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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.270.4476" ID-GBIF-Dataset="95bc8b32-baed-4351-b41b-9ce20cb5d804" ID-PMC="PMC3668421" ID-Pensoft-Pub="1313-2970-270-59" ID-PubMed="23730188" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-270-59" ModsDocOrigin="ZooKeys 270" ModsDocTitle="New spider flies from the Neotropical Region (Diptera, Acroceridae) with a key to New World genera" checkinTime="1451247659983" checkinUser="pensoft" docAuthor="Schlinger, Evert I., Gillung, Jessica P. &amp; Borkent, Christopher J." docDate="2013" docId="34DCD9B665AFA9DDA425D18103E996A8" docLanguage="en" docName="ZooKeys 270: 59-93" docOrigin="ZooKeys 270" docSource="http://dx.doi.org/10.3897/zookeys.270.4476" docTitle="Sphaerops micella Schlinger, sp. n." docType="treatment" docVersion="4" lastPageNumber="66" masterDocId="FFFCFF98FFAAFFC0544CFFDBFF97FFEF" masterDocTitle="New spider flies from the Neotropical Region (Diptera, Acroceridae) with a key to New World genera" masterLastPageNumber="93" masterPageNumber="59" pageNumber="64" updateTime="1668155376915" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New spider flies from the Neotropical Region (Diptera, Acroceridae) with a key to New World genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schlinger, Evert I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gillung, Jessica P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Borkent, Christopher J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>270</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>59</mods:start>
<mods:end>93</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.270.4476</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.270.4476</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-270-59</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040826" LSID="urn:lsid:zoobank.org:act:32AB0AE4-47D5-42C9-9BCC-2AC1CB4B281A" httpUri="http://treatment.plazi.org/id/34DCD9B665AFA9DDA425D18103E996A8" lastPageId="7" lastPageNumber="66" pageId="5" pageNumber="64">
<subSubSection pageId="5" pageNumber="64" type="nomenclature">
<paragraph pageId="5" pageNumber="64">
<taxonomicName LSID="urn:lsid:zoobank.org:act:32AB0AE4-47D5-42C9-9BCC-2AC1CB4B281A" authority="Schlinger" class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="micella">
<pageBreakToken pageId="5" pageNumber="64" start="start">Sphaerops</pageBreakToken>
micella Schlinger
</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="64">sp. n.</taxonomicNameLabel>
Figs 1224
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="64" type="material examined">
<paragraph pageId="5" pageNumber="64">Material examined.</paragraph>
<paragraph pageId="5" pageNumber="64">
Holotype male: Top label: &quot;CHILE, Region III,
<normalizedToken originalValue="Copiapó">Copiapo</normalizedToken>
Prov / 125 km SE
<normalizedToken originalValue="Copiapó">Copiapo</normalizedToken>
; Fundo La / Semilla; malaise on alluvium nr river; / 30.x-9.xi.2003; ME Irwin; FD Parker / 2358 m;
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-28.250668">28°15.04'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-69.741">69°44.46'W</geoCoordinate>
&quot;. Bottom label: red &quot;HOLOTYPE ♂/
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="micella">Sphaerops micella</taxonomicName>
/ Schlinger new species/ Det. E.I. Schlinger 2012&quot; (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="64" type="paratypes">
<paragraph pageId="5" pageNumber="64">Paratypes.</paragraph>
<paragraph pageId="5" pageNumber="64">
All from Chile and bearing yellow paratype labels. Some with genitalia dissected and placed in glycerin in microvial on pin with specimen. Same data as holotype (4 ♂, CAS); Same except: 2 km E Puente La Semilla: 119 km SE
<normalizedToken originalValue="Copiapó">Copiapo</normalizedToken>
; malaise nr wash with water; 11-17.x.2003; ME Irwin; 2082 m;
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-28.215">28°12.90'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-69.761">69°45.66'W</geoCoordinate>
(1 ♂, CAS); Quillota Province; Palma de Ocoa; Parque Nacional Campanas; malaise in hillside draw; 215 m; 29.xii.1999; ME Irwin, EI Schlinger;
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-32.9324">32.9324°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-71.0781">71.0781°W</geoCoordinate>
(EI Schlinger #010913; 1 ♂, CAS); same except: 2-10.i.2000 (EI Schlinger #013443-013449; 7 ♂, CAS); Region VI,
<normalizedToken originalValue="Limarí">Limari</normalizedToken>
Prov. Frey Jorge
<normalizedToken originalValue="Natl">Nat'l</normalizedToken>
Park, Quebrada Honda I; malaise in small wash; 15-31.x.2003; ME Irwin, FD Parker; 122 m;
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-30.69">30°41.4'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.63">71°37.8'W</geoCoordinate>
(8 ♂); same except: 1-7.xi.2003 (ME Irwin #174749-174752, #174754-174756; 6 ♂, 1?, CAS); same except: El Mineral, malaise in wash upstream of seep, 23.
