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<document id="E3DA002D6B9BB18BA2D4FCD616F20A30" ID-CLB-Dataset="9327" ID-DOI="10.11646/zootaxa.4894.3.4" ID-GBIF-Dataset="74322c33-5f04-4474-a9ff-0dbc181e7bfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="4315887" ID-ZooBank="451A3DBD-A70C-426B-8455-8A554DE99144" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607645634773" checkinUser="plazi" docAuthor="Oanh, Le Thi Kieu &amp; Boyko, Christopher B." docDate="2020" docId="34489947DA63CC1BFF703B7655C0B124" docLanguage="en" docName="zootaxa.4894.3.4.pdf" docOrigin="Zootaxa 4894 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stellatoniscus shieldsi Oanh &amp; Boyko 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="384" masterDocId="C871E13FDA72CC09FFE73868546FB428" masterDocTitle="Cancrion khanhensis sp. nov. (Crustacea: Isopoda: Entoniscidae) infesting Monomia haanii (Stimpson, 1858) (Crustacea: Brachyura: Portunidae) from Nha Trang Bay, Khanh Hoa, Vietnam, with remarks on larval stages of entoniscids and description of a new family, genus and two new species of hyperparasites" masterLastPageNumber="386" masterPageNumber="366" pageNumber="383" updateTime="1698847412080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A2EE8BE5B27D282687C0406A22B0EED7">Cancrion khanhensis sp. nov. (Crustacea: Isopoda: Entoniscidae) infesting Monomia haanii (Stimpson, 1858) (Crustacea: Brachyura: Portunidae) from Nha Trang Bay, Khanh Hoa, Vietnam, with remarks on larval stages of entoniscids and description of a new family, genus and two new species of hyperparasites</mods:title>
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<mods:namePart id="895B27788DA118D4E66C764285131417">Oanh, Le Thi Kieu</mods:namePart>
<mods:affiliation id="50C5451120CAB35D0944AC8BFE6A8A2E">Vietnam-Russia Tropical Centre, Nha Trang city, Khanh Hoa province, Vietnam.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="890BB6CA9A0A6CABE806A6EA0FE6C64E">Boyko, Christopher B.</mods:namePart>
<mods:affiliation id="1E6F925886E30B009870B93849469C7D">Department of Biology, Hofstra University, Hempstead, NY 11549, USA.</mods:affiliation>
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<mods:title id="D18E5821E10E03CE26F5D32F3F1530D4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="56A5743A39CEADF69838655B5C0DA5D5">
<mods:date id="191AC72B814A56B417635E088EBE1857">2020</mods:date>
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<mods:number id="24312F04B3000EE7F6C82C133CC2F6E2">2020-12-10</mods:number>
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<mods:detail id="309CCE49AFE29EB8DB3E67A323F1F61B" type="volume">
<mods:number id="A7E8FA7320119D705778E166DB411A94">4894</mods:number>
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<mods:classification id="42A727DAA730E3D8D7175AD6B71E1EED">journal article</mods:classification>
<mods:identifier id="A1558DB56456F149AA98284C5C4DFB70" type="CLB-Dataset">9327</mods:identifier>
<mods:identifier id="7E516BC86B7B81AE65D571E103E2BBED" type="DOI">10.11646/zootaxa.4894.3.4</mods:identifier>
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<treatment id="34489947DA63CC1BFF703B7655C0B124" ID-DOI="http://doi.org/10.5281/zenodo.4329026" ID-GBIF-Taxon="176084953" ID-Zenodo-Dep="4329026" LSID="urn:lsid:plazi:treatment:34489947DA63CC1BFF703B7655C0B124" httpUri="http://treatment.plazi.org/id/34489947DA63CC1BFF703B7655C0B124" lastPageId="18" lastPageNumber="384" pageId="17" pageNumber="383">
<subSubSection id="F4FB7BDADA63CC18FF703B765668B711" box="[151,519,798,825]" pageId="17" pageNumber="383" type="nomenclature">
<paragraph id="BC5E2851DA63CC18FF703B765668B711" blockId="17.[151,519,798,825]" box="[151,519,798,825]" pageId="17" pageNumber="383">
<heading id="E7169F3DDA63CC18FF703B765668B711" bold="true" box="[151,519,798,825]" fontSize="11" level="1" pageId="17" pageNumber="383" reason="1">
<emphasis id="8E95F443DA63CC18FF703B765668B711" bold="true" box="[151,519,798,825]" pageId="17" pageNumber="383">
