treatments-xml/data/34/50/CF/3450CF1A1323FF8FE1E6FF41B28A62B9.xml

<document id="7BC4F14CD486846964B4056EBCEA2228" ID-DOI="10.11646/zootaxa.3981.2.6" ID-GBIF-Dataset="4ecba3c3-55a4-46cf-b276-0b0fa6c4f24c" ID-ISSN="1175-5326" ID-Zenodo-Dep="237013" ID-ZooBank="7C5C4334-8C6C-473C-9C6C-0B92BCF90883" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461246775608" checkinUser="plazi" docAuthor="Boonsoong, Boonsatien &amp; Sartori, Michel" docDate="2015" docId="3450CF1A1323FF8FE1E6FF41B28A62B9" docLanguage="en" docName="zt03981p263.pdf" docOrigin="Zootaxa 3981 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gilliesia ratchaburiensis Boonsoong &amp; Sartori, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="263" masterDocId="C869B7621322FF85E171FF9DB00B634D" masterDocTitle="The nymph of Gilliesia Peters &amp; Edmunds, 1970 (Ephemeroptera: Leptophlebiidae), with description of a new species from Thailand" masterLastPageNumber="263" masterPageNumber="253" pageNumber="254" updateTime="1698606609343" updateUser="plazi">
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<mods:title id="CAA814051A8698708375F6DDB965CDC3">The nymph of Gilliesia Peters &amp; Edmunds, 1970 (Ephemeroptera: Leptophlebiidae), with description of a new species from Thailand</mods:title>
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<mods:namePart id="1DC09EE933A88CBE3F70F06076FED2B4">Boonsoong, Boonsatien</mods:namePart>
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<mods:namePart id="4B7BFAAF2A8D179DCB7F2EDD626C7E9E">Sartori, Michel</mods:namePart>
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<mods:date id="20EDB0BC782E7BB46E403BCD3AB753B1">2015</mods:date>
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<treatment id="3450CF1A1323FF8FE1E6FF41B28A62B9" ID-DOI="http://doi.org/10.5281/zenodo.5622171" ID-GBIF-Taxon="119635290" ID-Zenodo-Dep="5622171" LSID="urn:lsid:plazi:treatment:3450CF1A1323FF8FE1E6FF41B28A62B9" httpUri="http://treatment.plazi.org/id/3450CF1A1323FF8FE1E6FF41B28A62B9" lastPageId="10" lastPageNumber="263" pageId="1" pageNumber="254">
<subSubSection id="F4E32D871323FF84E1E6FF41B1136255" pageId="1" pageNumber="254" type="nomenclature">
<paragraph id="BC467E0C1323FF84E1E6FF41B22E63BA" blockId="1.[151,549,220,280]" box="[151,549,220,247]" pageId="1" pageNumber="254">
<heading id="E70EC9601323FF84E1E6FF41B22E63BA" bold="true" box="[151,549,220,247]" fontSize="11" level="1" pageId="1" pageNumber="254" reason="1">
<emphasis id="8E8DA21E1323FF84E1E6FF41B22E63BA" bold="true" box="[151,549,220,247]" pageId="1" pageNumber="254">
<taxonomicName id="7BF9058F1323FF84E1E6FF41B1CB63BB" box="[151,448,220,246]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1323FF84E1E6FF41B1CB63BB" bold="true" box="[151,448,220,246]" italics="true" pageId="1" pageNumber="254">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651323FF84E0B6FF40B22E63BA" box="[455,549,221,247]" pageId="1" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BC467E0C1323FF84E1E6FF62B1136255" blockId="1.[151,549,220,280]" box="[151,280,255,280]" pageId="1" pageNumber="254">
(
<figureCitation id="24C262891323FF84E1EEFF62B11B6255" box="[159,272,255,280]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 5" captionStart-2="FIGURES 6 – 9" captionStart-3="FIGURES 10 – 13" captionStart-4="FIGURES 14 – 22" captionStart-5="FIGURES 23 – 29" captionStart-6="FIGURES 30 – 34" captionStart-7="FIGURES 35 – 38" captionStart-8="FIGURES 39 – 40" captionStartId-0="1.[151,250,1586,1608]" captionStartId-1="2.[151,264,1630,1653]" captionStartId-2="3.[151,264,1404,1427]" captionStartId-3="4.[151,264,1844,1867]" captionStartId-4="5.[151,264,1953,1976]" captionStartId-5="6.[151,264,1788,1811]" captionStartId-6="7.[151,264,1859,1882]" captionStartId-7="8.[151,264,897,920]" captionStartId-8="8.[151,264,1586,1609]" captionTargetBox-0="[565,1022,660,1559]" captionTargetBox-1="[168,1417,661,1599]" captionTargetBox-2="[199,1385,445,1382]" captionTargetBox-3="[190,1396,917,1823]" captionTargetBox-4="[227,1338,217,1909]" captionTargetBox-5="[179,1418,212,1746]" captionTargetBox-6="[151,1436,221,1837]" captionTargetBox-7="[340,1247,193,875]" captionTargetBox-8="[340,1246,1000,1565]" captionTargetId-0="figure@1.[560,1027,657,1564]" captionTargetId-1="figure@2.[168,1418,661,1599]" captionTargetId-2="figure@3.[199,1385,445,1383]" captionTargetId-3="figure@4.[190,1397,917,1824]" captionTargetId-4="figure@5.[223,1364,193,1932]" captionTargetId-5="figure@6.[151,1436,193,1767]" captionTargetId-6="figure@7.[151,1436,193,1838]" captionTargetId-7="figure@8.[340,1248,193,876]" captionTargetId-8="figure@8.[340,1247,1000,1566]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="7" captionTargetPageId-7="8" captionTargetPageId-8="8" captionText-0="FIGURE 1. Gilliesia ratchaburiensis sp. n. (1) male mature nymph, dorsal view." captionText-1="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." captionText-2="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." captionText-3="FIGURES 10 – 13. Nymphal details (SEM) of Gilliesia ratchaburiensis sp. n. (10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view." captionText-4="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." captionText-5="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" captionText-6="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." captionText-7="FIGURES 35 – 38. SEM pictures of eggs extracted from a female imago of Gilliesia ratchaburiensis sp. n. (35) general outline; (36 – 37) detail of pole; (38) detail of the chorionic structure." captionText-8="FIGURES 39 – 40. Inner-apical angle of maxilla of Gilliesia ratchaburiensis sp. n. (39) and Habrophlebiodes prominens (40), view in optical microscope." httpUri-0="https://zenodo.org/record/237014/files/figure.png" httpUri-1="https://zenodo.org/record/237015/files/figure.png" httpUri-2="https://zenodo.org/record/237016/files/figure.png" httpUri-3="https://zenodo.org/record/237017/files/figure.png" httpUri-4="https://zenodo.org/record/237018/files/figure.png" httpUri-5="https://zenodo.org/record/237019/files/figure.png" httpUri-6="https://zenodo.org/record/237020/files/figure.png" httpUri-7="https://zenodo.org/record/237021/files/figure.png" httpUri-8="https://zenodo.org/record/237022/files/figure.png" pageId="1" pageNumber="254">Figs 1–40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F4E32D871323FF84E1E6FEDAB33F6111" pageId="1" pageNumber="254" type="materials_examined">
<paragraph id="BC467E0C1323FF84E1E6FEDAB2E26159" blockId="1.[151,1437,327,605]" pageId="1" pageNumber="254">
<emphasis id="8E8DA21E1323FF84E1E6FEDAB181622D" bold="true" box="[151,394,327,352]" pageId="1" pageNumber="254">Material examined.</emphasis>
<typeStatus id="6342C0AE1323FF84E0EAFED5B238622D" box="[411,563,328,352]" pageId="1" pageNumber="254" type="holotype">HOLOTYPE</typeStatus>
: male imago (reared from nymph), 
<collectingCountry id="C4EE3E9C1323FF84E28CFED5B49A622D" box="[1021,1169,328,352]" name="Thailand" pageId="1" pageNumber="254">THAILAND</collectingCountry>
, Ratchaburi Province, tributary of Phachi River, 
<geoCoordinate id="D9CD18CB1323FF84E0B2FEF1B24E62C9" box="[451,581,364,389]" direction="north" orientation="latitude" pageId="1" pageNumber="254" precision="92" value="13.476666">13°28.6’ N</geoCoordinate>
, 
<geoCoordinate id="D9CD18CB1323FF84E323FEF1B2DB62C9" box="[594,720,364,389]" direction="east" orientation="longitude" pageId="1" pageNumber="254" precision="92" value="99.24167">99°14.5’ E</geoCoordinate>
, 
<quantity id="7B01D3E91323FF84E3AFFEF1B32262C9" box="[734,809,364,388]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="254" unit="m" value="230.0">230 m</quantity>
, 
<date id="C84758CC1323FF84E246FEF1B3B862C8" box="[823,947,364,389]" pageId="1" pageNumber="254" value="2013-03-30">30.iii.2013</date>
, B. Boonsoong leg.. ALLOTYPE: female imago (reared from nymph), same data as 
<typeStatus id="6342C0AE1323FF84E3E5FE12B2F162E5" box="[660,762,399,424]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
. 
<typeStatus id="6342C0AE1323FF84E278FE0DB3A362E5" box="[777,936,400,424]" pageId="1" pageNumber="254" type="paratype">PARATYPES</typeStatus>
: 4 mature nymphs, same data as 
<typeStatus id="6342C0AE1323FF84E44BFE12B59762E5" box="[1338,1436,399,424]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
(
<collectionCode id="DAE8E6C91323FF84E1EFFE29B0FF6281" box="[158,244,436,460]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="1" pageNumber="254">ZMKU</collectionCode>
); 4 nymphs mature nymphs, same data as 
<typeStatus id="6342C0AE1323FF84E3A8FE29B3306280" box="[729,827,436,461]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
(
<collectionCode id="DAE8E6C91323FF84E23BFE29B3806281" box="[842,907,436,460]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="1" pageNumber="254">MZL</collectionCode>
); 9 nymph, 
<date id="C84758CC1323FF84E561FE29B47D6280" box="[1040,1142,436,461]" pageId="1" pageNumber="254" value="2013-02-04">4.ii.2013</date>
, same locale as 
<typeStatus id="6342C0AE1323FF84E45AFE29B5866280" box="[1323,1421,436,461]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
; 
<specimenCount id="AAFFB5851323FF84E1E6FE45B0EA62BD" box="[151,225,471,496]" pageId="1" pageNumber="254" type="male">1 male</specimenCount>
imago (reared from nymph), 
<date id="C84758CC1323FF84E35DFE4AB2AD62BD" box="[556,678,471,496]" pageId="1" pageNumber="254" value="2014-11-15">15.xi.2014</date>
, same locale as 
<typeStatus id="6342C0AE1323FF84E229FE4AB3B162BD" box="[856,954,471,496]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
(all in 
<collectionCode id="DAE8E6C91323FF84E579FE45B46B62BD" box="[1032,1120,472,496]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="1" pageNumber="254">ZMKU</collectionCode>
); 
<specimenCount id="AAFFB5851323FF84E503FE45B4B662BD" box="[1138,1213,471,496]" pageId="1" pageNumber="254" type="male">1 male</specimenCount>
imago (reared from nymph), 
<date id="C84758CC1323FF84E18DFE61B17D6158" box="[252,374,508,533]" pageId="1" pageNumber="254" value="2014-11-15">15.xi.2014</date>
, same locale as 
<typeStatus id="6342C0AE1323FF84E35BFE61B2876158" box="[554,652,508,533]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
(
<collectionCode id="DAE8E6C91323FF84E3EAFE61B2D76159" box="[667,732,508,532]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="1" pageNumber="254">MZL</collectionCode>
).
