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<document id="4A404699C7B025150044A3D897866297" ID-CLB-Dataset="21808" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C16AFF801C05F2A1FE03FC35" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus Attems 1914" docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="8" updateTime="1698841637397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B74442D0CA810AB8841F6137B2EAB266">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<treatment id="334F2769C16AFF801C05F2A1FE03FC35" ID-DOI="http://doi.org/10.5281/zenodo.3861187" ID-GBIF-Taxon="164362765" ID-Zenodo-Dep="3861187" LSID="urn:lsid:plazi:treatment:334F2769C16AFF801C05F2A1FE03FC35" httpUri="http://treatment.plazi.org/id/334F2769C16AFF801C05F2A1FE03FC35" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="F3FCC5F4C16AFF8F1C05F2A1FC42FC13" box="[594,994,918,944]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="BB59967FC16AFF8F1C05F2A1FC42FC13" blockId="7.[594,994,918,944]" box="[594,994,918,944]" pageId="7" pageNumber="8">
<heading id="E0112113C16AFF8F1C05F2A1FC42FC13" box="[594,994,918,944]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="7CE6EDFCC16AFF8F1CF3F2A1FC42FC13" ID-CoL="7NXRH" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[676,994,918,944]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1CF3F2A1FC9DFC13" box="[676,829,918,944]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
Attems, 1914
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C16AFF801EEAF2E0FE03FC35" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph id="BB59967FC16AFF8F1EEAF2E0FC35FBB6" blockId="7.[189,1399,983,1045]" pageId="7" pageNumber="8">
This genus was established by Attems (1914) for three new species, of which
<taxonomicName id="7CE6EDFCC16AFF8F1A11F2EFFAD1FC52" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[1094,1393,983,1010]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="89924A6DC16AFF8F1A11F2EFFB6CFC52" box="[1094,1228,983,1010]" italics="true" pageId="7" pageNumber="8">C. limbatus</emphasis>
Attems, 1914
</taxonomicName>
, was subsequently designated as
<typeStatus id="645D28DDC16AFF8F1C60F2CCFDCAFBB6" box="[567,618,1019,1045]" pageId="7" pageNumber="8">type</typeStatus>
species by
<bibRefCitation id="DF77EB8EC16AFF8F1CA5F2CCFC31FBB6" author="Kraus O." box="[754,913,1018,1045]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960)</bibRefCitation>
.
</paragraph>
<paragraph id="BB59967FC16AFF8F1EEAF50BFD43FB41" blockId="7.[189,1399,1084,1250]" pageId="7" pageNumber="8">
<taxonomicName id="7CE6EDFCC16AFF8F1EEAF50BFEF6FBF5" authorityName="Attems" authorityYear="1914" box="[189,342,1084,1110]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1EEAF50BFEF6FBF5" box="[189,342,1084,1110]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
</taxonomicName>
was treated in detail by
<bibRefCitation id="DF77EB8EC16AFF8F1CD5F50BFCB9FBF5" author="Kraus O." box="[642,793,1084,1110]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960</bibRefCitation>
,
<bibRefCitation id="DF77EB8EC16AFF8F1D70F50BFCC3FBF4" author="Kraus O." box="[807,867,1084,1111]" pageId="7" pageNumber="8" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">1966</bibRefCitation>
) and was recently addressed by
<bibRefCitation id="DF77EB8EC16AFF8F1ABBF50BFEBCFBDA" author="Frederiksen S. B." pageId="7" pageNumber="8" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen (2013a)</bibRefCitation>
.