<taxonomicName id="7BE153D2DA63CC18FF703B7655CEB711" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[151,417,798,825]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="383" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA63CC18FF703B7655CEB711" bold="true" box="[151,417,798,825]" italics="true" pageId="17" pageNumber="383">Stellatoniscus shieldsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA63CC18FE4E3B775668B711" box="[425,519,799,825]" pageId="17" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF703B0B57AFB753" box="[151,960,867,891]" pageId="17" pageNumber="383" type="description">
<paragraph id="BC5E2851DA63CC18FF703B0B57AFB753" blockId="17.[151,1377,867,922]" box="[151,960,867,891]" pageId="17" pageNumber="383">
LSID
<uri id="C8702453DA63CC18FF313B0B57AFB753" box="[214,960,867,891]" pageId="17" pageNumber="383">
urn:lsid:zoobank.org:act:
<uuid id="C8471284DA63CC18FE283B0B57AFB753" box="[463,960,867,891]" pageId="17" pageNumber="383">C3DF7EB6-2762-4A8A-810F-9B3BDE28D380</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF703BEA510EB7B2" box="[151,1377,898,922]" pageId="17" pageNumber="383" type="reference_group">
<paragraph id="BC5E2851DA63CC18FF703BEA510EB7B2" blockId="17.[151,1377,867,922]" box="[151,1377,898,922]" pageId="17" pageNumber="383">
<treatmentCitationGroup id="9CF10F7FDA63CC18FF703BEA510EB7B2" box="[151,1377,898,922]" pageId="17" pageNumber="383">
<treatmentCitation id="3D400E40DA63CC18FF703BEA56ACB7B2" author="Shields, J. D. &amp; Ward, L. A." box="[151,707,898,922]" page="594" pageId="17" pageNumber="383" year="1998">
<taxonomicName id="7BE153D2DA63CC18FF703BEA56ACB7B2" ID-CoL="7BYRC" authority="Shields &amp; Ward, 1998: 591 - 594" authorityName="Shields &amp; Ward" authorityPageNumber="591 - 594" authorityYear="1998" box="[151,707,898,922]" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="383" phylum="Arthropoda" rank="species" species="texopallium">
<emphasis id="8E95F443DA63CC18FF703BEA551BB7B2" box="[151,372,898,922]" italics="true" pageId="17" pageNumber="383">Tiarinion texopallium</emphasis>
<bibRefCitation id="D87055A0DA63CC18FE9D3BEA56ACB7B2" author="Shields, J. D. &amp; Ward, L. A." box="[378,707,898,922]" pageId="17" pageNumber="383" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward, 1998: 591–594</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 4.—
<treatmentCitation id="3D400E40DA63CC18FCF83BEA5066B7B2" author="Poore, G. C. B. &amp; Markham, J. C. &amp; Lew Tan, H. M." box="[799,1033,898,922]" page="132" pageId="17" pageNumber="383" year="2002">
<bibRefCitation id="D87055A0DA63CC18FCF83BEA5066B7B2" author="Poore, G. C. B. &amp; Markham, J. C. &amp; Lew Tan, H. M." box="[799,1033,898,922]" pageId="17" pageNumber="383" pagination="111 - 137" refId="ref13656" refString="Poore, G. C. B., Markham, J. C. &amp; Lew Tan, H. M. (2002) Superfamily: Bopyroidea. In: Houston, W. W. K. &amp; Beesley, P. L. (Eds.), Zoological Catalogue of Australia. Vol. 19.2 A. Crustacea: Malacostraca: Syncarida, Peracarida: Isopoda, Tanaidacea, Mictacea, Thermosbaenacea, Spelaeogriphacea. Vol. 19.2 A. CSIRO Publishing, Melbourne, pp. 111 - 137." type="book chapter" year="2002">
Poore
<emphasis id="8E95F443DA63CC18FC843BEB57FFB7B2" box="[867,912,898,922]" italics="true" pageId="17" pageNumber="383">et al</emphasis>
., 2002: 132
</bibRefCitation>
</treatmentCitation>
(in part, cryptoniscus larva only).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF703BA05610B066" pageId="17" pageNumber="383" type="materials_examined">
<paragraph id="BC5E2851DA63CC18FF703BA05610B066" blockId="17.[151,1437,967,2039]" pageId="17" pageNumber="383">
<materialsCitation id="0C89220CDA63CC18FF703BA05610B066" ID-GBIF-Occurrence="2995018308" collectingDate="1992-03-01" collectionCode="QM" collectorName="J. D. Shields" country="Australia" elevation="4" location="Shields &amp; Ward" pageId="17" pageNumber="383" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="8E95F443DA63CC18FF703BA055DAB7CA" bold="true" box="[151,437,967,994]" pageId="17" pageNumber="383">
Type material.