</paragraph>
<paragraph id="BC467E0C1323FF84E1B6FD82B33F6111" blockId="1.[151,1437,327,605]" pageId="1" pageNumber="254">
<emphasis id="8E8DA21E1323FF84E1B6FD82B1B66175" bold="true" box="[199,445,543,568]" pageId="1" pageNumber="254">Additional material.</emphasis>
7 nymphs, 
<date id="C84758CC1323FF84E338FD82B2A46175" box="[585,687,543,568]" pageId="1" pageNumber="254" value="2013-02-02">2.ii.2013</date>
, F. David leg.; 65 nymphs, 
<date id="C84758CC1323FF84E289FD82B4676175" box="[1016,1132,543,568]" pageId="1" pageNumber="254" value="2013-02-16">16.ii.2013</date>
, F. David leg.; 6 nymphs, 
<date id="C84758CC1323FF84E1E6FDD9B1006110" box="[151,267,580,605]" pageId="1" pageNumber="254" value="2013-02-30">30.ii.2013</date>
, F. David leg.; same locale as 
<typeStatus id="6342C0AE1323FF84E32FFDD9B2CB6110" box="[606,704,580,605]" pageId="1" pageNumber="254" type="holotype">holotype</typeStatus>
(
<collectionCode id="DAE8E6C91323FF84E3BEFDD9B3236111" box="[719,808,580,604]" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="1" pageNumber="254">ZMKU</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="E8862E841323FF84E1E6F9AFB3DA6505" box="[151,977,1586,1608]" httpUri="https://zenodo.org/record/237014/files/figure.png" pageId="1" pageNumber="254" targetBox="[565,1022,660,1559]" targetPageId="1">
<paragraph id="BC467E0C1323FF84E1E6F9AFB3DA6505" blockId="1.[151,977,1586,1608]" box="[151,977,1586,1608]" pageId="1" pageNumber="254">
<emphasis id="8E8DA21E1323FF84E1E6F9AFB118650A" bold="true" box="[151,275,1586,1608]" pageId="1" pageNumber="254">FIGURE 1.</emphasis>
<taxonomicName id="7BF9058F1323FF84E068F9AFB21E6505" box="[281,533,1586,1608]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="254" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1323FF84E068F9AFB21E6505" box="[281,533,1586,1608]" italics="true" pageId="1" pageNumber="254">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651323FF84E36DF9AFB25E650A" box="[540,597,1586,1607]" pageId="1" pageNumber="254" rank="species">
<emphasis id="8E8DA21E1323FF84E36DF9AFB25E650A" bold="true" box="[540,597,1586,1607]" pageId="1" pageNumber="254">sp. n.</emphasis>
</taxonomicNameLabel>
(1) male mature nymph, dorsal view.
</paragraph>
</caption>
<subSubSection id="F4E32D871323FF86E1B6F9F2B3A76441" lastPageId="3" lastPageNumber="256" pageId="1" pageNumber="254" type="description">
<paragraph id="BC467E0C1323FF87E1B6F9F2B351612D" blockId="1.[151,1437,1647,2032]" lastBlockId="2.[151,1437,151,608]" lastPageId="2" lastPageNumber="255" pageId="1" pageNumber="254">
<emphasis id="8E8DA21E1323FF84E1B6F9F2B15265C5" bold="true" box="[199,345,1647,1672]" pageId="1" pageNumber="254">Description.</emphasis>
<emphasis id="8E8DA21E1323FF84E011F9EDB20F65C5" box="[352,516,1647,1672]" italics="true" pageId="1" pageNumber="254">Mature nymph</emphasis>
(
<figureCitation id="24C262891323FF84E365F9F2B25E65CA" box="[532,597,1647,1672]" captionStart="FIGURE 1" captionStartId="1.[151,250,1586,1608]" captionTargetBox="[565,1022,660,1559]" captionTargetId="figure@1.[560,1027,657,1564]" captionTargetPageId="1" captionText="FIGURE 1. Gilliesia ratchaburiensis sp. n. (1) male mature nymph, dorsal view." httpUri="https://zenodo.org/record/237014/files/figure.png" pageId="1" pageNumber="254">Fig. 1</figureCitation>
) (in alcohol). Body length between 3.0–4.0 mm for male (n=10) and 3.5– 5.0 mm for female (n=10). Head prognathous, antennae more than 2 times width of head (
<figureCitation id="24C262891323FF84E5F3F90EB4C865E6" box="[1154,1219,1683,1708]" captionStart="FIGURE 1" captionStartId="1.[151,250,1586,1608]" captionTargetBox="[565,1022,660,1559]" captionTargetId="figure@1.[560,1027,657,1564]" captionTargetPageId="1" captionText="FIGURE 1. Gilliesia ratchaburiensis sp. n. (1) male mature nymph, dorsal view." httpUri="https://zenodo.org/record/237014/files/figure.png" pageId="1" pageNumber="254">Fig. 1</figureCitation>
). Mouthparts (
<figureCitation id="24C262891323FF84E41BF90EB0DF659D" captionStart-0="FIGURES 2 – 5" captionStart-1="FIGURES 6 – 9" captionStart-2="FIGURES 10 – 13" captionStart-3="FIGURES 14 – 22" captionStartId-0="2.[151,264,1630,1653]" captionStartId-1="3.[151,264,1404,1427]" captionStartId-2="4.[151,264,1844,1867]" captionStartId-3="5.[151,264,1953,1976]" captionTargetBox-0="[168,1417,661,1599]" captionTargetBox-1="[199,1385,445,1382]" captionTargetBox-2="[190,1396,917,1823]" captionTargetBox-3="[227,1338,217,1909]" captionTargetId-0="figure@2.[168,1418,661,1599]" captionTargetId-1="figure@3.[199,1385,445,1383]" captionTargetId-2="figure@4.[190,1397,917,1824]" captionTargetId-3="figure@5.[223,1364,193,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." captionText-1="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." captionText-2="FIGURES 10 – 13. Nymphal details (SEM) of Gilliesia ratchaburiensis sp. n. (10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view." captionText-3="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri-0="https://zenodo.org/record/237015/files/figure.png" httpUri-1="https://zenodo.org/record/237016/files/figure.png" httpUri-2="https://zenodo.org/record/237017/files/figure.png" httpUri-3="https://zenodo.org/record/237018/files/figure.png" pageId="1" pageNumber="254">Figs 2–16</figureCitation>
): Labrum with pronounced anteromedian emargination and row of setae on anterior margin directed ventrally (
<figureCitation id="24C262891323FF84E1EEF946B10E65BE" box="[159,261,1755,1780]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1630,1653]" captionTargetBox="[168,1417,661,1599]" captionTargetId="figure@2.[168,1418,661,1599]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." httpUri="https://zenodo.org/record/237015/files/figure.png" pageId="1" pageNumber="254">Figs.2, 3</figureCitation>
, 
<figureCitation id="24C262891323FF84E064F946B13F65BE" box="[277,308,1755,1779]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="1" pageNumber="254">14</figureCitation>
). Dorsal surface of labrum with scattered setae and no transverse row of setae. Ventral surface of labrum with distal row of bristles (
<figureCitation id="24C262891323FF84E358F962B270645A" box="[553,635,1791,1816]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1630,1653]" captionTargetBox="[168,1417,661,1599]" captionTargetId="figure@2.[168,1418,661,1599]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." httpUri="https://zenodo.org/record/237015/files/figure.png" pageId="1" pageNumber="254">Figs. 4</figureCitation>
, 
<figureCitation id="24C262891323FF84E3F8F962B2A3645A" box="[649,680,1791,1815]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="1" pageNumber="254">14</figureCitation>
). Left and right mandibles as in 
<figureCitation id="24C262891323FF84E555F962B4AD6455" box="[1060,1190,1791,1816]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="1" pageNumber="254">Figs 15–16</figureCitation>
, with well-developed prostheca, without seta on outer margin. Superlinguae of hypopharynx rounded, not extended laterally, with rows of setae along anterior margin; apex of lingua cleft (
<figureCitation id="24C262891323FF84E395F8DAB322642D" box="[740,809,1863,1888]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1630,1653]" captionTargetBox="[168,1417,661,1599]" captionTargetId="figure@2.[168,1418,661,1599]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." httpUri="https://zenodo.org/record/237015/files/figure.png" pageId="1" pageNumber="254">Fig. 5</figureCitation>
); with two dense tufts of setae. Segment 1 of maxillary palpi subequal in length to segment 3. Length of segment 3 more than ca 1.6 times of length segment 2, apex of segment 3 with numerous long and thin setae, progressively longer from base to apex (
<figureCitation id="24C262891323FF84E50EF812B4E364E5" box="[1151,1256,1935,1960]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="1" pageNumber="254">Figs 6–7</figureCitation>
). Maxillae with crown of setae covering inner 3/4 of apex of galea (
<figureCitation id="24C262891323FF84E399F82EB3446481" box="[744,847,1971,1996]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="1" pageNumber="254">Figs 6–8</figureCitation>
); with 3 pectinate submarginal setae (ps, 
<figureCitation id="24C262891323FF84E458F82EB5846486" box="[1321,1423,1971,1996]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="1" pageNumber="254">Figs 8–9</figureCitation>
), apex of maxilla with three reduced canines (c1–c3, 
<figureCitation id="24C262891323FF84E3AFF84AB32964A2" box="[734,802,2007,2032]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="1" pageNumber="254">Fig. 9</figureCitation>
), proximal dentiseta (pd) and distal dentiseta (dd) entire (
<figureCitation id="24C262891320FF87E1EEFF0AB10C63FD" box="[159,263,151,176]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="2" pageNumber="255">Figs 8–9</figureCitation>
). Labium as in 
<figureCitation id="24C262891320FF87E0C8FF0AB23063FD" box="[441,571,151,176]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,264,1844,1867]" captionTargetBox="[190,1396,917,1823]" captionTargetId="figure@4.[190,1397,917,1824]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Nymphal details (SEM) of Gilliesia ratchaburiensis sp. n. (10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view." httpUri="https://zenodo.org/record/237017/files/figure.png" pageId="2" pageNumber="255">Figs 10–13</figureCitation>
; segment 2 of palpi slightly longer than 1/2 length of segment 1; segment 3 subequal in length to segment 2, triangular, with distinct stout bristles and thin setae (
<figureCitation id="24C262891320FF87E51DFF21B4C96399" box="[1132,1218,188,213]" captionStart="FIGURES 10 – 13" captionStartId="4.[151,264,1844,1867]" captionTargetBox="[190,1396,917,1823]" captionTargetId="figure@4.[190,1397,917,1824]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Nymphal details (SEM) of Gilliesia ratchaburiensis sp. n. (10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view." httpUri="https://zenodo.org/record/237017/files/figure.png" pageId="2" pageNumber="255">Fig. 12</figureCitation>
); glossae distinctly ventral to paraglossae; narrow at top and wider at base, densely covered with long setae on ventral surface (
<figureCitation id="24C262891320FF87E41EFF42B0BD6251" captionStart="FIGURES 10 – 13" captionStartId="4.[151,264,1844,1867]" captionTargetBox="[190,1396,917,1823]" captionTargetId="figure@4.[190,1397,917,1824]" captionTargetPageId="4" captionText="FIGURES 10 – 13. Nymphal details (SEM) of Gilliesia ratchaburiensis sp. n. (10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view." httpUri="https://zenodo.org/record/237017/files/figure.png" pageId="2" pageNumber="255">Fig. 13</figureCitation>
). Legs (
<figureCitation id="24C262891320FF87E07EFE99B1996251" box="[271,402,260,285]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Figs 17–19</figureCitation>
): all legs with stout bristles on outer margin of femur and inner margin of tibia; tibio-patellar suture present on mid- and hindlegs; foreleg (
<figureCitation id="24C262891320FF87E3EDFEBAB2E5620D" box="[668,750,295,320]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 17</figureCitation>
) with dark pigmentation on dorsal surface of femur, apex of tibia, and in middle of tarsi; midleg (
<figureCitation id="24C262891320FF87E346FED1B2826229" box="[567,649,332,357]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 18</figureCitation>
) with stout bristles on dorsal surface of femur; hindleg (
<figureCitation id="24C262891320FF87E479FED1B5516229" box="[1288,1370,332,357]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 19</figureCitation>
) with few short bristles on outer margin; tarsal claw (
<figureCitation id="24C262891320FF87E3D2FEF2B2FF62C5" box="[675,756,367,392]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 20</figureCitation>
) rather stout and not elongated, apex moderately hooked and narrow, with row of ca 10 teeth increasing in size towards apex. Abdominal terga with distinctive color pattern; terga 2–7 with the same pattern, with two median spots and two lateral spots; terga 2–9 with median longitudinal pale line; tergum 10 with two median spots (
<figureCitation id="24C262891320FF87E3EEFE41B2E862B9" box="[671,739,476,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1586,1608]" captionTargetBox="[565,1022,660,1559]" captionTargetId="figure@1.[560,1027,657,1564]" captionTargetPageId="1" captionText="FIGURE 1. Gilliesia ratchaburiensis sp. n. (1) male mature nymph, dorsal view." httpUri="https://zenodo.org/record/237014/files/figure.png" pageId="2" pageNumber="255">Fig. 1</figureCitation>
). Posterolateral expansions present on abdominal segment 9 only (
<figureCitation id="24C262891320FF87E1A8FE62B1256155" box="[217,302,511,536]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 21</figureCitation>
). Gills (
<figureCitation id="24C262891320FF87E0FDFE62B1EA6155" box="[396,481,511,536]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="2" pageNumber="255">Fig. 22</figureCitation>
): gills on segments 1–7 alike; long, slender and slightly forked at 2/5 distance from base; tracheae unbranched; no setae on gill margins. Caudal filaments with whorls of fine hair-like setae; terminal filament (
<quantity id="7B01D3E91320FF87E072FDD5B187612D" box="[259,396,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="2" pageNumber="255" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 mm</quantity>
) slightly longer than cerci (
<quantity id="7B01D3E91320FF87E3B0FDD5B345612D" box="[705,846,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="2" pageNumber="255" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.9–3.3 mm</quantity>
).