<bibRefCitation id="DF77EB8EC16AFF8F1F72F568FDEDFBDA" author="Vohland K. &amp; Hamer M." box="[293,589,1119,1145]" pageId="7" pageNumber="8" pagination="251 - 304" refId="ref31257" refString="Vohland K. &amp; Hamer M. 2013. A review of the millipedes (Diplopoda) of Namibia, with identification keys and descriptions of two new genera and five new species. African Invertebrates 54: 251 - 304." type="journal article" year="2013">Vohland &amp; Hamer (2013)</bibRefCitation>
redescribed two
<taxonomicName id="7CE6EDFCC16AFF8F1D44F568FC0CFBDA" authorityName="Attems" authorityYear="1914" box="[787,940,1119,1145]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1D44F568FC0CFBDA" box="[787,940,1119,1145]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
</taxonomicName>
species from
<collectingCountry id="C3F1D6EFC16AFF8F1A1AF568FB16FBDA" box="[1101,1206,1119,1145]" name="Namibia" pageId="7" pageNumber="8">Namibia</collectingCountry>
. From the works of these authors the following diagnosis of
<taxonomicName id="7CE6EDFCC16AFF8F1C92F5B5FCFEFB3F" authorityName="Attems" authorityYear="1914" box="[709,862,1154,1180]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1C92F5B5FCFEFB3F" box="[709,862,1154,1180]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
</taxonomicName>
can be extracted, based entirely on gonopod structure, using the terminology suggested by
<bibRefCitation id="DF77EB8EC16AFF8F1C89F592FC67FB1C" author="Frederiksen S. B." box="[734,967,1189,1215]" pageId="7" pageNumber="8" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen (2013a</bibRefCitation>
,b;
<bibRefCitation id="DF77EB8EC16AFF8F1DB0F592FA9CFB1C" author="Frederiksen S. B. &amp; Enghoff H." box="[999,1340,1189,1215]" pageId="7" pageNumber="8" pagination="69 - 82" refId="ref30445" refString="Frederiksen S. B. &amp; Enghoff H. 2012. East African odontopygid millipedes 1: Five new species of Xystopyge (Attems, 1909) and a proposal for a new gonopod terminology (Diplopoda; Spirostreptida; Odontopygidae). Zootaxa 3485: 69 - 82." type="journal article" year="2012">Frederiksen &amp; Enghoff 2012</bibRefCitation>
) and emended in light of the species described here:
</paragraph>
<paragraph id="BB59967FC16AFF8F1EEAF43DFE71FA87" blockId="7.[189,1399,1290,1562]" box="[189,465,1290,1316]" pageId="7" pageNumber="8">
<taxonomicName id="7CE6EDFCC16AFF8F1EEAF43DFEC3FA87" authorityName="Hoffman" authorityYear="1991" box="[189,355,1290,1316]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Prionopetalini">Prionopetalini</taxonomicName>
in which
</paragraph>
<paragraph id="BB59967FC16AFF8F1EEAF419FE42FAC9" blockId="7.[189,1399,1290,1562]" pageId="7" pageNumber="8">• the proplica and metaplica of the gonopod coxa come together apically and form a hoodlike structure, here termed cucullus</paragraph>
<paragraph id="BB59967FC16AFF8F1EEAF443FB36FA76" blockId="7.[189,1399,1290,1562]" pageId="7" pageNumber="8">• the solenomere is very long, whiplike, at least twice as long as the telomere if stretched out, not spiralled terminally, and without accessory branches or outgrowths (except for sometimes at the very base, corresponding to the “Tibialdorn” of Kraus; see species descriptions)</paragraph>
<paragraph id="BB59967FC16AFF8F1EEAF4E9FE67F9B9" blockId="7.[189,1399,1290,1562]" pageId="7" pageNumber="8">• the telomere is proximally folded like a tube or a trough and distally separates into two or three diverging lamellae</paragraph>
