodo.13165807" figureDoi-1="http://doi.org/10.5281/zenodo.13165809" httpUri-0="https://zenodo.org/record/13165807/files/figure.png" httpUri-1="https://zenodo.org/record/13165809/files/figure.png" pageId="2" pageNumber="159">Figs 15</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F58E6522FFB5FFF0FF13C17AFDF36353" box="[226,588,581,600]" pageId="2" pageNumber="159" type="materials_examined">
<paragraph id="BD2B36A9FFB5FFF0FF13C17AFDF36353" blockId="2.[189,776,515,1187]" box="[226,588,581,600]" pageId="2" pageNumber="159">
<materialsCitation id="0DFC3CF4FFB5FFF0FF13C17AFDF36353" box="[226,588,581,600]" collectionCode="SIZK" pageId="2" pageNumber="159" specimenCode="UA-28067" specimenCount="1" typeStatus="holotype">
MATERIAL.
<typeStatus id="622F880BFFB5FFF0FEACC17AFE126353" box="[349,429,581,600]" pageId="2" pageNumber="159" type="holotype">Holotype</typeStatus>
<collectionCode id="DB85AE6CFFB5FFF0FE42C17AFE5C6353" box="[435,483,581,600]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="2" pageNumber="159">SIZK</collectionCode>
<specimenCode id="ED329ED2FFB5FFF0FE18C17AFDF76353" box="[489,584,581,600]" collectionCode="UA-" pageId="2" pageNumber="159">UA-28067</specimenCode>
.
</materialsCitation>
</paragraph>
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<subSubSection id="F58E6522FFB5FFF0FF13C162FC2762DA" pageId="2" pageNumber="159" type="description">
<paragraph id="BD2B36A9FFB5FFF0FF13C162FDD16242" blockId="2.[189,776,515,1187]" pageId="2" pageNumber="159">
DESCRIPTION. Habitus as illustrated (
<figureCitation id="25AF2A2CFFB5FFF0FD90C162FD076378" box="[609,696,605,627]" captionStart-0="Figs 13" captionStart-1="Figs 45" captionStartId-0="1.[226,263,2017,2036]" captionStartId-1="2.[226,263,2017,2036]" captionTargetBox-0="[188,1398,558,1980]" captionTargetBox-1="[189,1378,1219,1977]" captionTargetId-0="figure-165@1.[188,1398,558,1980]" captionTargetId-1="figure-502@2.[189,1378,1219,1968]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 13. Bacanius kirejtshuki sp.n.: 1, 3 — habitus, ventral and dorsal view; 2 — head, lateral view. Рис. 13. Bacanius kirejtshuki sp.n.: 1, 3 — габитус, сниЗу и сверху; 2 — голова, сбоку." captionText-1="Figs 45. Bacanius kirejtshuki sp.n., habitus: 4 — dorsal view; 5 — ventral view. Рис. 45. Bacanius kirejtshuki sp.n., габитус: 4 — сверху; 5 — сниЗу." figureDoi-0="http://doi.org/10.5281/zenodo.13165807" figureDoi-1="http://doi.org/10.5281/zenodo.13165809" httpUri-0="https://zenodo.org/record/13165807/files/figure.png" httpUri-1="https://zenodo.org/record/13165809/files/figure.png" pageId="2" pageNumber="159">Figs 15</figureCitation>
). PEL =
<quantity id="7A6C9B4CFFB5FFF0FF4CC147FEB26386" box="[189,269,632,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="159" unit="mm" value="1.2">1.2 mm</quantity>
. H =
<quantity id="7A6C9B4CFFB5FFF0FEBDC147FE236386" box="[332,412,632,654]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="159" unit="mm" value="0.6">0.6 mm</quantity>
. Colour dark brown. Dorsal surface without microsculpture. Broadly oval, moderately convex. Widest at elytral humeri. Head retracted in thorax, relatively big, form typical for the tribe
<taxonomicName id="7A944D2AFFB5FFF0FE29C1F7FD8A63D5" authorityName="Kryzhanovskij" authorityYear="1976" box="[472,565,712,734]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Bacaniini">Bacaniini</taxonomicName>
(