</paragraph>
<caption id="E8862E841320FF87E1E6F9C3B34465D9" httpUri="https://zenodo.org/record/237015/files/figure.png" pageId="2" pageNumber="255" targetBox="[168,1417,661,1599]" targetPageId="2">
<paragraph id="BC467E0C1320FF87E1E6F9C3B34465D9" blockId="2.[151,1436,1630,1684]" pageId="2" pageNumber="255">
<emphasis id="8E8DA21E1320FF87E1E6F9C3B1316539" bold="true" box="[151,314,1630,1653]" pageId="2" pageNumber="255">FIGURES 2–5.</emphasis>
Nymphal details of 
<taxonomicName id="7BF9058F1320FF87E37DF9C2B3016538" box="[524,778,1631,1653]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="255" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1320FF87E37DF9C2B3016538" box="[524,778,1631,1653]" italics="true" pageId="2" pageNumber="255">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
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<emphasis id="8E8DA21E1320FF87E261F9C2B3416539" bold="true" box="[784,842,1631,1652]" pageId="2" pageNumber="255">sp. n.</emphasis>
</taxonomicNameLabel>
(2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view.
</paragraph>
</caption>
<paragraph id="BC467E0C1320FF86E1B6F926B30E62C5" blockId="2.[151,1437,1723,2007]" lastBlockId="3.[151,1437,151,392]" lastPageId="3" lastPageNumber="256" pageId="2" pageNumber="255">
<emphasis id="8E8DA21E1320FF87E1B6F926B146659E" box="[199,333,1723,1748]" italics="true" pageId="2" pageNumber="255">Male imago</emphasis>
(
<typeStatus id="6342C0AE1320FF87E02DF926B1CB6599" box="[348,448,1723,1748]" pageId="2" pageNumber="255" type="holotype">holotype</typeStatus>
in alcohol). Body length 
<quantity id="7B01D3E91320FF87E3AEF921B34E659E" box="[735,837,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="4.24" pageId="2" pageNumber="255" unit="mm" value="4.24">4.24 mm</quantity>
, fore wing 
<quantity id="7B01D3E91320FF87E2B5F921B421659E" box="[964,1066,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="4.18" pageId="2" pageNumber="255" unit="mm" value="4.18">4.18 mm</quantity>
, hind wing 
<quantity id="7B01D3E91320FF87E5DFF921B51E659E" box="[1198,1301,1724,1748]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="255" unit="mm" value="0.4">0.40 mm</quantity>
. Compound eyes black; antennae pale, pedicel brown, ocelli white with dark base (
<figureCitation id="24C262891320FF87E2C6F942B40065BA" box="[951,1035,1759,1784]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="2" pageNumber="255">Fig. 23</figureCitation>
). Vertex of head and notum brown. Forefemora (
<figureCitation id="24C262891320FF87E05BF89EB1776451" box="[298,380,1795,1820]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="2" pageNumber="255">Fig. 26</figureCitation>
) brown, with two white spots basally and medially; tibiae pale, brown at base and apex; tarsi pale, each segment distally brownish; ratios of segments in fore legs, 0.50: 1.00 (
<quantity id="7B01D3E91320FF87E54FF8BAB4AD6472" box="[1086,1190,1831,1855]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="2" pageNumber="255" unit="mm" value="1.72">1.72 mm</quantity>
): 0.03: 0.37: 0.18: 0.15: 0.08. Mid- and hindlegs paler, with same markings as in foreleg. Midlegs (
<figureCitation id="24C262891320FF87E556F8D6B4776429" box="[1063,1148,1867,1892]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="2" pageNumber="255">Fig. 27</figureCitation>
): ratios of segments, 0.77: 1.00 (
<quantity id="7B01D3E91320FF87E196F8F2B14464CA" box="[231,335,1903,1927]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="255" unit="mm" value="0.89">0.89 mm</quantity>
): 0.13: 0.05: 0.05: 0.10. 
<collectingCountry id="C4EE3E9C1320FF87E3F9F8F2B2CB64C5" box="[648,704,1903,1928]" name="India" pageId="2" pageNumber="255">Hind</collectingCountry>
legs (
<figureCitation id="24C262891320FF87E274F8F2B35064CA" box="[773,859,1903,1928]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="2" pageNumber="255">Fig. 28</figureCitation>
): ratios of segments, 0.78: 1.00 (
<quantity id="7B01D3E91320FF87E5A4F8F2B53664CA" box="[1237,1341,1903,1927]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="255" unit="mm" value="0.93">0.93 mm</quantity>
): 0.11: 0.05: 0.05: 0.09. Forewing transparent, cross veins yellowish-brown; those in C-Sc, Sc-R1 and R1- Rs surrounded by dark brownish clouds forming markings as shown in 
<figureCitation id="24C262891320FF87E260F827B36F649F" box="[785,868,1978,2003]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="2" pageNumber="255">Fig. 24</figureCitation>
; MP2 independent of vein MP1; wing base brown; two long intercalaries between CuA and CuP (
<figureCitation id="24C262891321FF86E3D4FF0AB2F163FD" box="[677,762,151,176]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="3" pageNumber="256">Fig. 24</figureCitation>
). 
<collectingCountry id="C4EE3E9C1321FF86E27FFF0AB34D63FD" box="[782,838,151,176]" name="India" pageId="3" pageNumber="256">Hind</collectingCountry>
wing very small with distinct acute costal projection near apex (
<figureCitation id="24C262891321FF86E066FF21B1666398" box="[279,365,188,213]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="3" pageNumber="256">Fig. 25</figureCitation>
). Abdomen dark brown, terga 2–7 with one pair of median longitudinal white stripes arising from anterior margin; all terga with brown posterior margins; terga 8–9 entirely brown, tergum 10 with two pale spots (
<figureCitation id="24C262891321FF86E1AEFE99B1386251" box="[223,307,260,285]" captionStart="FIGURES 23 – 29" captionStartId="6.[151,264,1788,1811]" captionTargetBox="[179,1418,212,1746]" captionTargetId="figure@6.[151,1436,193,1767]" captionTargetPageId="6" captionText="FIGURES 23 – 29. Male imago of Gilliesia ratchaburiensis sp. n. (23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="3" pageNumber="256">Fig. 29</figureCitation>
). Caudal filaments missing. Genitalia (
<figureCitation id="24C262891321FF86E397FE99B3316251" box="[742,826,260,285]" captionStart="FIGURES 30 – 34" captionStartId="7.[151,264,1859,1882]" captionTargetBox="[151,1436,221,1837]" captionTargetId="figure@7.[151,1436,193,1838]" captionTargetPageId="7" captionText="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="3" pageNumber="256">Fig. 33</figureCitation>
): forceps length 
<quantity id="7B01D3E91321FF86E282FE99B4426251" box="[1011,1097,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="256" unit="mm" value="0.4">0.4 mm</quantity>
(ratios of segments 1.00 (
<quantity id="7B01D3E91321FF86E418FE99B0CF620D" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="256" unit="mm" value="0.32">0.32 mm</quantity>
): 0.15: 0.09), slightly curved, first segment long and curved, second broader than third segment. Styliger plate with deep, V-shaped, median incision. Penis anchor-like, apical portion of each lobe bent laterally but not ventrally, sharply pointed, no spine arising from tip of penis lobe.
</paragraph>
<caption id="E8862E841321FF86E1E6FAE1B2C4669D" httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="3" pageNumber="256" targetBox="[199,1385,445,1382]" targetPageId="3">
<paragraph id="BC467E0C1321FF86E1E6FAE1B2C4669D" blockId="3.[151,1436,1404,1488]" pageId="3" pageNumber="256">
<emphasis id="8E8DA21E1321FF86E1E6FAE1B13F66DC" bold="true" box="[151,308,1404,1427]" pageId="3" pageNumber="256">FIGURES 6–9</emphasis>
. Nymphal maxillae details of 
<taxonomicName id="7BF9058F1321FF86E31FFAE1B36766DF" box="[622,876,1404,1426]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1321FF86E31FFAE1B36766DF" box="[622,876,1404,1426]" italics="true" pageId="3" pageNumber="256">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651321FF86E205FAE0B3A566DF" box="[884,942,1405,1426]" pageId="3" pageNumber="256" rank="species">
<emphasis id="8E8DA21E1321FF86E205FAE0B3A566DF" bold="true" box="[884,942,1405,1426]" pageId="3" pageNumber="256">sp. n.</emphasis>
</taxonomicNameLabel>
(6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae.
</paragraph>
</caption>
<paragraph id="BC467E0C1321FF86E1B6FA6AB10B6589" blockId="3.[151,1436,1527,1912]" pageId="3" pageNumber="256">
<emphasis id="8E8DA21E1321FF86E1B6FA6AB1606542" box="[199,363,1527,1552]" italics="true" pageId="3" pageNumber="256">Female imago</emphasis>
(allotype in alcohol). Body length 
<quantity id="7B01D3E91321FF86E275FA6AB3666542" box="[772,877,1527,1551]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="256" unit="mm" value="4.7">4.70 mm</quantity>
, fore wing 
<quantity id="7B01D3E91321FF86E285FA6AB4566542" box="[1012,1117,1527,1552]" metricMagnitude="-3" metricUnit="m" metricValue="4.61" pageId="3" pageNumber="256" unit="mm" value="4.61">4.61 mm</quantity>
, hind wing 
<quantity id="7B01D3E91321FF86E599FA6AB5596542" box="[1256,1362,1527,1551]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="256" unit="mm" value="0.41">0.41 mm</quantity>
. Body color darker than in male. Vertex of head brown; compound eyes black; ocelli white with dark base (
<figureCitation id="24C262891321FF86E449F986B59B6579" box="[1336,1424,1563,1588]" captionStart="FIGURES 30 – 34" captionStartId="7.[151,264,1859,1882]" captionTargetBox="[151,1436,221,1837]" captionTargetId="figure@7.[151,1436,193,1838]" captionTargetPageId="7" captionText="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="3" pageNumber="256">Fig. 30</figureCitation>
). Pronotum pale, with dark brown pattern as in 
<figureCitation id="24C262891321FF86E3EEF9A2B2F8651A" box="[671,755,1599,1624]" captionStart="FIGURES 30 – 34" captionStartId="7.[151,264,1859,1882]" captionTargetBox="[151,1436,221,1837]" captionTargetId="figure@7.[151,1436,193,1838]" captionTargetPageId="7" captionText="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="3" pageNumber="256">Fig. 30</figureCitation>
. Meso- and metanotum yellowish-brown, lateral and ventral parts of abdomen pale. All legs missing. Abdominal tergal patterns similar to that of male. Color pattern of wing similar to that of male but crossveins darker (
<figureCitation id="24C262891321FF86E3D3F91AB2FC65D2" box="[674,759,1671,1696]" captionStart="FIGURES 30 – 34" captionStartId="7.[151,264,1859,1882]" captionTargetBox="[151,1436,221,1837]" captionTargetId="figure@7.[151,1436,193,1838]" captionTargetPageId="7" captionText="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="3" pageNumber="256">Fig. 31</figureCitation>
). Apex of sternum 9 with U-shaped, deep, median incision (
<figureCitation id="24C262891321FF86E1EEF936B0FF6589" box="[159,244,1707,1732]" captionStart="FIGURES 30 – 34" captionStartId="7.[151,264,1859,1882]" captionTargetBox="[151,1436,221,1837]" captionTargetId="figure@7.[151,1436,193,1838]" captionTargetPageId="7" captionText="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="3" pageNumber="256">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="BC467E0C1321FF86E1B6F94DB3A76441" blockId="3.[151,1436,1527,1912]" pageId="3" pageNumber="256">
<emphasis id="8E8DA21E1321FF86E1B6F94DB0FF65AA" box="[199,244,1744,1767]" italics="true" pageId="3" pageNumber="256">Egg</emphasis>
. General shape ovoid; chorionic surface with small uniform tubercles subequal in size, each turbercle with indistinct ridges and shallow groove between turbercles (
<figureCitation id="24C262891321FF86E267F96EB3946441" box="[790,927,1779,1804]" captionStart="FIGURES 35 – 38" captionStartId="8.[151,264,897,920]" captionTargetBox="[340,1247,193,875]" captionTargetId="figure@8.[340,1248,193,876]" captionTargetPageId="8" captionText="FIGURES 35 – 38. SEM pictures of eggs extracted from a female imago of Gilliesia ratchaburiensis sp. n. (35) general outline; (36 – 37) detail of pole; (38) detail of the chorionic structure." httpUri="https://zenodo.org/record/237021/files/figure.png" pageId="3" pageNumber="256">Figs. 35–38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F4E32D871321FF8DE1B6F88AB4F4647A" lastPageId="8" lastPageNumber="261" pageId="3" pageNumber="256" type="diagnosis">
<paragraph id="BC467E0C1321FF86E1B6F88AB4B2643A" blockId="3.[151,1436,1527,1912]" pageId="3" pageNumber="256">
<emphasis id="8E8DA21E1321FF86E1B6F88AB1FD647D" bold="true" box="[199,502,1815,1840]" pageId="3" pageNumber="256">Diagnosis and discussion.</emphasis>
Male imagos of 
<taxonomicName id="7BF9058F1321FF86E3C5F88AB3C7647D" box="[692,972,1815,1840]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1321FF86E3C5F88AB3C7647D" box="[692,972,1815,1840]" italics="true" pageId="3" pageNumber="256">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651321FF86E2A3F88AB41A647D" box="[978,1041,1815,1840]" pageId="3" pageNumber="256" rank="species">
<emphasis id="8E8DA21E1321FF86E2A3F88AB41A647D" bold="true" box="[978,1041,1815,1840]" pageId="3" pageNumber="256">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished from those of 
<taxonomicName id="7BF9058F1321FF86E1E6F8A1B14D6419" box="[151,326,1851,1876]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="hindustanica">
<emphasis id="8E8DA21E1321FF86E1E6F8A1B14D6419" box="[151,326,1851,1876]" italics="true" pageId="3" pageNumber="256">G. hindustanica</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BF9058F1321FF86E00CF8A1B1F86419" box="[381,499,1851,1876]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="256" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="8E8DA21E1321FF86E00CF8A1B1F86419" box="[381,499,1851,1876]" italics="true" pageId="3" pageNumber="256">G. pulchra</emphasis>
</taxonomicName>
by the following characters: 1) proportions of forelegs, 2) fore femur coloration, 3) abdominal terga patterns, 4) shape of apex of penis lobes, and 5) shape of hind wing (
<tableCitation id="F17B4BB71321FF86E525F8C2B4A3643A" box="[1108,1192,1887,1912]" captionStart="TABLE 1" captionStartId="4.[157,245,152,175]" captionTargetBox="[159,1410,202,338]" captionTargetPageId="4" captionText="TABLE 1. Comparison of imaginal characteristics of three species of Gilliesia (Gillies 1951, Zhou 2004)." httpUri="http://table.plazi.org/id/E8862E841326FF81E1ECFF05B50163E2" pageId="3" pageNumber="256" tableUuid="E8862E841326FF81E1ECFF05B50163E2">Table 1</tableCitation>
).