<paragraph id="BB59967FC16AFF801EEAF775FDEBFEEA" blockId="7.[189,1399,1602,2016]" lastBlockId="8.[189,1398,267,329]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<bibRefCitation id="DF77EB8EC16AFF8F1EEAF775FEF5F9FF" author="Kraus O." box="[189,341,1602,1628]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960</bibRefCitation>
,
<bibRefCitation id="DF77EB8EC16AFF8F1F34F775FE3FF9FE" author="Kraus O." box="[355,415,1602,1629]" pageId="7" pageNumber="8" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">1966</bibRefCitation>
) included 19 species in
<taxonomicName id="7CE6EDFCC16AFF8F1C90F775FCC0F9FF" authorityName="Attems" authorityYear="1914" box="[711,864,1602,1628]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1C90F775FCC0F9FF" box="[711,864,1602,1628]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
</taxonomicName>
, distributed in southern African as far north as
<collectingCountry id="C3F1D6EFC16AFF8F1E89F752FEFBF9DC" box="[222,347,1637,1663]" name="Zimbabwe" pageId="7" pageNumber="8">Zimbabwe</collectingCountry>
and southern
<collectingCountry id="C3F1D6EFC16AFF8F1C51F752FD04F9DC" box="[518,676,1637,1663]" name="Mozambique" pageId="7" pageNumber="8">Mozambique</collectingCountry>
, and
<bibRefCitation id="DF77EB8EC16AFF8F1CB3F752FC72F9DC" author="Frederiksen S. B." box="[740,978,1637,1663]" pageId="7" pageNumber="8" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen (2013a)</bibRefCitation>
added a 20
<superScript id="4C933B37C16AFF8F1A0AF754FBCAF9D2" attach="left" box="[1117,1130,1635,1649]" fontSize="6" pageId="7" pageNumber="8">th</superScript>
species from northern
<collectingCountry id="C3F1D6EFC16AFF8F1EEAF7BFFE8AF901" box="[189,298,1672,1698]" name="Tanzania" pageId="7" pageNumber="8">Tanzania</collectingCountry>
.
<bibRefCitation id="DF77EB8EC16AFF8F1F60F7BEFE6DF901" author="Kraus O." box="[311,461,1672,1699]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960</bibRefCitation>
,
<bibRefCitation id="DF77EB8EC16AFF8F1F8CF7BFFDB7F900" author="Kraus O." box="[475,535,1672,1699]" pageId="7" pageNumber="8" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">1966</bibRefCitation>
) did not include
<taxonomicName id="7CE6EDFCC16AFF8F1CB2F7BFFB16F900" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[741,1206,1672,1699]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC16AFF8F1CB2F7BFFBBAF901" box="[741,1050,1672,1698]" italics="true" pageId="7" pageNumber="8">Chaleponcus dabagaensis</emphasis>
<bibRefCitation id="DF77EB8EC16AFF8F1A75F7BEFB16F900" author="Kraus O." box="[1058,1206,1672,1699]" pageId="7" pageNumber="8" pagination="1 - 16" refId="ref30842" refString="Kraus O. 1958. Myriapoden von Ostafrika (Tanganyika Territory). Veroffentlichungen des Uberseemuseums in Bremen, Reihe A 3 (1): 1 - 16." type="journal article" year="1958">Kraus, 1958</bibRefCitation>
</taxonomicName>
, described from the Udzungwa Mts (not W Usambaras as stated by
<bibRefCitation id="DF77EB8EC16AFF8F1D15F79CFB85F965" author="Frederiksen S. B." box="[834,1061,1707,1734]" pageId="7" pageNumber="8" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen 2013a</bibRefCitation>
); this species
<bibRefCitation id="DF77EB8EC16AFF8F1A81F79BFAD6F965" author="Kraus O." box="[1238,1398,1707,1734]" pageId="7" pageNumber="8" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus (1960)</bibRefCitation>
excluded from the genus and tentatively assigned to another predominantly S African genus,