<normalizedToken originalValue="xi">xi-</normalizedToken>
12.xii.2003; ME Irwin, FD Parker; 224 m;
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-30.657333">30°39.44'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-71.665">71°39.90'W</geoCoordinate>
(2 ♂, CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="64" type="diagnosis">
<paragraph pageId="5" pageNumber="64">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="64">
This species is much smaller than
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops appendiculata" order="Diptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="appendiculata">Sphaerops appendiculata</taxonomicName>
(1.9 mm versus 6.3 mm) and has shorter pile on the thorax and abdomen. It is also unique in having the antennae placed immediately adjacent to the ocellar tubercle, the wing veins pale yellow, and wing veins CuA2 and A1 remaining separate.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="64" type="description">
<paragraph pageId="5" pageNumber="64">Description.</paragraph>
<paragraph pageId="5" pageNumber="64">
Male with small body length: 1.9
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6 mm (1.4-2.4 mm, n = 10) and wing just longer than body: 2.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5 mm (1.7 -2.5 mm, n = 10) setae covering body and legs is fine and short (Fig. 1). Female unknown. Head. Eye dark brown, occiput black, covered with pale yellow pile; ocellar tubercle black; ocelli white, frons dark brown, antennae light brown and placed on mediodorsal eye margin, face dark brown with pale yellow pile, clypeus light brown, bare and shorter than the antennae, mouthparts yellow and strongly reduced. Thorax. Uniformly dark brown with covering pale yellow setae; coxae light brown, femora light brown with apex yellow, tibia light brown with basal third yellow, tarsi light brown, basal and apical tarsomeres longer than middle tarsomeres, lower calypter translucent white and covered with pale yellow setae, halter pale yellow. Wing. (Fig. 2) All wing veins pale yellow except costa, subcosta and R1 pale brown; pterostigma darker. Abdomen. Background color brown tergites I-II entirely brown, tergite III with posteriomedial portion yellow, tergites IV-VI medially yellow, sternites brown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="66" pageId="5" pageNumber="64" type="comments">
<paragraph pageId="5" pageNumber="64">Comments.</paragraph>
<paragraph lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64">
Based on an examination of the voucher specimens of
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops" order="Diptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="genus">Sphaerops</taxonomicName>
sequenced by
<bibRefCitation author="Winterton, SL" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="24" pageNumber="83" pagination="808 - 832" title="Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers." url="10.1016/j.ympev.2006.08.015" volume="43" year="2007">Winterton et al. (2007)</bibRefCitation>
these were representatives of
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="micella">Sphaerops micella</taxonomicName>
sp. n. not
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops appendiculata" order="Diptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="appendiculata">
<pageBreakToken pageId="6" pageNumber="65" start="start">Sphaerops</pageBreakToken>
appendiculata
</taxonomicName>
as reported. Therefore the taxon associated with Genbank accession numbers AY140877, AF539875, AY144436, and AY144403 will be changed to
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="micella">Sphaerops micella</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="7" pageNumber="66">
<pageBreakToken pageId="7" pageNumber="66" start="start">Etymology</pageBreakToken>
. The species epithet is derived from the Latin: micella (diminutive feminine) meaning; little, crumb, or small, in reference to the minute size of this species relative to
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops appendiculata" order="Diptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="appendiculata">Sphaerops appendiculata</taxonomicName>
.
</paragraph>
<caption pageId="7" pageNumber="66">
<paragraph pageId="7" pageNumber="66">
Figure 1.
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="micella">Sphaerops micella</taxonomicName>
Schlinger sp. n., male paratype, lateral view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption pageId="7" pageNumber="66">
<paragraph pageId="7" pageNumber="66">
Figure 2.
<taxonomicName class="Insecta" family="Acroceridae" genus="Sphaerops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerops micella" order="Diptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="micella">Sphaerops micella</taxonomicName>
Schlinger sp. n., wing. Scale bar = 1.0 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>