<emphasis id="8E95F443DA63CC18FEAA3BAF55DAB7CA" bold="true" box="[333,437,967,994]" italics="true" pageId="17" pageNumber="383">
<typeStatus id="635A96F3DA63CC18FEAA3BAF55DAB7CA" box="[333,437,967,994]" pageId="17" pageNumber="383" type="holotype">Holotype</typeStatus>
</emphasis>
</emphasis>
cryptoniscus larva,
<quantity id="7B1985B4DA63CC18FD7F3BA0569EB7CB" box="[664,753,968,995]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="383" unit="mm" value="800.0">800 µm</quantity>
, ex mature holotype female
<taxonomicName id="7BE153D2DA63CC18FBD63BA15577B02E" authority="Shields &amp; Ward, 1998" authorityName="Shields &amp; Ward" authorityYear="1998" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="383" phylum="Arthropoda" rank="species" species="texopallium">
<emphasis id="8E95F443DA63CC18FBD63BA1514AB7CA" box="[1073,1317,968,994]" italics="true" pageId="17" pageNumber="383">Tiarinion texopallium</emphasis>
<bibRefCitation id="D87055A0DA63CC18FACA3BA05577B02E" author="Shields, J. D. &amp; Ward, L. A." pageId="17" pageNumber="383" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward, 1998</bibRefCitation>
</taxonomicName>
(QM W222015),
<quantity id="7B1985B4DA63CC18FE383B84562BB02F" box="[479,580,1004,1031]" metricMagnitude="-3" metricUnit="m" metricValue="7.59" pageId="17" pageNumber="383" unit="mm" value="7.59">7.59 mm</quantity>
, ex female
<taxonomicName id="7BE153D2DA63CC18FD253B855026B02F" authority="(Latreille 1825)" baseAuthorityName="Latreille" baseAuthorityYear="1825" box="[706,1097,1004,1031]" class="Malacostraca" family="Majidae" genus="Tiarinia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="383" phylum="Arthropoda" rank="species" species="cornigera">
<emphasis id="8E95F443DA63CC18FD253B8557FFB02E" box="[706,912,1005,1030]" italics="true" pageId="17" pageNumber="383">Tiarinia cornigera</emphasis>
(
<bibRefCitation id="D87055A0DA63CC18FC793B84502FB02F" author="Latreille, P. A." box="[926,1088,1004,1031]" pageId="17" pageNumber="383" refId="ref13319" refString="Latreille, P. A. (1825) Familles Naturelles du Regne Animal, Expose Succinctement et dans un Ordre Analytique avec l'indication de Leurs Genres. J. B. Bailliere, Paris, 570 pp. https: // doi. org / 10.5962 / bhl. title. 16094" type="book" year="1825">Latreille 1825</bibRefCitation>
)
</taxonomicName>
,
<quantity id="7B1985B4DA63CC18FBB33B8450D8B02F" box="[1108,1207,1004,1031]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="17" pageNumber="383" unit="mm" value="10.8">10.8 mm</quantity>
CL, Mermaid Cove, Lizard Island,
<collectingRegion id="7E25E6B3DA63CC18FEDE3C7855AFB002" box="[313,448,1040,1066]" country="Australia" name="Queensland" pageId="17" pageNumber="383">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA63CC18FE2D3C78565BB002" box="[458,564,1040,1066]" name="Australia" pageId="17" pageNumber="383">Australia</collectingCountry>
,
<quantity id="7B1985B4DA63CC18FDD83C7856E8B003" box="[575,647,1040,1067]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="17" pageNumber="383" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<elevation id="37CCCF62DA63CC18FDD83C7856E8B003" box="[575,647,1040,1067]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="17" pageNumber="383" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</elevation>
</quantity>
, 14.40’S, 145.28’E,
<date id="C85F0E91DA63CC18FC893C78579BB002" box="[878,1012,1040,1067]" pageId="17" pageNumber="383" value="1992-03-01">
<collectingDate id="D81BF779DA63CC18FC893C78579BB002" box="[878,1012,1040,1067]" pageId="17" pageNumber="383" value="1992-03-01">1 Mar 1992</collectingDate>
</date>
, coll.