<taxonomicName id="7CE6EDFCC16AFF8F1ABBF7F8FEFFF8AF" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1ABBF7F8FAD7F94A" box="[1260,1399,1743,1769]" italics="true" pageId="7" pageNumber="8">Spinotarsus</emphasis>
Attems, 1909
</taxonomicName>
. In fact, he described the species as
<taxonomicName id="7CE6EDFCC16AFF8F1D58F7C5FB58F8AF" authorityName="Kraus" authorityYear="1958" box="[783,1272,1778,1804]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dabagaensis" subGenus="Storthoporus">
<emphasis id="89924A6DC16AFF8F1D58F7C5FC08F8AF" box="[783,936,1778,1804]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
(
<emphasis id="89924A6DC16AFF8F1DEDF7C5FBF3F8AF" box="[954,1107,1778,1804]" italics="true" pageId="7" pageNumber="8">Storthoporus</emphasis>
)
<emphasis id="89924A6DC16AFF8F1A32F7C5FB58F8AF" box="[1125,1272,1778,1804]" italics="true" pageId="7" pageNumber="8">dabagaensis</emphasis>
</taxonomicName>
, and as he (1960) listed
<emphasis id="89924A6DC16AFF8F1F36F622FE5AF88C" box="[353,506,1813,1839]" italics="true" pageId="7" pageNumber="8">Storthoporus</emphasis>
Attems, 1928, as a synonym of
<emphasis id="89924A6DC16AFF8F1DDBF622FBBEF88C" box="[908,1054,1813,1839]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="7CE6EDFCC16AFF8F1DDBF622FBB9F88C" authorityName="Attems" authorityYear="1909" box="[908,1049,1813,1839]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Spinotarsus</taxonomicName>
,
</emphasis>
the transfer seemed logical. However,
<taxonomicName id="7CE6EDFCC16AFF8F1F61F60EFE4BF8F1" authorityName="Kraus" authorityYear="1958" box="[310,491,1848,1875]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC16AFF8F1F61F60EFE4BF8F1" box="[310,491,1848,1875]" italics="true" pageId="7" pageNumber="8">C. dabagaensis</emphasis>
</taxonomicName>
lacks most characteristics of
<taxonomicName id="7CE6EDFCC16AFF8F1D12F60EFC70F8F0" authorityName="Attems" authorityYear="1909" box="[837,976,1849,1875]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1D12F60EFC70F8F0" box="[837,976,1849,1875]" italics="true" pageId="7" pageNumber="8">Spinotarsus</emphasis>
</taxonomicName>
, including a spiny proximal lamella on the telomere and the strongly sclerotized lamella (“Bogenlamelle”) on the posterior surface of the telomere.
<taxonomicName id="7CE6EDFCC16AFF8F1F62F648FE4CF83A" authorityName="Kraus" authorityYear="1958" box="[309,492,1919,1945]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC16AFF8F1F62F648FE4CF83A" box="[309,492,1919,1945]" italics="true" pageId="7" pageNumber="8">C. dabagaensis</emphasis>
</taxonomicName>
does have a trait seen in many
<taxonomicName id="7CE6EDFCC16AFF8F1D39F648FC59F83A" authorityName="Attems" authorityYear="1909" box="[878,1017,1919,1945]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1D39F648FC59F83A" box="[878,1017,1919,1945]" italics="true" pageId="7" pageNumber="8">Spinotarsus</emphasis>
</taxonomicName>
species, viz., the longitudinally fluted solenomere; this is a character not seen in other previously described
<taxonomicName id="7CE6EDFCC16AFF8F1A07F695FB49F81F" authorityName="Attems" authorityYear="1914" box="[1104,1257,1954,1980]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC16AFF8F1A07F695FB49F81F" box="[1104,1257,1954,1980]" italics="true" pageId="7" pageNumber="8">Chaleponcus</emphasis>
</taxonomicName>
species but found in most of the species described here.