<figureCitation id="25AF2A2CFFB5FFF0FDB1C1F7FDC063D6" box="[576,639,712,734]" captionStart="Figs 13" captionStartId="1.[226,263,2017,2036]" captionTargetBox="[188,1398,558,1980]" captionTargetId="figure-165@1.[188,1398,558,1980]" captionTargetPageId="1" captionText="Figs 13. Bacanius kirejtshuki sp.n.: 1, 3 — habitus, ventral and dorsal view; 2 — head, lateral view. Рис. 13. Bacanius kirejtshuki sp.n.: 1, 3 — габитус, сниЗу и сверху; 2 — голова, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13165807" httpUri="https://zenodo.org/record/13165807/files/figure.png" pageId="2" pageNumber="159">Figs 2</figureCitation>
,
<figureCitation id="25AF2A2CFFB5FFF0FD79C1F7FD2863D6" box="[648,663,712,733]" captionStart="Figs 45" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1378,1219,1977]" captionTargetId="figure-502@2.[189,1378,1219,1968]" captionTargetPageId="2" captionText="Figs 45. Bacanius kirejtshuki sp.n., habitus: 4 — dorsal view; 5 — ventral view. Рис. 45. Bacanius kirejtshuki sp.n., габитус: 4 — сверху; 5 — сниЗу." figureDoi="http://doi.org/10.5281/zenodo.13165809" httpUri="https://zenodo.org/record/13165809/files/figure.png" pageId="2" pageNumber="159">4</figureCitation>
). Frons and epistoma with uniform punctures coarse and sparse (23). Frons separate from epistoma by distinct stria, arcuate inwardly to vertex. First antennal segment poorly distinguishable, rest parts of antenna retracted in thorax.
</paragraph>
<paragraph id="BD2B36A9FFB5FFF0FF13C071FDB062E2" blockId="2.[189,776,515,1187]" pageId="2" pageNumber="159">
PL =
<quantity id="7A6C9B4CFFB5FFF0FEE9C072FECE6268" box="[280,369,845,867]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="159" unit="mm" value="0.38">0.38 mm</quantity>
. PPW =
<quantity id="7A6C9B4CFFB5FFF0FE37C072FDAD6268" box="[454,530,845,867]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="159" unit="mm" value="0.9">0.9 mm</quantity>
. APW =
<quantity id="7A6C9B4CFFB5FFF0FD9AC072FD076268" box="[619,696,845,867]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="159" unit="mm" value="0.8">0.8 mm</quantity>
. PPW =
<quantity id="7A6C9B4CFFB5FFF0FF4CC057FEB06276" box="[189,271,872,893]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="159" unit="mm" value="0.9">0.9 mm</quantity>
. Small part of pronotum visible only, impossible define features of punctures on the disc (
<figureCitation id="25AF2A2CFFB5FFF0FDA2C0BCFD2B6293" box="[595,660,899,921]" captionStart="Figs 13" captionStartId="1.[226,263,2017,2036]" captionTargetBox="[188,1398,558,1980]" captionTargetId="figure-165@1.[188,1398,558,1980]" captionTargetPageId="1" captionText="Figs 13. Bacanius kirejtshuki sp.n.: 1, 3 — habitus, ventral and dorsal view; 2 — head, lateral view. Рис. 13. Bacanius kirejtshuki sp.n.: 1, 3 — габитус, сниЗу и сверху; 2 — голова, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13165807" httpUri="https://zenodo.org/record/13165807/files/figure.png" pageId="2" pageNumber="159">Figs 2</figureCitation>
,
<figureCitation id="25AF2A2CFFB5FFF0FD51C0BCFD106293" box="[672,687,899,920]" captionStart="Figs 45" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1378,1219,1977]" captionTargetId="figure-502@2.[189,1378,1219,1968]" captionTargetPageId="2" captionText="Figs 45. Bacanius kirejtshuki sp.n., habitus: 4 — dorsal view; 5 — ventral view. Рис. 45. Bacanius kirejtshuki sp.n., габитус: 4 — сверху; 5 — сниЗу." figureDoi="http://doi.org/10.5281/zenodo.13165809" httpUri="https://zenodo.org/record/13165809/files/figure.png" pageId="2" pageNumber="159">4</figureCitation>
). Lateral surface with fine and sparse (24) punctures. Marginal pronotal stria complete laterally and anteriorly. Antescutellar stria absent. Scutellum not visible.