<collectorName id="11144D87DA63CC18FBD03C7950ABB002" box="[1079,1220,1040,1067]" pageId="17" pageNumber="383">J. D. Shields</collectorName>
(
<collectionCode id="DAF0B094DA63CC18FB343C78516DB002" box="[1235,1282,1040,1066]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="17" pageNumber="383" type="Museum">QM</collectionCode>
W22272 [not W22016 as cited in
<bibRefCitation id="D87055A0DA63CC18FE903C5C561AB066" author="Shields, J. D. &amp; Ward, L. A." box="[375,629,1075,1102]" pageId="17" pageNumber="383" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">
<location id="B93E7E8ADA63CC18FE903C5C5643B066" LSID="urn:lsid:plazi:treatment:34489947DA63CC1BFF703B7655C0B124:B93E7E8ADA63CC18FE903C5C5643B066" box="[375,556,1076,1102]" country="Australia" name="Shields &amp; Ward" pageId="17" pageNumber="383" stateProvince="Queensland">Shields &amp; Ward</location>
, 1998
</bibRefCitation>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF203C30554BB0BE" pageId="17" pageNumber="383" type="etymology">
<paragraph id="BC5E2851DA63CC18FF203C30554BB0BE" blockId="17.[151,1437,967,2039]" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203C305521B05A" bold="true" box="[199,334,1112,1138]" pageId="17" pageNumber="383">Etymology.</emphasis>
The new species name is given after Jeffrey D. Shields in honor of his many years of parasitological research.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF203CC85030B387" pageId="17" pageNumber="383" type="description">
<paragraph id="BC5E2851DA63CC18FF203CC850A4B1BA" blockId="17.[151,1437,967,2039]" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203CC85650B092" bold="true" box="[199,575,1183,1210]" pageId="17" pageNumber="383">
Description.
<emphasis id="8E95F443DA63CC18FE843CC85650B092" bold="true" box="[355,575,1183,1210]" italics="true" pageId="17" pageNumber="383">Cryptoniscus larva</emphasis>
</emphasis>
body fusiform, strongly convex dorsally, length
<quantity id="7B1985B4DA63CC18FB913CC850BDB093" box="[1142,1234,1184,1211]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="383" unit="mm" value="800.0">800 µm</quantity>
, maximum width
<quantity id="7B1985B4DA63CC18FF703CAC549BB0F7" box="[151,244,1220,1247]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="383" unit="mm" value="125.0">125 µm</quantity>
.
<emphasis id="8E95F443DA63CC18FEE63CAC5501B0F6" box="[257,366,1220,1246]" italics="true" pageId="17" pageNumber="383">Cephalon</emphasis>
anterior margin round; eyes positioned dorsolaterally.
<emphasis id="8E95F443DA63CC18FC013CAC504FB0F6" box="[998,1056,1220,1246]" italics="true" pageId="17" pageNumber="383">Body</emphasis>
pigmentation lacking.
<emphasis id="8E95F443DA63CC18FACE3CAC51F3B0F6" box="[1321,1436,1220,1246]" italics="true" pageId="17" pageNumber="383">Antennula</emphasis>
expanded distolaterally, mesially approximated, covering almost entire ventral cephalon; first segment flattened, broad, 6 distal, rounded to subacute projections; proximomesial area demarcated, smooth; proximal margin with single spine and 2 short setae slightly distally; second segment cylindrical, projecting distolaterally, smooth; exopodite short, cylindrical, endopodite longer, distally tapered, epipodite short, subtriangular, distally extended, all three segments with numerous long setae.
<emphasis id="8E95F443DA63CC18FDD43D1056FFB1BA" box="[563,656,1400,1426]" italics="true" pageId="17" pageNumber="383">Antenna</emphasis>
with 4 peduncular and 5 flagellar articles, smooth.
</paragraph>
<paragraph id="BC5E2851DA63CC18FF203DF555D4B336" blockId="17.[151,1437,967,2039]" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203DF55578B19E" box="[199,279,1437,1462]" italics="true" pageId="17" pageNumber="383">Pereon</emphasis>
with at least some coxal plates covered with scales (see Remarks).