<taxonomicName id="7CE6EDFCC16AFF8F1C93F6F2FCD9F87C" authorityName="Kraus" authorityYear="1958" box="[708,889,1989,2015]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC16AFF8F1C93F6F2FCD9F87C" box="[708,889,1989,2015]" italics="true" pageId="7" pageNumber="8">C. dabagaensis</emphasis>
</taxonomicName>
also has the spine-bearing shelf, see below, and without doubt belongs together with the other Udzungwan species treated in the present paper. I here reallocate it back to
<taxonomicName id="7CE6EDFCC165FF801FFCF018FDE4FEEA" authorityName="Attems" authorityYear="1914" box="[427,580,303,329]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801FFCF018FDE4FEEA" box="[427,580,303,329]" italics="true" pageId="8" pageNumber="9">Chaleponcus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BB59967FC165FF801EEAF047FC44FD49" blockId="8.[189,1399,368,747]" pageId="8" pageNumber="9">
The Udzungwa species described here and referred to
<taxonomicName id="7CE6EDFCC165FF801D14F047FC7CFE29" authorityName="Attems" authorityYear="1914" box="[835,988,368,394]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801D14F047FC7CFE29" box="[835,988,368,394]" italics="true" pageId="8" pageNumber="9">Chaleponcus</emphasis>
</taxonomicName>
differ from their congeners in one conspicuous character, the spine-bearing ‘shelf’ on the gonopodal coxal metaplica. Many odontopygids have spines in various positions on the gonopod coxa, including the subdistal position on the metaplica where the spine-bearing shelf is situated in the Udzungwan species. This is, e.g., true for
<taxonomicName id="7CE6EDFCC165FF801A8AF0EDFE5BFDB4" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="parensis">
<emphasis id="89924A6DC165FF801A8AF0EDFEBFFDB4" italics="true" pageId="8" pageNumber="9">Chaleponcus parensis</emphasis>
Frederiksen, 2013
</taxonomicName>
,
<taxonomicName id="7CE6EDFCC165FF801C51F0CAFB38FDB4" authority="VandenSpiegel &amp; Pierrard, 2009" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2009" box="[518,1176,509,535]" class="Diplopoda" family="Odontopygidae" genus="Prionopetalum" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bifidum">
<emphasis id="89924A6DC165FF801C51F0CAFCB5FDB4" box="[518,789,509,535]" italics="true" pageId="8" pageNumber="9">Prionopetalum bifidum</emphasis>
VandenSpiegel &amp; Pierrard, 2009
</taxonomicName>
,
<taxonomicName id="7CE6EDFCC165FF801AF4F0C9FEF3FD99" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" class="Diplopoda" family="Odontopygidae" genus="Patinatius" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="attemsi">
<emphasis id="89924A6DC165FF801AF4F0C9FAD7FDB4" box="[1187,1399,510,535]" italics="true" pageId="8" pageNumber="9">Patinatius attemsi</emphasis>
<bibRefCitation id="DF77EB8EC165FF801EEAF317FEF3FD99" author="Kraus O." box="[189,339,544,570]" pageId="8" pageNumber="9" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7CE6EDFCC165FF801F34F316FD00FD99" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" box="[355,672,544,570]" class="Diplopoda" family="Odontopygidae" genus="Patinatius" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="89924A6DC165FF801F34F316FE5EFD99" box="[355,510,544,570]" italics="true" pageId="8" pageNumber="9">P. bidentatus</emphasis>
<bibRefCitation id="DF77EB8EC165FF801C5DF317FD00FD99" author="Kraus O." box="[522,672,544,570]" pageId="8" pageNumber="9" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7CE6EDFCC165FF801CE7F317FB93FD99" authority="Kraus, 1966" authorityName="Kraus" authorityYear="1966" box="[688,1075,544,570]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="89924A6DC165FF801CE7F317FC31FD99" box="[688,913,544,570]" italics="true" pageId="8" pageNumber="9">Spinotarsus viridis</emphasis>
<bibRefCitation id="DF77EB8EC165FF801DCAF317FB93FD99" author="Kraus O." box="[925,1075,544,570]" pageId="8" pageNumber="9" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">Kraus, 1966</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7CE6EDFCC165FF801A14F317FB7FFD99" box="[1091,1247,544,570]" class="Diplopoda" family="Odontopygidae" genus="Allantogonus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801A14F317FB7FFD99" box="[1091,1247,544,570]" italics="true" pageId="8" pageNumber="9">Allantogonus</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="DF77EB8EC165FF801B78F317FF5AFDFE" author="Kraus O." pageId="8" pageNumber="9" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus 1960</bibRefCitation>