</paragraph>
<caption id="E8862E841326FF81E1ECFF05B50163E2" ID-Table-UUID="E8862E841326FF81E1ECFF05B50163E2" box="[157,1290,151,175]" httpUri="http://table.plazi.org/id/E8862E841326FF81E1ECFF05B50163E2" pageId="4" pageNumber="257" targetBox="[159,1410,202,338]" targetIsTable="true" targetPageId="4">
<paragraph id="BC467E0C1326FF81E1ECFF05B50163E2" blockId="4.[157,1290,151,175]" box="[157,1290,151,175]" pageId="4" pageNumber="257">
<emphasis id="8E8DA21E1326FF81E1ECFF05B11B63E2" bold="true" box="[157,272,152,175]" pageId="4" pageNumber="257">TABLE 1.</emphasis>
Comparison of imaginal characteristics of three species of 
<taxonomicName id="7BF9058F1326FF81E2F8FF0AB3EE63E2" box="[905,997,151,175]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E1326FF81E2F8FF0AB3EE63E2" box="[905,997,151,175]" italics="true" pageId="4" pageNumber="257">Gilliesia</emphasis>
</taxonomicName>
(Gillies 1951, Zhou 2004).
</paragraph>
</caption>
<paragraph id="BC467E0C1326FF81E0B3FF56B589621F" pageId="4" pageNumber="257">
<table id="CEF98CAC1326007AE1EEFF57B589621F" box="[159,1410,202,338]" gridcols="3" gridrows="3" pageId="4" pageNumber="257">
<tr id="02C97C4E1326007AE1EEFF57B58963AD" box="[159,1410,202,224]" gridrow="0" pageId="4" pageNumber="257" rowspan-0="1">
<th id="411815321326007AE0B3FF57B3FC63AD" box="[450,1015,202,224]" gridcol="1" gridrow="0" pageId="4" pageNumber="257">
<emphasis id="8E8DA21E1326FF81E0B3FF56B37263AD" box="[450,889,202,224]" italics="true" pageId="4" pageNumber="257">
<taxonomicName id="7BF9058F1326FF81E0B3FF56B26A63AD" box="[450,609,202,224]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="hindustanica">G. hindustanica</taxonomicName>
<taxonomicName id="7BF9058F1326FF81E27FFF56B37263AD" box="[782,889,202,224]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="pulchra">G. pulchra</taxonomicName>
</emphasis>
</th>
<th id="411815321326007AE52BFF57B58963AD" box="[1114,1410,202,224]" gridcol="2" gridrow="0" pageId="4" pageNumber="257">
<taxonomicName id="7BF9058F1326FF81E52BFF56B51E63AD" box="[1114,1301,202,224]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="ratchaburiensis">
<emphasis id="8E8DA21E1326FF81E52BFF56B51E63AD" box="[1114,1301,202,224]" italics="true" pageId="4" pageNumber="257">G. ratchaburiensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="02C97C4E1326007AE1EEFF69B5896247" box="[159,1410,244,266]" gridrow="1" pageId="4" pageNumber="257">
<th id="411815321326007AE1EEFF69B1016247" box="[159,266,244,266]" gridcol="0" gridrow="1" pageId="4" pageNumber="257">forelegs</th>
<td id="411815321326007AE0B3FF69B3FC6247" box="[450,1015,244,266]" gridcol="1" gridrow="1" pageId="4" pageNumber="257">tibiae equal in length to tarsi tibiae shorter than tarsi</td>
<td id="411815321326007AE52BFF69B5896247" box="[1114,1410,244,266]" gridcol="2" gridrow="1" pageId="4" pageNumber="257">tibiae longer than tarsi</td>
</tr>
<tr id="02C97C4E1326007AE1EEFEA1B589621F" box="[159,1410,316,338]" gridrow="2" pageId="4" pageNumber="257">
<th id="411815321326007AE1EEFEA1B101621F" box="[159,266,316,338]" gridcol="0" gridrow="2" pageId="4" pageNumber="257">fore femur</th>
<td id="411815321326007AE0B3FEA1B3FC621F" box="[450,1015,316,338]" gridcol="1" gridrow="2" pageId="4" pageNumber="257">dark brown grey to brown</td>
<td id="411815321326007AE52BFEA1B589621F" box="[1114,1410,316,338]" gridcol="2" gridrow="2" pageId="4" pageNumber="257">dark brown with median pale</td>
</tr>
</table>
</paragraph>
<paragraph id="BC467E0C1326FF81E1EEFE19B59962D7" blockId="4.[159,1426,388,440]" box="[159,1426,388,410]" pageId="4" pageNumber="257">abdominal terga dark brown with pitch brown translucent white on terga 1–8, dark brown on terga 1–10, with</paragraph>
<paragraph id="BC467E0C1326FF8DE0B3FE3EB4F4647A" blockId="4.[159,1426,388,440]" lastBlockId="8.[151,1437,1679,2028]" lastPageId="8" lastPageNumber="261" pageId="4" pageNumber="257">
on terga 1–8 with distinct markings distinct pale markings Nymphs of 
<taxonomicName id="7BF9058F132AFF8DE028F912B1B165E5" box="[345,442,1679,1704]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE028F912B1B165E5" box="[345,442,1679,1704]" italics="true" pageId="8" pageNumber="261">Gilliesia</emphasis>
</taxonomicName>
can be distinguished from those of the two 
<taxonomicName id="7BF9058F132AFF8DE295F912B49165E5" box="[996,1178,1679,1704]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="subFamily" subFamily="Leptophlebiinae">Leptophlebiinae</taxonomicName>
genera distributed in Southeast Asia (
<taxonomicName id="7BF9058F132AFF8DE021F92EB2146581" box="[336,543,1715,1740]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE021F92EB2146581" box="[336,543,1715,1740]" italics="true" pageId="8" pageNumber="261">Dipterophlebiodes</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BF9058F132AFF8DE329F92EB3126581" box="[600,793,1715,1740]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE329F92EB3126581" box="[600,793,1715,1740]" italics="true" pageId="8" pageNumber="261">Habrophlebiodes</emphasis>
</taxonomicName>
) by the posterolateral spines of the abdomen, gill shape, characteristics of the glossae, and maxillary palpi segment 3 (
<tableCitation id="F17B4BB7132AFF8DE201F94AB3C765A2" box="[880,972,1751,1776]" captionStart="TABLE 2" captionStartId="9.[151,239,548,571]" captionTargetBox="[159,1415,774,1288]" captionTargetPageId="9" captionText="TABLE 2. Comparison of nymphal characteristics of three related genera (Habrophlebiodes, Dipterophlebiodes and Gilliesia)." httpUri="http://table.plazi.org/id/E8862E84132BFF8CE1E6FDB9B1096116" pageId="8" pageNumber="261" tableUuid="E8862E84132BFF8CE1E6FDB9B1096116">Table 2</tableCitation>
). 