<emphasis id="8E95F443DA63CC18FBFF3DF45149B19E" box="[1048,1318,1436,1462]" italics="true" pageId="17" pageNumber="383">Pereopods 1, 2 and 3–7</emphasis>
dissimilar in size and shape. Pereopod 1 stout, dactylus approximately propodus length, with scale-like spinules on outer surface, propodus rounded, carpus with area of ridges ventrally, merus mostly smooth with few ridges distoventrally, ischium smooth, basis smooth. Pereopod 2 larger than 1, dactylus reaching to distal margin of merus, surface smooth; propodus smooth with 1 spine-like seta on ventral margin; carpus smooth with 1 distoventral spine-like seta. Pereopods 3–7 subequal in size and shape, much slenderer than 1 and 2; dactylus smooth, recurved, approximately half length of propodus, those of pereopods 6 and 7 with small setae on dorsal and ventral surface; propodus slender, slightly medially inflated, 2 stout spine-like setae on distoventral surface; carpus subtriangular with single distoventral spine-like seta on at least pereopod 6; merus approximately twice as long as carpus, distally expanded with 1 distoventral seta and 1 distodorsal seta; ischium and basis subequal in size and shape, dorsomedial surface expanded into broad lobe.
</paragraph>
<paragraph id="BC5E2851DA63CC18FF203F405030B387" blockId="17.[151,1437,967,2039]" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203F405567B36A" box="[199,264,1832,1858]" italics="true" pageId="17" pageNumber="383">Pleon</emphasis>
ventrally smooth; five pairs of biramous subequal pleopods; sympod subquadrate, distomesial corner with 2 long setae; exopods and endopods short, subquadrate, each with 5 long terminal setae. Pleotelson rounded.
<emphasis id="8E95F443DA63CC18FF703F185497B3A2" box="[151,248,1904,1930]" italics="true" pageId="17" pageNumber="383">Uropods</emphasis>
biramous, composed of subquadrate sympod with single long seta at distolateral corner, endopod nearly twice as long as exopod, exopod with 2 long distal setae and pair of lateral short setae.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC18FF203FD05732B3FB" box="[199,861,1976,2003]" pageId="17" pageNumber="383" type="distribution">
<paragraph id="BC5E2851DA63CC18FF203FD05732B3FB" blockId="17.[151,1437,967,2039]" box="[199,861,1976,2003]" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203FD0550EB3FA" bold="true" box="[199,353,1976,2002]" pageId="17" pageNumber="383">Distribution.</emphasis>
Lizard Island,
<collectingRegion id="7E25E6B3DA63CC18FDED3FD056FEB3FA" box="[522,657,1976,2002]" country="Australia" name="Queensland" pageId="17" pageNumber="383">Queensland</collectingRegion>
,
<collectingCountry id="C4F668C1DA63CC18FD7C3FD0576AB3FA" box="[667,773,1976,2002]" name="Australia" pageId="17" pageNumber="383">Australia</collectingCountry>
,
<quantity id="7B1985B4DA63CC18FCE83FD05736B3FB" box="[783,857,1976,2003]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="17" pageNumber="383" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
3–
<specimenCount id="AAE7E3D8DA63CC18FCCB3FD05736B3FB" box="[812,857,1976,2003]" pageId="17" pageNumber="383" type="generic">5 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F4FB7BDADA63CC1BFF203FB455C0B124" lastPageId="18" lastPageNumber="384" pageId="17" pageNumber="383" type="discussion">
<paragraph id="BC5E2851DA63CC1BFF203FB4579BB070" blockId="17.[151,1437,967,2039]" lastBlockId="18.[151,1437,150,1292]" lastPageId="18" lastPageNumber="384" pageId="17" pageNumber="383">
<emphasis id="8E95F443DA63CC18FF203FB45554B3DE" bold="true" box="[199,315,2012,2038]" pageId="17" pageNumber="383">Remarks.</emphasis>
The cryptoniscus larva now designated as the
<typeStatus id="635A96F3DA63CC18FCA23FB457C8B3DE" box="[837,935,2012,2038]" pageId="17" pageNumber="383" type="holotype">holotype</typeStatus>
of
<taxonomicName id="7BE153D2DA63CC18FC2B3FB450AAB3DE" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[972,1221,2012,2038]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="383" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA63CC18FC2B3FB450AAB3DE" box="[972,1221,2012,2038]" italics="true" pageId="17" pageNumber="383">Stellatoniscus shieldsi</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA63CC18FB2C3FB4514AB3DE" bold="true" box="[1227,1317,2012,2038]" pageId="17" pageNumber="383">
<taxonomicNameLabel id="95A64938DA63CC18FB2C3FB4514AB3DE" box="[1227,1317,2012,2038]" pageId="17" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was originally described as one of two
<typeStatus id="635A96F3DA60CC1BFE0338FF562BB499" box="[484,580,151,177]" pageId="18" pageNumber="384" type="paratype">paratype</typeStatus>
cryptoniscus larvae of
<taxonomicName id="7BE153D2DA60CC1BFCAF38FF5153B498" authority="Shields &amp; Ward, 1998" authorityName="Shields &amp; Ward" authorityYear="1998" box="[840,1340,150,176]" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="texopallium">
<emphasis id="8E95F443DA60CC1BFCAF38FF5054B498" box="[840,1083,150,176]" italics="true" pageId="18" pageNumber="384">Tiarinion texopallium</emphasis>
<bibRefCitation id="D87055A0DA60CC1BFBA538FE5153B498" author="Shields, J. D. &amp; Ward, L. A." box="[1090,1340,150,176]" pageId="18" pageNumber="384" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward, 1998</bibRefCitation>
</taxonomicName>
. Despite the description being incomplete and the illustrations containing numerous errors (see below), it is clear from the biramous condition of the pleopods that this is a cryptoniscoid cryptoniscus, not a bopyroid cryptoniscus and was therefore incorrectly considered a larval stage of
<taxonomicName id="7BE153D2DA60CC1BFD35396B5718B534" authorityName="Shields &amp; Ward" authorityYear="1998" box="[722,887,258,285]" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="texopallium">
<emphasis id="8E95F443DA60CC1BFD35396B568DB534" box="[722,738,259,284]" italics="true" pageId="18" pageNumber="384">T</emphasis>
.