, Attems 1935),
<taxonomicName id="7CE6EDFCC165FF801FE4F373FDF9FDFE" authorityName="Attems" authorityYear="1935" box="[435,601,580,605]" class="Diplopoda" family="Odontopygidae" genus="Kompsoprium" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801FE4F373FDF9FDFE" box="[435,601,580,605]" italics="true" pageId="8" pageNumber="9">Kompsoprium</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="DF77EB8EC165FF801CC8F374FC8BFDFE" author="Kraus O." box="[671,811,579,605]" pageId="8" pageNumber="9" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus 1960</bibRefCitation>
, Attems 1935) and
<taxonomicName id="7CE6EDFCC165FF801A59F374FF5BFD23" authority="Kraus, 1960" authorityName="Kraus" authorityYear="1960" class="Diplopoda" family="Odontopygidae" genus="Odontopygista" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="natalica">
<emphasis id="89924A6DC165FF801A59F374FA82FDFE" box="[1038,1314,579,605]" italics="true" pageId="8" pageNumber="9">Odontopygista natalica</emphasis>
<bibRefCitation id="DF77EB8EC165FF801B7EF373FF5BFD23" author="Kraus O." pageId="8" pageNumber="9" pagination="1 - 207" refId="ref30872" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">Kraus, 1960</bibRefCitation>
</taxonomicName>
. In all these species except one, however, the spine is not associated with a shelflike structure and are most likely not homologous with the ‘shelf-borne’ spine in the Udzungwan species. The exception is
<taxonomicName id="7CE6EDFCC165FF801E8EF39AFE16FD64" authorityName="Kraus" authorityYear="1966" box="[217,438,685,711]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="89924A6DC165FF801E8EF39AFE16FD64" box="[217,438,685,711]" italics="true" pageId="8" pageNumber="9">Spinotarsus viridis</emphasis>
</taxonomicName>
, where the spine does seem to originate from a horizontal shelf (
<bibRefCitation id="DF77EB8EC165FF801AE1F39AFAE4FD64" author="Kraus O." box="[1206,1348,685,711]" pageId="8" pageNumber="9" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">Kraus 1966</bibRefCitation>
: fig. 350); in other characters, however,
<taxonomicName id="7CE6EDFCC165FF801C0BF3E7FD64FD49" authorityName="Kraus" authorityYear="1966" box="[604,708,720,746]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="89924A6DC165FF801C0BF3E7FD64FD49" box="[604,708,720,746]" italics="true" pageId="8" pageNumber="9">S. viridis</emphasis>
</taxonomicName>
is a typical
<taxonomicName id="7CE6EDFCC165FF801D05F3E7FC7DFD49" authorityName="Attems" authorityYear="1909" box="[850,989,720,746]" class="Diplopoda" family="Odontopygidae" genus="Spinotarsus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801D05F3E7FC7DFD49" box="[850,989,720,746]" italics="true" pageId="8" pageNumber="9">Spinotarsus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BB59967FC165FF801EEAF226FE03FC35" blockId="8.[189,1399,785,918]" pageId="8" pageNumber="9">
Based on the probably autapomorphic spine-bearing shelf and the longitudinally fluted solenomere, a species group can be defined for the Udzungwan species of
<taxonomicName id="7CE6EDFCC165FF801DE7F202FBE9FCEC" authorityName="Attems" authorityYear="1914" box="[944,1097,821,847]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC165FF801DE7F202FBE9FCEC" box="[944,1097,821,847]" italics="true" pageId="8" pageNumber="9">Chaleponcus</emphasis>
</taxonomicName>
. Following tradition, the group is named after the first described species in the group and hence will be named the
<taxonomicName id="7CE6EDFCC165FF801A8AF26FFEF0FC36" authorityName="Kraus" authorityYear="1958" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC165FF801A8AF26FFEF0FC36" italics="true" pageId="8" pageNumber="9">Chaleponcus dabagaensis</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>
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