<taxonomicName id="7BF9058F132AFF8DE294F94AB44D65BD" box="[997,1094,1751,1776]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE294F94AB44D65BD" box="[997,1094,1751,1776]" italics="true" pageId="8" pageNumber="261">Gilliesia</emphasis>
</taxonomicName>
seems to be more similar to 
<taxonomicName id="7BF9058F132AFF8DE1E6F966B16D6459" box="[151,358,1787,1812]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE1E6F966B16D6459" box="[151,358,1787,1812]" italics="true" pageId="8" pageNumber="261">Dipterophlebiodes</emphasis>
</taxonomicName>
than to 
<taxonomicName id="7BF9058F132AFF8DE0B6F966B2826459" box="[455,649,1787,1812]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE0B6F966B2826459" box="[455,649,1787,1812]" italics="true" pageId="8" pageNumber="261">Habrophlebiodes</emphasis>
</taxonomicName>
by the following characters: posterolateral spines well-developed on abdominal segment 9 only, and gills slender and slightly forked at&gt;1/3 distance from base (
<tableCitation id="F17B4BB7132AFF8DE5EBF882B4F9647A" box="[1178,1266,1823,1848]" captionStart="TABLE 2" captionStartId="9.[151,239,548,571]" captionTargetBox="[159,1415,774,1288]" captionTargetPageId="9" captionText="TABLE 2. Comparison of nymphal characteristics of three related genera (Habrophlebiodes, Dipterophlebiodes and Gilliesia)." httpUri="http://table.plazi.org/id/E8862E84132BFF8CE1E6FDB9B1096116" pageId="8" pageNumber="261" tableUuid="E8862E84132BFF8CE1E6FDB9B1096116">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<paragraph id="BC467E0C1326FF81E1EEFE77B4B46005" pageId="4" pageNumber="257">
<table id="CEF98CAC1326007AE1EEFE77B57D6005" box="[159,1398,490,840]" gridcols="4" gridrows="3" pageId="4" pageNumber="257">
<tr id="02C97C4E1326007AE1EEFE77B57D6173" box="[159,1398,490,574]" gridrow="0" pageId="4" pageNumber="257">
<th id="411815321326007AE1EEFE77B1AB6173" box="[159,416,490,574]" gridcol="0" gridrow="0" pageId="4" pageNumber="257">penis</th>
<th id="411815321326007AE0B3FE77B2FA6173" box="[450,753,490,574]" gridcol="1" gridrow="0" pageId="4" pageNumber="257">apex of the penis lobes broad, each lobe bent laterally and then ventrally</th>
<th id="411815321326007AE27FFE77B43C6173" box="[782,1079,490,574]" gridcol="2" gridrow="0" pageId="4" pageNumber="257">apex of the penis taped, each lobe bent laterally and then ventrally</th>
<th id="411815321326007AE52BFE77B57D6173" box="[1114,1398,490,574]" gridcol="3" gridrow="0" pageId="4" pageNumber="257">apex of the penis lobes sharpened, each lobe bent laterally</th>
</tr>
<tr id="02C97C4E1326007AE1EEFDEDB57D61E9" box="[159,1398,624,676]" gridrow="1" pageId="4" pageNumber="257">
<th id="411815321326007AE1EEFDEDB1AB61E9" box="[159,416,624,676]" gridcol="0" gridrow="1" pageId="4" pageNumber="257">apex of female sternum 9</th>
<td id="411815321326007AE0B3FDEDB2FA61E9" box="[450,753,624,676]" gridcol="1" gridrow="1" pageId="4" pageNumber="257">with V-shaped deep median cleft</td>
<td id="411815321326007AE27FFDEDB43C61E9" box="[782,1079,624,676]" gridcol="2" gridrow="1" pageId="4" pageNumber="257">with V-shaped deep median cleft</td>
<td id="411815321326007AE52BFDEDB57D61E9" box="[1114,1398,624,676]" gridcol="3" gridrow="1" pageId="4" pageNumber="257">with U-shaped deep median cleft</td>
</tr>
<tr id="02C97C4E1326007AE1EEFD4BB57D6005" box="[159,1398,726,840]" gridrow="2" pageId="4" pageNumber="257">
<th id="411815321326007AE1EEFD4BB1AB6005" box="[159,416,726,840]" gridcol="0" gridrow="2" pageId="4" pageNumber="257">shape of the hind wing</th>
<td id="411815321326007AE0B3FD4BB2FA6005" box="[450,753,726,840]" gridcol="1" gridrow="2" pageId="4" pageNumber="257">costal projection well developed and rounded, apex located about 1/2 distance from base</td>
<td id="411815321326007AE27FFD4BB43C6005" box="[782,1079,726,840]" gridcol="2" gridrow="2" pageId="4" pageNumber="257">costal projection well developed and rounded, apex located about 2/3 distance from base</td>
<td id="411815321326007AE52BFD4BB57D6005" box="[1114,1398,726,840]" gridcol="3" gridrow="2" pageId="4" pageNumber="257">costal projection well developed and acute, apex located about 3/4 distance from base</td>
</tr>
</table>
</paragraph>
<caption id="E8862E841326FF81E1E6F8A9B3426427" httpUri="https://zenodo.org/record/237017/files/figure.png" pageId="4" pageNumber="257" targetBox="[190,1396,917,1823]" targetPageId="4">
<paragraph id="BC467E0C1326FF81E1E6F8A9B3426427" blockId="4.[151,1436,1844,1898]" pageId="4" pageNumber="257">
<emphasis id="8E8DA21E1326FF81E1E6F8A9B1476407" bold="true" box="[151,332,1844,1867]" pageId="4" pageNumber="257">FIGURES 10–13</emphasis>
. Nymphal details (SEM) of 
<taxonomicName id="7BF9058F1326FF81E31BF8A8B36D6406" box="[618,870,1845,1867]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1326FF81E31BF8A8B36D6406" box="[618,870,1845,1867]" italics="true" pageId="4" pageNumber="257">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<emphasis id="8E8DA21E1326FF81E21DF8ABB3AF6406" bold="true" box="[876,932,1846,1867]" pageId="4" pageNumber="257">
<taxonomicNameLabel id="95BE1F651326FF81E21DF8ABB3AF6406" box="[876,932,1846,1867]" pageId="4" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(10) labium in ventral view; (11) labium in dorsal view; (12) apical segment of labium; (13) paraglossae, ventral view.
</paragraph>
</caption>
<caption id="E8862E841327FF80E1E6F83CB3E564B9" httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="5" pageNumber="258" targetBox="[227,1338,217,1909]" targetPageId="5">
<paragraph id="BC467E0C1327FF80E1E6F83CB3E564B9" blockId="5.[151,1436,1953,2036]" pageId="5" pageNumber="258">
<emphasis id="8E8DA21E1327FF80E1E6F83CB14764FB" bold="true" box="[151,332,1953,1976]" pageId="5" pageNumber="258">FIGURES 14–22</emphasis>
. Nymph of 
<taxonomicName id="7BF9058F1327FF80E0B7F83CB2C964FA" box="[454,706,1953,1975]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1327FF80E0B7F83CB2C964FA" box="[454,706,1953,1975]" italics="true" pageId="5" pageNumber="258">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651327FF80E3B8F83FB30A64FA" box="[713,769,1954,1975]" pageId="5" pageNumber="258" rank="species">
<emphasis id="8E8DA21E1327FF80E3B8F83FB30A64FA" bold="true" box="[713,769,1954,1975]" pageId="5" pageNumber="258">sp. n.</emphasis>
</taxonomicNameLabel>
(14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw;(21) abdominal segment 8–10, dorsal view; (22) gill 4.
</paragraph>
</caption>
<caption id="E8862E841324FF83E1E6F961B343647F" httpUri="https://zenodo.org/record/237019/files/figure.png" pageId="6" pageNumber="259" targetBox="[179,1418,212,1746]" targetPageId="6">
<paragraph id="BC467E0C1324FF83E1E6F961B343647F" blockId="6.[151,1435,1788,1842]" pageId="6" pageNumber="259">
<emphasis id="8E8DA21E1324FF83E1E6F961B145645F" bold="true" box="[151,334,1788,1811]" pageId="6" pageNumber="259">FIGURES 23–29</emphasis>
. Male imago of 
<taxonomicName id="7BF9058F1324FF83E08AF960B2F2645E" box="[507,761,1789,1811]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1324FF83E08AF960B2F2645E" box="[507,761,1789,1811]" italics="true" pageId="6" pageNumber="259">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<emphasis id="8E8DA21E1324FF83E271F960B331645F" bold="true" box="[768,826,1789,1810]" pageId="6" pageNumber="259">
<taxonomicNameLabel id="95BE1F651324FF83E271F960B331645F" box="[768,826,1789,1810]" pageId="6" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(23) head, dorsal view; (24) forewing; (25) hindwing; (26) foreleg; (27) middle leg; (28) hind leg; (29) abdomen in dorsal view
</paragraph>
</caption>
<caption id="E8862E841325FF82E1E6F8DEB0C564DA" httpUri="https://zenodo.org/record/237020/files/figure.png" pageId="7" pageNumber="260" targetBox="[151,1436,221,1837]" targetPageId="7">
<paragraph id="BC467E0C1325FF82E1E6F8DEB0C564DA" blockId="7.[151,1436,1859,1943]" pageId="7" pageNumber="260">
<emphasis id="8E8DA21E1325FF82E1E6F8DEB1466414" bold="true" box="[151,333,1859,1882]" pageId="7" pageNumber="260">FIGURES 30–34</emphasis>
. Imagos of 
<taxonomicName id="7BF9058F1325FF82E0B6F8D9B2CF6417" box="[455,708,1860,1882]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1325FF82E0B6F8D9B2CF6417" box="[455,708,1860,1882]" italics="true" pageId="7" pageNumber="260">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F651325FF82E3BBF8D9B3086414" box="[714,771,1860,1881]" pageId="7" pageNumber="260" rank="species">
<emphasis id="8E8DA21E1325FF82E3BBF8D9B3086414" bold="true" box="[714,771,1860,1881]" pageId="7" pageNumber="260">sp. n.</emphasis>
</taxonomicNameLabel>
(30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view.
</paragraph>
</caption>
<caption id="E8862E84132AFF8DE1E6FC1CB2FD60FB" httpUri="https://zenodo.org/record/237021/files/figure.png" pageId="8" pageNumber="261" targetBox="[340,1247,193,875]" targetPageId="8">
<paragraph id="BC467E0C132AFF8DE1E6FC1CB2FD60FB" blockId="8.[151,1436,897,950]" pageId="8" pageNumber="261">
<emphasis id="8E8DA21E132AFF8DE1E6FC1CB14760DB" bold="true" box="[151,332,897,920]" pageId="8" pageNumber="261">FIGURES 35–38</emphasis>
. SEM pictures of eggs extracted from a female imago of 
<taxonomicName id="7BF9058F132AFF8DE2FFFC1CB48160DA" box="[910,1162,897,919]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E132AFF8DE2FFFC1CB48160DA" box="[910,1162,897,919]" italics="true" pageId="8" pageNumber="261">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<emphasis id="8E8DA21E132AFF8DE5E1FC1FB4C360DA" bold="true" box="[1168,1224,898,919]" pageId="8" pageNumber="261">
<taxonomicNameLabel id="95BE1F65132AFF8DE5E1FC1FB4C360DA" box="[1168,1224,898,919]" pageId="8" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(35) general outline; (36–37) detail of pole; (38) detail of the chorionic structure.
</paragraph>
</caption>
<caption id="E8862E84132AFF8DE1E6F9AFB1BB6525" httpUri="https://zenodo.org/record/237022/files/figure.png" pageId="8" pageNumber="261" targetBox="[340,1246,1000,1565]" targetPageId="8">
<paragraph id="BC467E0C132AFF8DE1E6F9AFB1BB6525" blockId="8.[151,1436,1586,1640]" pageId="8" pageNumber="261">
<emphasis id="8E8DA21E132AFF8DE1E6F9AFB1466505" bold="true" box="[151,333,1586,1609]" pageId="8" pageNumber="261">FIGURES 39–40</emphasis>
. Inner-apical angle of maxilla of 
<taxonomicName id="7BF9058F132AFF8DE3D5F9AEB3AA6504" box="[676,929,1587,1609]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E132AFF8DE3D5F9AEB3AA6504" box="[676,929,1587,1609]" italics="true" pageId="8" pageNumber="261">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132AFF8DE2D9F9AEB3E96505" box="[936,994,1587,1608]" pageId="8" pageNumber="261" rank="species">
<emphasis id="8E8DA21E132AFF8DE2D9F9AEB3E96505" bold="true" box="[936,994,1587,1608]" pageId="8" pageNumber="261">sp. n.</emphasis>
</taxonomicNameLabel>
(39) and 
<taxonomicName id="7BF9058F132AFF8DE534F9AEB56E6504" box="[1093,1381,1587,1609]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="8E8DA21E132AFF8DE534F9AEB56E6504" box="[1093,1381,1587,1609]" italics="true" pageId="8" pageNumber="261">Habrophlebiodes prominens</emphasis>
</taxonomicName>
(40), view in optical microscope.
</paragraph>
</caption>
<subSubSection id="F4E32D87132AFF8FE1B6F8DEB36F6399" lastPageId="10" lastPageNumber="263" pageId="8" pageNumber="261" type="reference_group">
<paragraph id="BC467E0C132AFF8DE1B6F8DEB29F6485" blockId="8.[151,1437,1679,2028]" pageId="8" pageNumber="261">
<taxonomicName id="7BF9058F132AFF8DE1B6F8DEB1236411" ID-CoL="8V5TD" box="[199,296,1859,1884]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132AFF8DE1B6F8DEB1236411" box="[199,296,1859,1884]" italics="true" pageId="8" pageNumber="261">Gilliesia</emphasis>
</taxonomicName>
nymphs have all attributes of the subfamily 
<taxonomicName id="7BF9058F132AFF8DE25EF8DEB3EE6411" box="[815,997,1859,1884]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="subFamily" subFamily="Leptophlebiinae">Leptophlebiinae</taxonomicName>
as defined by 
<bibRefCitation id="D86803FD132AFF8DE5E3F8DEB52F6411" author="Kluge" box="[1170,1316,1859,1884]" pageId="8" pageNumber="261" refString="Kluge, N. J. (1994) Habrophlebiinae subfam. n. with description of a new species of Habroleptoides from the Caucasus (Ephemeroptera: Leptophlebiidae). Zoosystematica Rossica, 3, 35 - 43." type="journal article" year="1994">Kluge (1994</bibRefCitation>
, 
<bibRefCitation id="D86803FD132AFF8DE443F8D9B5646411" author="Kluge" box="[1330,1391,1860,1884]" pageId="8" pageNumber="261" refString="Kluge, N. J. (2009) Higher system of Atalophlebiinae (Leptophlebiidae) with description of three new species of Terpides s. l. from Peruvian Amazonia. Russian Entomological Journal, 18 (4), 243 - 256." type="journal article" year="2009">2009</bibRefCitation>
): 1) apex of maxilla with three reduced canines; proximal and distal dentisetae present and entire; 2) no transverse row of setae on the dorsal face of labrum; 3) tibio-patellar suture present on mid- and hindlegs; and 4) superlinguae of hypopharynx rounded, not extended laterally.