<emphasis id="8E95F443DA60CC1BFD14396A5718B534" box="[755,887,258,284]" italics="true" pageId="18" pageNumber="384">texopallium</emphasis>
</taxonomicName>
. However, until the discovery of
<taxonomicName id="7BE153D2DA60CC1BFAE6396A557CB568" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="khanhensis" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFAE6396A557CB568" italics="true" pageId="18" pageNumber="384">Stellatoniscus khanhensis</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFEFB394E5518B568" bold="true" box="[284,375,294,320]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFEFB394E5518B568" box="[284,375,294,320]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the affinities of this larva could not be determined. It is clear that the cryptoniscus of
<taxonomicName id="7BE153D2DA60CC1BFAB4394E5503B54C" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFAB4394E5503B54C" italics="true" pageId="18" pageNumber="384">Stellatoniscus tentaculus</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFE95392255A5B54C" bold="true" box="[370,458,330,356]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFE95392255A5B54C" box="[370,458,330,356]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="7BE153D2DA60CC1BFE1939235601B54C" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[510,622,330,357]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFE1939235663B54D" box="[510,524,331,357]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFDFF39225601B54C" box="[536,622,330,356]" italics="true" pageId="18" pageNumber="384">shieldsi</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFD93392256A3B54C" bold="true" box="[628,716,330,356]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFD93392256A3B54C" box="[628,716,330,356]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share many characters in common and belong to the same genus. These characters include the shape of the antennular plate (expanded with demarcated mesial region), pereopod 1 with ridges on dactylus, carpus and merus, pereopod 2 larger than pereopod 1 and morphologically different (no ridges, much larger dactylus, tapered propodus), pereopods 3–7 different from 1 and 2 (proportionally shorter dactylus than pereopod 2, scarcely recurved, ischium and basis with dorsomedial expansion), scales on coxal plates. None of these characters are known from any other cryptoniscoid larvae. However, it needs to be pointed out that the description of the cryptoniscus larva from
<taxonomicName id="7BE153D2DA60CC1BFD463A4B57FAB614" box="[673,917,546,572]" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="texapallium">
<emphasis id="8E95F443DA60CC1BFD463A4B57FAB614" box="[673,917,546,572]" italics="true" pageId="18" pageNumber="384">Tiarinion texapallium</emphasis>
</taxonomicName>
is incomplete in several characters and the illustrations are wrong or misleading (based on examination of the badly damaged specimen). Errors include: there are only 2 distal antennular segments not 3 as drawn (
<bibRefCitation id="D87055A0DA60CC1BFD023A0257B7B6AC" author="Shields, J. D. &amp; Ward, L. A." box="[741,984,618,644]" pageId="18" pageNumber="384" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward 1998</bibRefCitation>
: fig. 4C), the peduncular segments of the antennae are very much like those seen in
<taxonomicName id="7BE153D2DA60CC1BFD893AE75693B680" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[622,764,654,681]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFD893AE75613B681" box="[622,636,655,681]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFD6E3AE65693B680" box="[649,764,654,680]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFCE43AE657BDB680" box="[771,978,654,681]" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFCE43AE65704B680" bold="true" box="[771,875,654,680]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFC9E3AE657BDB680" bold="true" box="[889,978,654,680]" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
and not as drawn (
<bibRefCitation id="D87055A0DA60CC1BFB433AE651F6B680" author="Shields, J. D. &amp; Ward, L. A." box="[1188,1433,654,680]" pageId="18" pageNumber="384" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward 1998</bibRefCitation>
: fig. 4B), pereopod 1 is much larger compared to pereopod 2, nearly the same size as the first 4 antennal peduncular segments, than drawn (
