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<document id="C0582DBFCBA995B1F400D7A0362337FB" ID-DOI="10.5852/ejt.2014.100" ID-GBIF-Dataset="5329bfc4-61b2-48f5-a96a-caf4ba31039b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860450" ID-ZooBank="B3E6C489-6D96-4AF5-A33D-EE8329A9321B" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1590595554426" checkinUser="carolina" docAuthor="Enghoff, Henrik" docDate="2014" docId="334F2769C12FFFC01EEAF728FED8FEEB" docLanguage="en" docName="ejt-2014-100.pdf.imf" docOrigin="European Journal of Taxonomy 100 (100)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chaleponcus dabagaensis Kraus 1958" docType="key" docVersion="10" lastPageNumber="70" masterDocId="CF765F11C16DFF881E57F137FFA0FFA3" masterDocTitle="A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="67" updateTime="1736355869612" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CD0460BA566E5A994F83C88E1688B1B2">A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="5C5E1D23D984737617225387EC1BAD6F">Enghoff, Henrik</mods:namePart>
<mods:affiliation id="B75180906B67F684854120265233D541">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:title id="D2BBC6BE59C59900E62DFF616ABB36BF">European Journal of Taxonomy</mods:title>
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<treatment id="334F2769C12FFFC01EEAF728FED8FEEB" ID-DOI="http://doi.org/10.5281/zenodo.14617429" ID-GBIF-Taxon="164362788" ID-Zenodo-Dep="14617429" LSID="urn:lsid:plazi:treatment:334F2769C12FFFC01EEAF728FED8FEEB" httpUri="http://treatment.plazi.org/id/334F2769C12FFFC01EEAF728FED8FEEB" lastPageId="72" lastPageNumber="70" pageId="66" pageNumber="67" scope_class="Diplopoda" scope_family="Odontopygidae" scope_order="Spirostreptida" scope_subFamily="Archepyginae" scope_tribe="Prionopetalini">
<subSubSection id="F3FCC5F4C12FFFCA1EEAF728FC23F998" box="[189,899,1566,1595]" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="BB59967FC12FFFCA1EEAF728FC23F998" blockId="66.[189,899,1566,1595]" box="[189,899,1566,1595]" pageId="66" pageNumber="67">
<heading id="E0112113C12FFFCA1EEAF728FC23F998" box="[189,899,1566,1595]" pageId="66" pageNumber="67">
<emphasis id="89924A6DC12FFFCA1EEAF728FC23F998" bold="true" box="[189,899,1566,1595]" pageId="66" pageNumber="67">
<emphasis id="89924A6DC12FFFCA1EEAF728FE68F998" bold="true" box="[189,456,1566,1595]" italics="true" pageId="66" pageNumber="67">Key to species of the</emphasis>
<taxonomicName id="7CE6EDFCC12FFFCA1F87F729FC8DF998" ID-CoL="TH64" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[464,813,1566,1595]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dabagaensis">Chaleponcus dabagaensis</taxonomicName>
<emphasis id="89924A6DC12FFFCA1D62F728FC23F998" bold="true" box="[821,899,1567,1595]" italics="true" pageId="66" pageNumber="67">group</emphasis>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="F3FCC5F4C12FFFCA1EEAF765FC4AF8E3" pageId="66" pageNumber="67" type="description">
<paragraph id="BB59967FC12FFFCA1EEAF765FC4AF8E3" blockId="66.[189,1399,1618,1856]" pageId="66" pageNumber="67">
The 21 known species of the 
<taxonomicName id="7CE6EDFCC12FFFCA1C17F764FD5CF9CF" authorityName="Kraus" authorityYear="1958" box="[576,764,1618,1645]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC12FFFCA1C17F764FD5CF9CF" box="[576,764,1618,1645]" italics="true" pageId="66" pageNumber="67">C. dabagaensis</emphasis>
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group are almost all quite distinctive in gonopod morphology. A key to the species is given below, but comparison with the gonopod coxa profiles in 
<figureCitation id="23DD8AFAC12FFFCA1B1FF741FF6CF910" captionStart="Fig" captionStartId="15.[187,230,1816,1842]" captionTargetBox="[247,1356,507,1717]" captionTargetId="figure@15.[200,1385,488,1723]" captionTargetPageId="15" captionText="Fig. 9. Chaleponcus spp., outline of coxite of left gonopod, posterior view. (The ’transverse’ plane of the gonopod is usually slightly at angles with the exact transverse plane of the millipede, so it may be necessary to tilt the gonopod slightly in order to obtain the view shown on the drawings.) The stippled lines indicate the place where the telopodital basomere emerges from the coxal cavity. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3860472" httpUri="https://zenodo.org/record/3860472/files/figure.png" pageId="66" pageNumber="67">Fig. 9</figureCitation>
will often lead directly to the right species. As new species of the group are likely to be discovered, and as the species in the key may prove to be more variable with regard to number of podous rings and body diameter, any identification should be checked against the detailed gonopod description and illustrations, and the size (no. of podous rings vs. body diameter) should be compared with the diagrams, 