</paragraph>
<paragraph id="BC467E0C132AFF8CE1B6F84EB3276250" blockId="8.[151,1437,1679,2028]" lastBlockId="9.[151,1436,151,501]" lastPageId="9" lastPageNumber="262" pageId="8" pageNumber="261">
All these characters are considered by 
<bibRefCitation id="D86803FD132AFF8DE30EF84EB31C64A1" author="Kluge" box="[639,791,2003,2028]" pageId="8" pageNumber="261" refString="Kluge, N. J. (1994) Habrophlebiinae subfam. n. with description of a new species of Habroleptoides from the Caucasus (Ephemeroptera: Leptophlebiidae). Zoosystematica Rossica, 3, 35 - 43." type="journal article" year="1994">Kluge (1994)</bibRefCitation>
as plesiomorphic, hence the non-ranking plesiomorphon 
<taxonomicName id="7BF9058F132BFF8CE1E6FF0AB12763FD" ID-CoL="5D97" box="[151,300,151,176]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">Leptophlebia</taxonomicName>
/fg2 proposed by him (
<bibRefCitation id="D86803FD132BFF8CE345FF0AB2CA63FD" author="Kluge" box="[564,705,151,176]" pageId="9" pageNumber="262" refString="Kluge, N. J. (2009) Higher system of Atalophlebiinae (Leptophlebiidae) with description of three new species of Terpides s. l. from Peruvian Amazonia. Russian Entomological Journal, 18 (4), 243 - 256." type="journal article" year="2009">Kluge 2009</bibRefCitation>
). Besides 
<taxonomicName id="7BF9058F132BFF8CE247FF0AB39D63FD" ID-CoL="8V5TD" box="[822,918,151,176]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE247FF0AB39D63FD" box="[822,918,151,176]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
, 
<taxonomicName id="7BF9058F132BFF8CE2D9FF0AB45563FD" box="[936,1118,151,176]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Leptophlebiinae">Leptophlebiinae</taxonomicName>
encompasses the following genera: 
<taxonomicName id="7BF9058F132BFF8CE182FF21B2446399" ID-CoL="5D97" authority="Westwood, 1940" authorityName="Westwood" authorityYear="1940" box="[243,591,188,213]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE182FF21B18C6398" box="[243,391,188,213]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
Westwood, 1940
</taxonomicName>
(with three subgenera 
<taxonomicName id="7BF9058F132BFF8CE22DFF21B3FB6398" ID-CoL="5D97" box="[860,1008,188,213]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8E8DA21E132BFF8CE22DFF21B3FB6398" box="[860,1008,188,213]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE288FF20B4156399" box="[1017,1054,189,212]" pageId="9" pageNumber="262" sensu="stricto">s.s.</taxonomicNameLabel>
, 
<taxonomicName id="7BF9058F132BFF8CE55FFF21B5976399" ID-CoL="6F8B" authority="Lestage, 1917" authorityName="Lestage" authorityYear="1917" box="[1070,1436,188,213]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE55FFF21B4F96398" box="[1070,1266,188,213]" italics="true" pageId="9" pageNumber="262">Paraleptophlebia</emphasis>
Lestage, 1917
</taxonomicName>
and 
<taxonomicName id="7BF9058F132BFF8CE1B6FF42B21363B5" authority="Kluge, 1997" authorityName="Kluge" authorityYear="1997" box="[199,536,223,248]" class="Insecta" family="Leptophlebiidae" genus="Neoleptophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1B6FF42B18B63B5" box="[199,384,223,248]" italics="true" pageId="9" pageNumber="262">Neoleptophlebia</emphasis>
Kluge, 1997
</taxonomicName>
), 
<taxonomicName id="7BF9058F132BFF8CE35CFF42B38F63B5" ID-CoL="4S9S" authority="Ulmer, 1920" authorityName="Ulmer" authorityYear="1920" box="[557,900,223,248]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE35CFF42B2E563B5" box="[557,750,223,248]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
Ulmer, 1920
</taxonomicName>
and 
<taxonomicName id="7BF9058F132BFF8CE2CDFF42B54563B5" ID-CoL="46J3" authority="Demoulin, 1954" authorityName="Demoulin" authorityYear="1954" box="[956,1358,223,248]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE2CDFF42B48063B5" box="[956,1163,223,248]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
Demoulin, 1954
</taxonomicName>
. Many authors treat 
<taxonomicName id="7BF9058F132BFF8CE058FE99B1E66250" ID-CoL="6F8B" box="[297,493,260,285]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE058FE99B1E66250" box="[297,493,260,285]" italics="true" pageId="9" pageNumber="262">Paraleptophlebia</emphasis>
</taxonomicName>
at the genus level, however.
</paragraph>
<paragraph id="BC467E0C132BFF8CE1B6FEBAB53262C5" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="262">
<taxonomicName id="7BF9058F132BFF8CE1B6FEBAB123620D" ID-CoL="8V5TD" box="[199,296,295,320]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1B6FEBAB123620D" box="[199,296,295,320]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
gills resemble somewhat those of 
<taxonomicName id="7BF9058F132BFF8CE3C1FEBAB34F620D" ID-CoL="5D97" box="[688,836,295,320]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8E8DA21E132BFF8CE3C1FEBAB34F620D" box="[688,836,295,320]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE23DFEB5B37B6272" box="[844,880,296,319]" pageId="9" pageNumber="262" sensu="stricto">s.s.</taxonomicNameLabel>
, being large in the proximal part before the fork, but contrary to 
<taxonomicName id="7BF9058F132BFF8CE023FED1B1ED6228" ID-CoL="5D97" box="[338,486,332,357]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8E8DA21E132BFF8CE023FED1B1ED6228" box="[338,486,332,357]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE083FED0B21D6229" box="[498,534,333,356]" pageId="9" pageNumber="262" sensu="stricto">s.s.</taxonomicNameLabel>
the fork is not suddenly constricted to form a single filament but regularly decreasing, much more similar to what is found in 
<taxonomicName id="7BF9058F132BFF8CE3A1FEF2B53E62C5" ID-CoL="62TYG" authority="Schonemund, 1929 (Habrophlebiinae)" authorityName="Schonemund, 1929 (Habrophlebiinae" box="[720,1333,367,392]" class="Insecta" family="Leptophlebiidae" genus="Habroleptoides" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE3A1FEF2B37662C5" box="[720,893,367,392]" italics="true" pageId="9" pageNumber="262">Habroleptoides</emphasis>
Schönemund, 1929 (Habrophlebiinae)
</taxonomicName>
.
</paragraph>
<paragraph id="BC467E0C132BFF8CE1B6FE09B21F62B8" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="262">
<taxonomicName id="7BF9058F132BFF8CE1B6FE09B12362E0" ID-CoL="8V5TD" box="[199,296,404,429]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1B6FE09B12362E0" box="[199,296,404,429]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
tarsal claws are rather stout and not as elongated as in 
<taxonomicName id="7BF9058F132BFF8CE2F6FE09B41062E0" ID-CoL="5D97" box="[903,1051,404,429]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="8E8DA21E132BFF8CE2F6FE09B41062E0" box="[903,1051,404,429]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE550FE09B44962E0" box="[1057,1090,404,429]" pageId="9" pageNumber="262" sensu="lato">s.l.</taxonomicNameLabel>
They are more similar to those of 
<taxonomicName id="7BF9058F132BFF8CE1CAFE2AB181629D" ID-CoL="46J3" box="[187,394,439,464]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1CAFE2AB181629D" box="[187,394,439,464]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
or 
<taxonomicName id="7BF9058F132BFF8CE0CDFE2AB276629D" ID-CoL="4S9S" box="[444,637,439,464]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE0CDFE2AB276629D" box="[444,637,439,464]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
, the row of teeth extending over ¾ of the claw length, and teeth increasing slightly in size distally.
</paragraph>
<caption id="E8862E84132BFF8CE1E6FDB9B1096116" ID-Table-UUID="E8862E84132BFF8CE1E6FDB9B1096116" httpUri="http://table.plazi.org/id/E8862E84132BFF8CE1E6FDB9B1096116" pageId="9" pageNumber="262" targetBox="[159,1415,774,1288]" targetIsTable="true" targetPageId="9">
<paragraph id="BC467E0C132BFF8CE1E6FDB9B1096116" blockId="9.[151,1436,547,603]" pageId="9" pageNumber="262">
<emphasis id="8E8DA21E132BFF8CE1E6FDB9B10C6176" bold="true" box="[151,263,548,571]" pageId="9" pageNumber="262">TABLE 2</emphasis>
. Comparison of nymphal characteristics of three related genera (
<taxonomicName id="7BF9058F132BFF8CE2ADFDBEB49F6176" ID-CoL="4S9S" box="[988,1172,547,571]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE2ADFDBEB49F6176" box="[988,1172,547,571]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="7BF9058F132BFF8CE5D4FDBEB5616176" ID-CoL="46J3" box="[1189,1386,547,571]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE5D4FDBEB5616176" box="[1189,1386,547,571]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BF9058F132BFF8CE1E6FDDEB0F86116" ID-CoL="8V5TD" box="[151,243,579,603]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1E6FDDEB0F86116" box="[151,243,579,603]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="BC467E0C132BFF8CE0BEFDEBB48C61C1" blockId="9.[159,1371,630,724]" box="[463,1159,630,652]" pageId="9" pageNumber="262">
<emphasis id="8E8DA21E132BFF8CE0BEFDEBB48C61C1" box="[463,1159,630,652]" italics="true" pageId="9" pageNumber="262">
<taxonomicName id="7BF9058F132BFF8CE0BEFDEBB27561C1" ID-CoL="4S9S" box="[463,638,630,652]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">Habrophlebiodes</taxonomicName>
<taxonomicName id="7BF9058F132BFF8CE38EFDEBB3B061C1" ID-CoL="46J3" box="[767,955,630,652]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">Dipterophlebiodes</taxonomicName>
<taxonomicName id="7BF9058F132BFF8CE541FDEBB48C61C1" ID-CoL="8V5TD" box="[1072,1159,630,652]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">Gilliesia</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="BC467E0C132BFF8CE1EEFD3DB4D86199" blockId="9.[159,1371,630,724]" pageId="9" pageNumber="262">posterolateral expansions of well developed on well developed on well developed on abdominal abdomen abdominal segments 8 and 9 abdominal segments 9 only segments 9 only</paragraph>
<paragraph id="BC467E0C132BFF8CE1EEFC9BB55267A7" pageId="9" pageNumber="262">
<table id="CEF98CAC132B007AE1EEFC9BB58C6645" box="[159,1415,774,1288]" gridcols="4" gridrows="4" pageId="9" pageNumber="262">
<tr id="02C97C4E132B007AE1EEFC9BB58C6076" box="[159,1415,774,827]" gridrow="0" pageId="9" pageNumber="262">
<th id="41181532132B007AE1EEFC9BB1996076" box="[159,402,774,827]" gridcol="0" gridrow="0" pageId="9" pageNumber="262">gills 4</th>
<th id="41181532132B007AE0BEFC9BB2E16076" box="[463,746,774,827]" gridcol="1" gridrow="0" pageId="9" pageNumber="262">gills long, more slender and deeply forked at 1/6 basally</th>
<th id="41181532132B007AE38EFC9BB4106076" box="[767,1051,774,827]" gridcol="2" gridrow="0" pageId="9" pageNumber="262">gills long, slender and deeply forked at 1/3 basally</th>
<th id="41181532132B007AE541FC9BB58C6076" box="[1072,1415,774,827]" gridcol="3" gridrow="0" pageId="9" pageNumber="262">gills long, slender and slightly forked at 2/5 basally</th>
</tr>
<tr id="02C97C4E132B007AE1EEFCF0B58C6092" box="[159,1415,877,991]" gridrow="1" pageId="9" pageNumber="262">
<th id="41181532132B007AE1EEFCF0B1996092" box="[159,402,877,991]" gridcol="0" gridrow="1" pageId="9" pageNumber="262">hypopharynx</th>
<td id="41181532132B007AE0BEFCF0B2E16092" box="[463,746,877,991]" gridcol="1" gridrow="1" pageId="9" pageNumber="262">lingua rectangular or cordate with two dense tufts of setae, superlingua slightly extended laterally</td>
<td id="41181532132B007AE38EFCF0B4106092" box="[767,1051,877,991]" gridcol="2" gridrow="1" pageId="9" pageNumber="262">lingua rectangular with two dense tufts of setae, superlingua not extended laterally</td>
<td id="41181532132B007AE541FCF0B58C6092" box="[1072,1415,877,991]" gridcol="3" gridrow="1" pageId="9" pageNumber="262">lingua rectangular with two dense tufts of setae,, superlingua not extended laterally</td>
</tr>
<tr id="02C97C4E132B007AE1EEFB8CB58C6729" box="[159,1415,1041,1124]" gridrow="2" pageId="9" pageNumber="262">
<th id="41181532132B007AE1EEFB8CB1996729" box="[159,402,1041,1124]" gridcol="0" gridrow="2" pageId="9" pageNumber="262">glossae</th>
<td id="41181532132B007AE0BEFB8CB2E16729" box="[463,746,1041,1124]" gridcol="1" gridrow="2" pageId="9" pageNumber="262">taped, with scattered long hairs on ventral surface</td>
<td id="41181532132B007AE38EFB8CB4106729" box="[767,1051,1041,1124]" gridcol="2" gridrow="2" pageId="9" pageNumber="262">conical, with marginal long hairs on ventral surface</td>
<td id="41181532132B007AE541FB8CB58C6729" box="[1072,1415,1041,1124]" gridcol="3" gridrow="2" pageId="9" pageNumber="262">narrow taped, with dense thickened-long hairs on ventral surface</td>
</tr>