<bibRefCitation id="D87055A0DA60CC1BFE7A3ABE56F8B6D8" author="Shields, J. D. &amp; Ward, L. A." box="[413,663,726,752]" pageId="18" pageNumber="384" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward 1998</bibRefCitation>
: fig. 1A). Unfortunately, the examined specimen is so damaged that the coxal plates of the pereomeres cannot be discerned and pereomeres 2 and 3 are completely missing. Despite this, the
<typeStatus id="635A96F3DA60CC1BFF263B76554CB710" box="[193,291,798,824]" pageId="18" pageNumber="384" type="holotype">holotype</typeStatus>
of
<taxonomicName id="7BE153D2DA60CC1BFEAD3B7755D2B710" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[330,445,798,825]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFEAD3B775537B711" box="[330,344,799,825]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFE803B7655D2B710" box="[359,445,798,824]" italics="true" pageId="18" pageNumber="384">shieldsi</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFE233B765670B710" bold="true" box="[452,543,798,824]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFE233B765670B710" box="[452,543,798,824]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the cryptoniscus of
<taxonomicName id="7BE153D2DA60CC1BFBD63B7750AEB710" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1073,1217,798,825]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFBD63B775050B711" box="[1073,1087,799,825]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFBA93B7650AEB710" box="[1102,1217,798,824]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFB2E3B7651F3B710" box="[1225,1436,798,825]" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFB2E3B76515CB710" bold="true" box="[1225,1331,798,824]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFAA53B7651F3B710" bold="true" box="[1346,1436,798,824]" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
by the following characters: antennular plate mesial demarcated region smooth, general shape rounded, and with 7+ distal teeth vs. with horizontal ridges, general shape angular, and with 7 distal teeth in
<taxonomicName id="7BE153D2DA60CC1BFB8C3B0F5095B7A8" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1131,1274,870,897]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFB8C3B0F5016B7A9" box="[1131,1145,871,897]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFB603B0E5095B7A8" box="[1159,1274,870,896]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFAE53B0E54A7B78C" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFAE53B0E5103B7A8" bold="true" box="[1282,1388,870,896]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFA9C3B0E54A7B78C" bold="true" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
; coxal plates with small region of scales vs. covered with scales in
<taxonomicName id="7BE153D2DA60CC1BFC253BE3503EB78C" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[962,1105,906,933]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFC253BE357BFB78D" box="[962,976,907,933]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFC393BE2503EB78C" box="[990,1105,906,932]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFBBF3BE25145B78C" box="[1112,1322,906,933]" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFBBF3BE250ADB78C" bold="true" box="[1112,1218,906,932]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFB373BE25145B78C" bold="true" box="[1232,1322,906,932]" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
, posterior pereopods with weakly recurved dactylus vs. strongly recurved dactylus in
<taxonomicName id="7BE153D2DA60CC1BFC103BC750E7B7E0" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1015,1160,942,969]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFC103BC7506AB7E1" box="[1015,1029,943,969]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFBF23BC650E7B7E0" box="[1045,1160,942,968]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFB763BC65107B7E0" box="[1169,1384,942,969]" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFB763BC65092B7E0" bold="true" box="[1169,1277,942,968]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFAEA3BC65107B7E0" bold="true" box="[1293,1384,942,968]" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
The sizes of the larvae are also very different:
<quantity id="7B1985B4DA60CC1BFD843BBA5686B7C5" box="[611,745,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="18" pageNumber="384" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
for
<taxonomicName id="7BE153D2DA60CC1BFCF23BBB57CCB7C4" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[789,931,978,1005]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="tentaculus" status="gen. nov., sp. nov.">