</paragraph>
<paragraph id="BD2B36A9FFB5FFF0FF13C0D1FC2C6300" blockId="2.[189,776,515,1187]" lastBlockId="2.[812,1399,315,1031]" pageId="2" pageNumber="159">
EW = 1.0 mm. Elytral punctures fine and sparse (24), distinguishable partly from middle to apex. Impossible to descern punctures in basal part of elytra and along suture. Outer subhumeral stria well impressed, complete, prolong to base. Epipleural stria distinct, situated closer to subhumeral striae. Prosternal lobe broad and prominent, with conspicuous sparse punctures (
<figureCitation id="25AF2A2CFFB5FFF0FE50C7B2FE5B65A9" box="[417,484,1165,1187]" captionStart="Figs 13" captionStartId="1.[226,263,2017,2036]" captionTargetBox="[188,1398,558,1980]" captionTargetId="figure-165@1.[188,1398,558,1980]" captionTargetPageId="1" captionText="Figs 13. Bacanius kirejtshuki sp.n.: 1, 3 — habitus, ventral and dorsal view; 2 — head, lateral view. Рис. 13. Bacanius kirejtshuki sp.n.: 1, 3 — габитус, сниЗу и сверху; 2 — голова, сбоку." figureDoi="http://doi.org/10.5281/zenodo.13165807" httpUri="https://zenodo.org/record/13165807/files/figure.png" pageId="2" pageNumber="159">Figs 1</figureCitation>
,
<figureCitation id="25AF2A2CFFB5FFF0FE04C7B2FDBC65A8" box="[501,515,1165,1187]" captionStart="Figs 45" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1378,1219,1977]" captionTargetId="figure-502@2.[189,1378,1219,1968]" captionTargetPageId="2" captionText="Figs 45. Bacanius kirejtshuki sp.n., habitus: 4 — dorsal view; 5 — ventral view. Рис. 45. Bacanius kirejtshuki sp.n., габитус: 4 — сверху; 5 — сниЗу." figureDoi="http://doi.org/10.5281/zenodo.13165809" httpUri="https://zenodo.org/record/13165809/files/figure.png" pageId="2" pageNumber="159">5</figureCitation>
). Mesometasternal suture looks like line of coarse punctures. Punctures of meso- and metasternum invisible mostly. Lateral parts of metasternum with various punctures, relatively dense (1.01.5). Hind edge of metaventrite arcuate to abdomen. There are outer and inner striae of 1st abdominal ventrite. Its surface coarsely punctured laterally and along anterior margin, rest part finely punctured. Probably pygidium smooth or with very weak punctures.
</paragraph>
<paragraph id="BD2B36A9FFB5FFF0FCA3C12EFB37637D" blockId="2.[812,1399,315,1031]" pageId="2" pageNumber="159">
Legs of typical form for
<taxonomicName id="7A944D2AFFB5FFF0FBB9C12EFB17632C" authorityName="Kryzhanovskij" authorityYear="1976" box="[1096,1192,529,551]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Bacaniini">Bacaniini</taxonomicName>
. Foretibia broadened with unique tooth near distal angle. Mesotibia and metatibia narrowed, not strongly broadened apically, mesotibia only with several fine spurs on the apex.
</paragraph>
<paragraph id="BD2B36A9FFB5FFF0FCA3C144FC2762DA" blockId="2.[812,1399,315,1031]" pageId="2" pageNumber="159">
COMPARATIVE REMARKS. The described species belongs to
<taxonomicName id="7A944D2AFFB5FFF0FC52C1A9FBBF63A7" authorityName="Kryzhanovskij" authorityYear="1976" box="[931,1024,662,684]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Bacaniini">Bacaniini</taxonomicName>
by not truncate elytra that covered exposed tergites, dorsal striae absent. Generic system of
<taxonomicName id="7A944D2AFFB5FFF0FCDDC1F3FC3963EA" authorityName="J. L. LeConte" authorityYear="1853" box="[812,902,716,737]" class="Insecta" family="Histeridae" genus="Bacanius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="8FE0EABBFFB5FFF0FCDDC1F3FC3963EA" box="[812,902,716,737]" italics="true" pageId="2" pageNumber="159">Bacanius</emphasis>
</taxonomicName>
is developed not so good now, many extant genera of