<tr id="02C97C4E132B007AE1EEFB0BB58C6645" box="[159,1415,1174,1288]" gridrow="3" pageId="9" pageNumber="262">
<th id="41181532132B007AE1EEFB0BB1996645" box="[159,402,1174,1288]" gridcol="0" gridrow="3" pageId="9" pageNumber="262">maxilla palpi segment 3</th>
<td id="41181532132B007AE0BEFB0BB2E16645" box="[463,746,1174,1288]" gridcol="1" gridrow="3" pageId="9" pageNumber="262">length shorter than 1.6 times the length of segment 2; apical-pointed, with scattered setae</td>
<td id="41181532132B007AE38EFB0BB4106645" box="[767,1051,1174,1288]" gridcol="2" gridrow="3" pageId="9" pageNumber="262">length shorter than 1.6 times the length of segment 2; apical-blunted, with few setae</td>
<td id="41181532132B007AE541FB0BB58C6645" box="[1072,1415,1174,1288]" gridcol="3" gridrow="3" pageId="9" pageNumber="262">length more than 1.6 times the length of segment 2; apicalblunted, with numerous setae</td>
</tr>
</table>
</paragraph>
<paragraph id="BC467E0C132BFF8CE1B6FAD3B34E652F" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="262">
In 
<taxonomicName id="7BF9058F132BFF8CE199FAD0B143662B" ID-CoL="8V5TD" box="[232,328,1357,1382]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE199FAD0B143662B" box="[232,328,1357,1382]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
, the posterolateral spines are only developed on abdominal segment 9, as in 
<taxonomicName id="7BF9058F132BFF8CE5B4FAD0B59F662B" ID-CoL="46J3" box="[1221,1428,1357,1382]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE5B4FAD0B59F662B" box="[1221,1428,1357,1382]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
, whereas they are also present on segment 
<quantity id="7B01D3E9132BFF8CE302FAEFB29466C7" box="[627,671,1393,1418]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="9" pageNumber="262" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="7BF9058F132BFF8CE3D7FAECB33166C7" ID-CoL="5D97" box="[678,826,1393,1418]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="8E8DA21E132BFF8CE3D7FAECB33166C7" box="[678,826,1393,1418]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE233FAECB36866C7" box="[834,867,1393,1418]" pageId="9" pageNumber="262" sensu="lato">s.l.</taxonomicNameLabel>
(strongly developed in 
<taxonomicName id="7BF9058F132BFF8CE501FAECB50F66C7" ID-CoL="5D97" box="[1136,1284,1393,1418]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="8E8DA21E132BFF8CE501FAECB50F66C7" box="[1136,1284,1393,1418]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE47AFAEFB52466C4" box="[1291,1327,1394,1417]" pageId="9" pageNumber="262" sensu="stricto">s.s.</taxonomicNameLabel>
but more reduced in some 
<taxonomicName id="7BF9058F132BFF8CE013FA08B22D66E3" ID-CoL="6F8B" box="[354,550,1429,1454]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE013FA08B22D66E3" box="[354,550,1429,1454]" italics="true" pageId="9" pageNumber="262">Paraleptophlebia</emphasis>
</taxonomicName>
) and 
<taxonomicName id="7BF9058F132BFF8CE31CFA08B32566E3" ID-CoL="4S9S" box="[621,814,1429,1454]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE31CFA08B32566E3" box="[621,814,1429,1454]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
. In addition, maxillary canines of 
<taxonomicName id="7BF9058F132BFF8CE5B8FA08B52266E3" ID-CoL="8V5TD" box="[1225,1321,1429,1454]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE5B8FA08B52266E3" box="[1225,1321,1429,1454]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
are more reduced than in 
<taxonomicName id="7BF9058F132BFF8CE020FA24B218669F" ID-CoL="4S9S" box="[337,531,1465,1490]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE020FA24B218669F" box="[337,531,1465,1490]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
, (
<figureCitation id="24C26289132BFF8CE35BFA24B2BB669F" box="[554,688,1465,1490]" captionStart="FIGURES 39 – 40" captionStartId="8.[151,264,1586,1609]" captionTargetBox="[340,1246,1000,1565]" captionTargetId="figure@8.[340,1247,1000,1566]" captionTargetPageId="8" captionText="FIGURES 39 – 40. Inner-apical angle of maxilla of Gilliesia ratchaburiensis sp. n. (39) and Habrophlebiodes prominens (40), view in optical microscope." httpUri="https://zenodo.org/record/237022/files/figure.png" pageId="9" pageNumber="262">Figs 39–40</figureCitation>
), whereas 
<bibRefCitation id="D86803FD132BFF8CE25BFA27B3C9669F" author="Peters" box="[810,962,1466,1490]" pageId="9" pageNumber="262" refString="Peters, W. L. (1972) The nymph of Dipterophlebiodes sp. (Leptophlebiidae: Ephemeroptera). Entomological News, 83, 53 - 56." type="journal article" year="1972">Peters (1972)</bibRefCitation>
did not mention the details of maxilla in 
<taxonomicName id="7BF9058F132BFF8CE1E6FA40B16D66BB" ID-CoL="46J3" box="[151,358,1501,1526]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1E6FA40B16D66BB" box="[151,358,1501,1526]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
. The shape of 
<taxonomicName id="7BF9058F132BFF8CE352FA40B28F66BB" ID-CoL="8V5TD" box="[547,644,1501,1526]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE352FA40B28F66BB" box="[547,644,1501,1526]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
superlinguae resembles those of 
<taxonomicName id="7BF9058F132BFF8CE564FA40B4EF66BB" ID-CoL="46J3" box="[1045,1252,1501,1526]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE564FA40B4EF66BB" box="[1045,1252,1501,1526]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
, which are not extended laterally, whereas in 
<taxonomicName id="7BF9058F132BFF8CE086F99CB2B36557" ID-CoL="4S9S" box="[503,696,1537,1562]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE086F99CB2B36557" box="[503,696,1537,1562]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
they are slightly extended (
<bibRefCitation id="D86803FD132BFF8CE28FF99FB59B6557" author="Peters" box="[1022,1424,1537,1562]" pageId="9" pageNumber="262" refString="Peters, W. L. &amp; Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters &amp; Edmunds 1970, Fig. 200</bibRefCitation>
). 
<taxonomicName id="7BF9058F132BFF8CE1E6F9B8B0F36573" ID-CoL="8V5TD" box="[151,248,1573,1598]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1E6F9B8B0F36573" box="[151,248,1573,1598]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
possess two dense tufts of setae on ventral side of the lingua, as in 
<emphasis id="8E8DA21E132BFF8CE29FF9B8B4B26573" box="[1006,1209,1573,1598]" italics="true" pageId="9" pageNumber="262">
<taxonomicName id="7BF9058F132BFF8CE29FF9B8B4B96573" ID-CoL="4S9S" box="[1006,1202,1573,1598]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">Habrophlebiodes</taxonomicName>
;
</emphasis>
this character is not mentioned for 
<taxonomicName id="7BF9058F132BFF8CE04DF9D4B200652F" ID-CoL="46J3" box="[316,523,1609,1634]" class="Insecta" family="Leptophlebiidae" genus="Dipterophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE04DF9D4B200652F" box="[316,523,1609,1634]" italics="true" pageId="9" pageNumber="262">Dipterophlebiodes</emphasis>
</taxonomicName>
(
<bibRefCitation id="D86803FD132BFF8CE368F9D7B333652F" author="Peters" box="[537,824,1609,1634]" pageId="9" pageNumber="262" refString="Peters, W. L. (1972) The nymph of Dipterophlebiodes sp. (Leptophlebiidae: Ephemeroptera). Entomological News, 83, 53 - 56." type="journal article" year="1972" yearSuffix="p">
Peters 1972, 
<figureCitation id="24C26289132BFF8CE3DCF9D4B2FB652F" box="[685,752,1609,1634]" captionStart="FIGURES 2 – 5" captionStartId="2.[151,264,1630,1653]" captionTargetBox="[168,1417,661,1599]" captionTargetId="figure@2.[168,1418,661,1599]" captionTargetPageId="2" captionText="FIGURES 2 – 5. Nymphal details of Gilliesia ratchaburiensis sp. n. (2) labrum in dorsal view; (3) labrum in ventral view; (4) median cleft of labrum, ventral view; (5) hypopharynx, ventral view." httpUri="https://zenodo.org/record/237015/files/figure.png" pageId="9" pageNumber="262">Fig. 5</figureCitation>
, p. 54
</bibRefCitation>
).
</paragraph>
<paragraph id="BC467E0C132BFF8CE1B6F9F3B3326583" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="262">
At the imaginal stages, too, 
<taxonomicName id="7BF9058F132BFF8CE36AF9F0B27065CB" ID-CoL="8V5TD" box="[539,635,1645,1670]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE36AF9F0B27065CB" box="[539,635,1645,1670]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
is easily distinguished from 
<taxonomicName id="7BF9058F132BFF8CE2A8F9F0B46665CB" ID-CoL="5D97" box="[985,1133,1645,1670]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="8E8DA21E132BFF8CE2A8F9F0B46665CB" box="[985,1133,1645,1670]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE508F9F0B49165CB" box="[1145,1178,1645,1670]" pageId="9" pageNumber="262" sensu="lato">s.l.</taxonomicNameLabel>
by the presence of a hindwing costal process, and by the male penis lobes which do not possess “a ventral appendage arising from apex of each penis lobe” sensu 
<bibRefCitation id="D86803FD132BFF8CE0CBF92BB33E6583" author="Peters" box="[442,821,1717,1742]" pageId="9" pageNumber="262" refString="Peters, W. L. &amp; Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970" yearSuffix="p">Peters &amp; Edmunds (1970, p. 177)</bibRefCitation>
.
</paragraph>
<paragraph id="BC467E0C132BFF8CE1B6F944B4726477" blockId="9.[151,1437,1357,2030]" pageId="9" pageNumber="262">
The eggs of 
<taxonomicName id="7BF9058F132BFF8CE015F944B1CF65BF" ID-CoL="8V5TD" box="[356,452,1753,1778]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE015F944B1CF65BF" box="[356,452,1753,1778]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
, here also described for the first time, resemble those of 
<taxonomicName id="7BF9058F132BFF8CE5F3F944B54865BF" ID-CoL="4S9S" box="[1154,1347,1753,1778]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE5F3F944B54865BF" box="[1154,1347,1753,1778]" italics="true" pageId="9" pageNumber="262">Habrophlebiodes</emphasis>
</taxonomicName>
by the arrangement of chorionic tubercules (see 
<bibRefCitation id="D86803FD132BFF8CE302F963B35D645B" author="Kang" box="[627,854,1790,1814]" pageId="9" pageNumber="262" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 82." type="journal article" year="1994">Kang &amp; Yang 1994</bibRefCitation>
, 
<figureCitation id="24C26289132BFF8CE212F960B3B3645B" box="[867,952,1789,1814]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="9" pageNumber="262">Fig. 17</figureCitation>
). Eggs of 
<taxonomicName id="7BF9058F132BFF8CE541F960B4CF645B" ID-CoL="5D97" box="[1072,1220,1789,1814]" class="Insecta" family="Leptophlebiidae" genus="Leptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="8E8DA21E132BFF8CE541F960B4CF645B" box="[1072,1220,1789,1814]" italics="true" pageId="9" pageNumber="262">Leptophlebia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95BE1F65132BFF8CE5BCF960B4E5645B" box="[1229,1262,1789,1814]" pageId="9" pageNumber="262" sensu="lato">s.l.</taxonomicNameLabel>
possess a very different chorionic structure (see for instance 
<bibRefCitation id="D86803FD132BFF8CE3E7F8BFB31B6477" author="Koss" box="[662,784,1825,1850]" pageId="9" pageNumber="262" refString="Koss, R. W. (1968) Morphology and taxonomic use of Ephemeroptera eggs. Annals of the Entomological Society of America, 61 (3), 696 - 721. http: // dx. doi. org / 10.1093 / aesa / 61.3.696" type="journal article" year="1968">Koss 1968</bibRefCitation>
, 
<figureCitation id="24C26289132BFF8CE26AF8BCB3976477" box="[795,924,1825,1850]" captionStart-0="FIGURES 30 – 34" captionStart-1="FIGURES 35 – 38" captionStartId-0="7.[151,264,1859,1882]" captionStartId-1="8.[151,264,897,920]" captionTargetBox-0="[151,1436,221,1837]" captionTargetBox-1="[340,1247,193,875]" captionTargetId-0="figure@7.[151,1436,193,1838]" captionTargetId-1="figure@8.[340,1248,193,876]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 30 – 34. Imagos of Gilliesia ratchaburiensis sp. n. (30) head of female imago, dorsal view; (31) forewing of female imago; (32) hindwing of female imago; (33) male genitalia, ventral view (inset in lateral view); (34) female genitalia, ventral view." captionText-1="FIGURES 35 – 38. SEM pictures of eggs extracted from a female imago of Gilliesia ratchaburiensis sp. n. (35) general outline; (36 – 37) detail of pole; (38) detail of the chorionic structure." httpUri-0="https://zenodo.org/record/237020/files/figure.png" httpUri-1="https://zenodo.org/record/237021/files/figure.png" pageId="9" pageNumber="262">Figs 30–36</figureCitation>
), without tubercles.