<emphasis id="8E95F443DA60CC1BFCF23BBB574CB7C5" box="[789,803,979,1005]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFCD73BBA57CCB7C4" box="[816,931,978,1004]" italics="true" pageId="18" pageNumber="384">tentaculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95A64938DA60CC1BFC4E3BBA5019B7C4" box="[937,1142,978,1005]" pageId="18" pageNumber="384" rank="species">
<emphasis id="8E95F443DA60CC1BFC4E3BBA507FB7C4" bold="true" box="[937,1040,978,1004]" pageId="18" pageNumber="384">gen. nov.</emphasis>
,
<emphasis id="8E95F443DA60CC1BFBFA3BBA5019B7C4" bold="true" box="[1053,1142,978,1004]" pageId="18" pageNumber="384">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<quantity id="7B1985B4DA60CC1BFB4D3BBA516FB7C5" box="[1194,1280,978,1005]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="384" unit="mm" value="800.0">800 µm</quantity>
for
<taxonomicName id="7BE153D2DA60CC1BFACB3BBB51F2B7C4" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[1324,1437,978,1005]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFACB3BBB51F2B7C4" box="[1324,1437,978,1005]" italics="true" pageId="18" pageNumber="384">S. shieldsi</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFF703B9E549DB038" bold="true" box="[151,242,1014,1040]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFF703B9E549DB038" box="[151,242,1014,1040]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
It should be noted that we were unable to examine the other cryptoniscus larva cited by
<bibRefCitation id="D87055A0DA60CC1BFB0F3B9E548DB01C" author="Shields, J. D. &amp; Ward, L. A." pageId="18" pageNumber="384" pagination="590 - 596" refId="ref14082" refString="Shields, J. D. &amp; Ward, L. A. (1998) Tiarinion texopallium, new species, an entoniscid isopod infesting majid crabs (Tiarinia spp.) from the Great Barrier Reef, Australia. Journal of Crustacean Biology, 18 (3), 590 - 596. https: // doi. org / 10.2307 / 1549423" type="journal article" year="1998">Shields &amp; Ward (1998)</bibRefCitation>
due to the onset of the COVID-19 pandemic; this specimen (USNM 284171) should be examined to confirm its conspecificity with the larva designated as
<typeStatus id="635A96F3DA60CC1BFD7C3C565692B070" box="[667,765,1086,1112]" pageId="18" pageNumber="384" type="holotype">holotype</typeStatus>
of
<taxonomicName id="7BE153D2DA60CC1BFCC53C5757FBB070" authority="Oanh &amp; Boyko, 2020" authorityName="Oanh &amp; Boyko" authorityYear="2020" box="[802,916,1086,1113]" class="Malacostraca" family="Entoniscidae" genus="Stellatoniscus" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="shieldsi" status="sp. nov.">
<emphasis id="8E95F443DA60CC1BFCC53C57575FB071" box="[802,816,1087,1113]" italics="true" pageId="18" pageNumber="384">S</emphasis>
.
<emphasis id="8E95F443DA60CC1BFCD93C5657FBB070" box="[830,916,1086,1112]" italics="true" pageId="18" pageNumber="384">shieldsi</emphasis>
</taxonomicName>
<emphasis id="8E95F443DA60CC1BFC7C3C56579BB070" bold="true" box="[923,1012,1086,1112]" pageId="18" pageNumber="384">
<taxonomicNameLabel id="95A64938DA60CC1BFC7C3C56579BB070" box="[923,1012,1086,1112]" pageId="18" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="BC5E2851DA60CC1BFF203C0A55C0B124" blockId="18.[151,1437,150,1292]" pageId="18" pageNumber="384">
The description and illustrations of the female, male, and epicaridium larva of
<taxonomicName id="7BE153D2DA60CC1BFBCB3C0B50A3B054" authorityName="Shields &amp; Ward" authorityYear="1998" box="[1068,1228,1122,1149]" class="Malacostraca" family="Entoniscidae" genus="Tiarinion" kingdom="Animalia" order="Isopoda" pageId="18" pageNumber="384" phylum="Arthropoda" rank="species" species="texopallium">
<emphasis id="8E95F443DA60CC1BFBCB3C0B5053B054" box="[1068,1084,1123,1148]" italics="true" pageId="18" pageNumber="384">T</emphasis>
.
<emphasis id="8E95F443DA60CC1BFBAF3C0A50A3B054" box="[1096,1228,1122,1148]" italics="true" pageId="18" pageNumber="384">texopallium</emphasis>
</taxonomicName>
are reasonably accurate except that the epicaridium larva was described and illustrated as having biramous pleopods when it actually has uniramous pleopods as do all other epicaridium larvae of entoniscids described to date. There are two segments present on each pleopod but they are the sympod and exopod (the endopod is lacking) and this is considered to be the uniramous condition.
</paragraph>
</subSubSection>
</treatment>
</document>
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