<taxonomicName id="7A944D2AFFB5FFF0FCB9C1D9FC1B63F7" authorityName="Kryzhanovskij" authorityYear="1976" box="[840,932,742,764]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="tribe" tribe="Bacaniini">Bacaniini</taxonomicName>
consist with species of former
<taxonomicName id="7A944D2AFFB5FFF0FB2EC1D8FA8663F7" authorityName="J. L. LeConte" authorityYear="1853" box="[1247,1337,743,764]" class="Insecta" family="Histeridae" genus="Bacanius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8FE0EABBFFB5FFF0FB2EC1D8FA8663F7" box="[1247,1337,743,764]" italics="true" pageId="2" pageNumber="159">Bacanius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94D357C0FFB5FFF0FAB0C1D9FCD3621C" pageId="2" pageNumber="159" sensu="stricto">sensu stricto</taxonomicNameLabel>
. Nonetheless
<taxonomicNameLabel id="94D357C0FFB5FFF0FC06C03EFBD0621C" box="[1015,1135,769,791]" pageId="2" pageNumber="159" rank="species">new species</taxonomicNameLabel>
has all key characters for attribute it to genus
<taxonomicName id="7A944D2AFFB5FFF0FBF4C023FBE0623A" authorityName="J. L. LeConte" authorityYear="1853" box="[1029,1119,796,817]" class="Insecta" family="Histeridae" genus="Bacanius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="8FE0EABBFFB5FFF0FBF4C023FBE0623A" box="[1029,1119,796,817]" italics="true" pageId="2" pageNumber="159">Bacanius</emphasis>
</taxonomicName>
. In 1984 Slawomir Mazur established subgenus
<taxonomicName id="7A944D2AFFB5FFF0FBFBC009FB406240" authority="Mazur, 1984" authorityName="Mazur" authorityYear="1984" box="[1034,1279,822,843]" class="Insecta" family="Histeridae" genus="Gomyister" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="8FE0EABBFFB5FFF0FBFBC009FBD16240" box="[1034,1134,822,843]" italics="true" pageId="2" pageNumber="159">Gomyister</emphasis>
<bibRefCitation id="D9054B58FFB5FFF0FB8AC009FB406240" author="Mazur S." box="[1147,1279,822,843]" pageId="2" pageNumber="159" pagination="1 - 376" refId="ref2760" refString="Mazur S. 1984. A world catalogue of Histeridae // Polskie Pismo Entomologiczne. Vol. 54. Nos 3 - 4. P. 1 - 376." type="book chapter" year="1984">Mazur, 1984</bibRefCitation>
</taxonomicName>
, distributed mostly in tropical areas of the Old and New World [
<bibRefCitation id="D9054B58FFB5FFF0FADEC06EFCDE628B" author="Mazur S." pageId="2" pageNumber="159" pagination="1 - 376" refId="ref2760" refString="Mazur S. 1984. A world catalogue of Histeridae // Polskie Pismo Entomologiczne. Vol. 54. Nos 3 - 4. P. 1 - 376." type="book chapter" year="1984">Mazur, 1984</bibRefCitation>
]. They can be recognized by complete subhumeral stria and absence of antescutellar stria.
<taxonomicName id="7A944D2AFFB5FFF0FB74C0B8FAF26290" authorityName="Sokolov et Perkovsky" authorityYear="2020" box="[1157,1357,902,924]" class="Insecta" family="Histeridae" genus="Bacanius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="species" species="kirejtshuki">
<emphasis id="8FE0EABBFFB5FFF0FB74C0B8FAF26290" box="[1157,1357,902,924]" italics="true" pageId="2" pageNumber="159">Bacanius kirejtshuki</emphasis>
</taxonomicName>
has that features, so we suppose to place it close to recent
<taxonomicName id="7A944D2AFFB5FFF0FCDDC083FC2E62DA" authorityName="Mazur" authorityYear="1984" box="[812,913,956,977]" class="Insecta" family="Histeridae" genus="Gomyister" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="8FE0EABBFFB5FFF0FCDDC083FC2E62DA" box="[812,913,956,977]" italics="true" pageId="2" pageNumber="159">Gomyister</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F58E6522FFB5FFF0FCA3C0E9FB18650C" pageId="2" pageNumber="159" type="etymology">
<paragraph id="BD2B36A9FFB5FFF0FCA3C0E9FB18650C" blockId="2.[812,1399,315,1031]" pageId="2" pageNumber="159">ETYMOLOGY. The new species is dedicated to coleopterist Dr. Sc. Alexander G. Kirejtshuk.</paragraph>
</subSubSection>
</treatment>
</document>