</paragraph>
<paragraph id="BC467E0C132BFF8FE1B6F8D8B36F6399" blockId="9.[151,1437,1357,2030]" lastBlockId="10.[151,1436,151,501]" lastPageId="10" lastPageNumber="263" pageId="9" pageNumber="262">
The genus 
<taxonomicName id="7BF9058F132BFF8CE037F8D8B1AD6413" ID-CoL="8V5TD" box="[326,422,1861,1886]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE037F8D8B1AD6413" box="[326,422,1861,1886]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
was previously known from northern 
<collectingCountry id="C4EE3E9C132BFF8CE210F8D8B3906413" box="[865,923,1861,1886]" name="India" pageId="9" pageNumber="262">India</collectingCountry>
(
<bibRefCitation id="D86803FD132BFF8CE2DDF8D8B4306413" author="Gillies" box="[940,1083,1861,1886]" pageId="9" pageNumber="262" refString="Gillies, M. T. (1951) Further notes on Ephemeroptera from India and South East Asia. Proceedings of the Royal Society of London (B), 20, 121 - 130. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1951. tb 01009. x" type="journal article" year="1951">Gillies 1951</bibRefCitation>
, 
<bibRefCitation id="D86803FD132BFF8CE538F8DBB5696413" author="Peters" box="[1097,1378,1861,1886]" pageId="9" pageNumber="262" refString="Peters, W. L. &amp; Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters &amp; Edmunds 1970</bibRefCitation>
) and mainland 
<collectingCountry id="C4EE3E9C132BFF8CE076F8F4B14164CF" box="[263,330,1897,1922]" name="China" pageId="9" pageNumber="262">China</collectingCountry>
(
<bibRefCitation id="D86803FD132BFF8CE02BF8F4B1D064CF" author="Zhou" box="[346,475,1897,1922]" pageId="9" pageNumber="262" refString="Zhou, C. - F. (2004) A new species of genus Gilliesia Peters &amp; Edmunds from China (Ephemeroptera: Leptophlebiidae). Zootaxa, 421, 1 - 4." type="journal article" year="2004">Zhou 2004</bibRefCitation>
). The discovery of a new species of this genus in 
<collectingCountry id="C4EE3E9C132BFF8CE563F8F4B47D64CF" box="[1042,1142,1897,1922]" name="Thailand" pageId="9" pageNumber="262">Thailand</collectingCountry>
extends its distribution to part of the Indo-Malayan mountain system in Southeast Asia, the locality of the new species being located in the Tenasserim Mountain Range. We may expect that some Oriental species described under the generic name 
<taxonomicName id="7BF9058F132BFF8CE1E6F848B15064A3" ID-CoL="6F8B" box="[151,347,2005,2030]" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE1E6F848B15064A3" box="[151,347,2005,2030]" italics="true" pageId="9" pageNumber="262">Paraleptophlebia</emphasis>
</taxonomicName>
at the nymphal stage only may actually be members of 
<taxonomicName id="7BF9058F132BFF8CE2B8F848B42164A3" ID-CoL="8V5TD" box="[969,1066,2005,2030]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="8E8DA21E132BFF8CE2B8F848B42164A3" box="[969,1066,2005,2030]" italics="true" pageId="9" pageNumber="262">Gilliesia</emphasis>
</taxonomicName>
. Among them, 
<taxonomicName id="7BF9058F132BFF8FE5A9F848B1B263FD" ID-CoL="4CYNP" authority="Kang &amp; Yang, 1994" authorityName="Kang &amp; Yang" authorityYear="1994" class="Insecta" family="Leptophlebiidae" genus="Paraleptophlebia" kingdom="Animalia" lastPageId="10" lastPageNumber="263" order="Ephemeroptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="spina">
<emphasis id="8E8DA21E132BFF8FE5A9F848B0DF63FD" italics="true" lastPageId="10" lastPageNumber="263" pageId="9" pageNumber="262">Paraleptophlebia spina</emphasis>
<bibRefCitation id="D86803FD1328FF8FE1ABFF05B1B263FD" author="Kang" box="[218,441,152,176]" pageId="10" pageNumber="263" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 82." type="journal article" year="1994">Kang &amp; Yang, 1994</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="C4EE3E9C1328FF8FE08DFF0AB24563FD" box="[508,590,151,176]" name="Taiwan" pageId="10" pageNumber="263">Taiwan</collectingCountry>
could be a good candidate, because of the reduced maxillary canines (
<bibRefCitation id="D86803FD1328FF8FE42FFF05B19A6399" author="Kang" pageId="10" pageNumber="263" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 82." type="journal article" year="1994">
Kang &amp; Yang 1994, 
<figureCitation id="24C262891328FF8FE04BFF21B1706399" box="[314,379,188,213]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1404,1427]" captionTargetBox="[199,1385,445,1382]" captionTargetId="figure@3.[199,1385,445,1383]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Nymphal maxillae details of Gilliesia ratchaburiensis sp. n. (6) right maxilla, ventral view; (7) left maxilla, dorsal view; (8) ventral view of maxilla; (9) SEM view of canines and dentisetae; abbreviation: c: canine, dd: distal dentiseta, pd: proximal dentiseta, ps: pectinate submarginal setae." httpUri="https://zenodo.org/record/237016/files/figure.png" pageId="10" pageNumber="263">Fig. 8</figureCitation>
D
</bibRefCitation>
) and the shape of the eggs (ibid, 
<figureCitation id="24C262891328FF8FE274FF21B35C6399" box="[773,855,188,213]" captionStart="FIGURES 14 – 22" captionStartId="5.[151,264,1953,1976]" captionTargetBox="[227,1338,217,1909]" captionTargetId="figure@5.[223,1364,193,1932]" captionTargetPageId="5" captionText="FIGURES 14 – 22. Nymph of Gilliesia ratchaburiensis sp. n. (14) labrum, dorsal view (left) and ventral view (right); (15) right mandible, ventral view; (16) left mandible, dorsal view; (17) foreleg, dorsal view; (18) foreleg, dorsal view; (19) foreleg, dorsal view; (20) forleg claw; (21) abdominal segment 8 – 10, dorsal view; (22) gill 4." httpUri="https://zenodo.org/record/237018/files/figure.png" pageId="10" pageNumber="263">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F4E32D871328FF8FE1B6FF42B28A62B9" pageId="10" pageNumber="263" type="etymology">
<paragraph id="BC467E0C1328FF8FE1B6FF42B1796250" blockId="10.[151,1436,151,501]" pageId="10" pageNumber="263">
<emphasis id="8E8DA21E1328FF8FE1B6FF42B14563B5" bold="true" box="[199,334,223,248]" pageId="10" pageNumber="263">Etymology.</emphasis>
The specific epithet is named for the province, where the 
<typeStatus id="6342C0AE1328FF8FE2AEFF7DB40463B5" box="[991,1039,224,248]" pageId="10" pageNumber="263">type</typeStatus>
material was collected: Ratchaburi province, 
<collectingCountry id="C4EE3E9C1328FF8FE076FE99B1656250" box="[263,366,260,285]" name="Thailand" pageId="10" pageNumber="263">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="BC467E0C1328FF8FE1B6FEBAB28A62B9" blockId="10.[151,1436,151,501]" pageId="10" pageNumber="263">
<emphasis id="8E8DA21E1328FF8FE1B6FEBAB182620D" bold="true" box="[199,393,295,320]" pageId="10" pageNumber="263">Biological notes.</emphasis>
The type-locality is located in a bamboo forest. The stream is characterized by a pronounced slope (25 % on the 
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) and an alternation of riffles, pools and waterfalls up to 
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high. The stream bed is mostly composed of boulders, cobbles, sand and leaf litter. The nymphs of 
<taxonomicName id="7BF9058F1328FF8FE288FEF2B51962C5" box="[1017,1298,367,392]" class="Insecta" family="Leptophlebiidae" genus="Gilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="263" phylum="Arthropoda" rank="species" species="ratchaburiensis" status="sp. nov.">
<emphasis id="8E8DA21E1328FF8FE288FEF2B51962C5" box="[1017,1298,367,392]" italics="true" pageId="10" pageNumber="263">Gilliesia ratchaburiensis</emphasis>
</taxonomicName>
<emphasis id="8E8DA21E1328FF8FE46AFEF2B55762C5" bold="true" box="[1307,1372,367,392]" pageId="10" pageNumber="263">
<taxonomicNameLabel id="95BE1F651328FF8FE46AFEF2B55762C5" box="[1307,1372,367,392]" pageId="10" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
were found in leaf bags of 
<taxonomicName id="7BF9058F1328FF8FE0FDFE09B25F62E0" box="[396,596,404,429]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="10" pageNumber="263" phylum="Tracheophyta" rank="species" species="bambos">
<emphasis id="8E8DA21E1328FF8FE0FDFE09B25F62E0" box="[396,596,404,429]" italics="true" pageId="10" pageNumber="263">Bambusa bambos</emphasis>
</taxonomicName>
and 
<taxonomicName id="7BF9058F1328FF8FE3FCFE08B3A462E0" box="[653,943,404,429]" class="Magnoliopsida" family="Lythraceae" genus="Lagerstroemia" kingdom="Plantae" order="Myrtales" pageId="10" pageNumber="263" phylum="Tracheophyta" rank="species" species="floribunda">
<emphasis id="8E8DA21E1328FF8FE3FCFE08B3A462E0" box="[653,943,404,429]" italics="true" pageId="10" pageNumber="263">Lagerstroemia floribunda</emphasis>
</taxonomicName>
in both pools and riffles habitats (
<bibRefCitation id="D86803FD1328FF8FE449FE09B156629D" author="David" pageId="10" pageNumber="263" refString="David, F. &amp; Boonsoong, B. (2014) Colonisation of leaf litter by lotic macroinvertebrates in a headwater stream of the Phachi River (western Thailand). Fundamental and Applied Limnology, 184 (2), 109 - 124. http: // dx. doi. org / 10.1127 / 1863 - 9135 / 2014 / 0596" type="journal article" year="2014">David &amp; Boonsoong 2014</bibRefCitation>
). Nymphs were found mostly together with those of 
<taxonomicName id="7BF9058F1328FF8FE2B2FE2AB597629D" authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" box="[963,1436,439,464]" class="Insecta" family="Leptophlebiidae" genus="Habrophlebiodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="263" phylum="Arthropoda" rank="species" species="prominens">
<emphasis id="8E8DA21E1328FF8FE2B2FE2AB508629D" box="[963,1283,439,464]" italics="true" pageId="10" pageNumber="263">Habrophlebiodes prominens</emphasis>
Ulmer, 1939
</taxonomicName>
and 
<taxonomicName id="7BF9058F1328FF8FE1B7FE40B27562B9" authority="Peters &amp; Edmunds, 1970" authorityName="Peters &amp; Edmunds" authorityYear="1970" box="[198,638,476,501]" class="Insecta" family="Leptophlebiidae" genus="Isca" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="263" phylum="Arthropoda" rank="species" species="janiceae">
<emphasis id="8E8DA21E1328FF8FE1B7FE40B15062B9" box="[198,347,477,500]" italics="true" pageId="10" pageNumber="263">Isca janiceae</emphasis>
<bibRefCitation id="D86803FD1328FF8FE013FE40B27562B9" author="Peters" box="[354,638,476,501]" pageId="10" pageNumber="263" refString="Peters, W. L. &amp; Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters &amp; Edmunds, 1970</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>