<figureCitation id="23DD8AFAC12FFFCA1EEAF611FE8EF8E3" box="[189,302,1830,1856]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1930,1956]" captionStartId-1="9.[189,232,1932,1958]" captionTargetBox-0="[204,1366,1100,1853]" captionTargetBox-1="[211,1383,1371,1882]" captionTargetId-0="figure@8.[189,1398,1047,1869]" captionTargetId-1="figure@9.[189,1398,1353,1891]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 2. Body size of males of 13 species of the Chaleponcus dabagaensis group: Number of podous rings and vertical midbody diameter in large, small and ‘long, thin’ species (cf. text), as well as medium sized species in which the gonopod coxa has an obvious lateral process." captionText-1="Fig. 3. Body size of males of 8 species of the Chaleponcus dabagaensis group: Number of podous rings and vertical midbody diameter in medium-sized species in which the gonopod coxa has no obvious lateral process." figureDoi-0="http://doi.org/10.5281/zenodo.3860456" figureDoi-1="http://doi.org/10.5281/zenodo.3860458" httpUri-0="https://zenodo.org/record/3860456/files/figure.png" httpUri-1="https://zenodo.org/record/3860458/files/figure.png" pageId="66" pageNumber="67">Figs. 2–3</figureCitation>
. The limbus (
<figureCitation id="23DD8AFAC12FFFCA1F99F611FDB4F8E3" box="[462,532,1830,1856]" captionStart="Fig" captionStartId="11.[189,231,1754,1780]" captionTargetBox="[189,1398,416,1713]" captionTargetId="figure@11.[189,1398,416,1713]" captionTargetPageId="11" captionText="Fig. 4. Limbus types in Chaleponcus species, SEM micrographs, external view, except I: internal view. — A. Lobes pointed, more than twice as long as broad (C. scopus sp. nov.). B. Lobes pointed, slightly less than twice as long as broad (C. howelli sp. nov.). C. Lobes pointed, almost equilateral triangles (C. netus sp. nov.), D. Lobes rounded, slightly longer than broad (C. dabagaensis Kraus, 1958). E. Lobes broadly rounded, slightly shorter than broad (C. hamerae sp. nov.). F. Lobes spatulate (C. gracilior sp. nov.). G. Lobes rounded, broader than long, spinose, ridged (C. krai sp. nov.). H. Lobes ca. 3 times as broad as high, rounded, spinose (C. teres sp. nov.). I. Lobes rounded, much broader than high, smooth (C. malleolus sp. nov.). J. No lobes (C. circumvallatus sp. nov.). Scales: 0.01 mm (A– F, J), 0.001 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.3860460" httpUri="https://zenodo.org/record/3860460/files/figure.png" pageId="66" pageNumber="67">Fig. 4</figureCitation>
) offers additional diagnostic characters.
</paragraph>
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<subSubSection id="F3FCC5F4C12FFFCA1EEAF650FD9BF822" box="[189,571,1895,1922]" pageId="66" pageNumber="67" type="multiple">
<paragraph id="BB59967FC12FFFCA1EEAF650FD9BF822" blockId="66.[189,571,1895,1922]" box="[189,571,1895,1922]" pageId="66" pageNumber="67">The key is based on adult males.</paragraph>
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<keyLead id="0017354AC12FFFCA1EEAF69EFAD6F860" box="[189,1398,1960,1987]" pageId="66" pageNumber="67">
1. Body diameter&gt; 
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…………………………………………………………………………2
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– Body diameter &lt;
<quantity id="7C1E3B9AC12FFFCA1F96F6FBFD83F845" box="[449,547,1996,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="66" pageNumber="67" unit="mm" value="2.5">2.5 mm</quantity>
…………………………………………………………………………5
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2. Gonopod coxa with two lateral processes ………………………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1AC8F03DFAB3FE87" ID-CoL="TH5X" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1183,1299,266,292]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="basiliscus" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1AC8F03DFAB3FE87" box="[1183,1299,266,292]" italics="true" pageId="67" pageNumber="68">basiliscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B4CF03DFAD7FE87" box="[1307,1399,266,292]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF019FAD7FEE4" blockId="67.[187,1399,266,328]" box="[187,1399,301,328]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF019FAD7FEE4" box="[187,1399,301,328]" pageId="67" pageNumber="68">– Gonopod coxa without lateral processes ……………………………………………………………3</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF058FAD7FE51" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF058FAD7FE0F" blockId="67.[187,1399,367,499]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF058FAD7FE0F" pageId="67" pageNumber="68">
3. Anterior lamella of telomere divided into a horizontal lobe with deeply laciniate edges and a long, straight, distal spine-like part (
<figureCitation id="23DD8AFAC12EFFCB1C63F0A5FD3BFE0F" box="[564,667,402,428]" captionStart="Fig" captionStartId="37.[187,230,1859,1885]" captionTargetBox="[189,1398,511,1821]" captionTargetId="figure@37.[189,1399,510,1822]" captionTargetPageId="37" captionText="Fig. 19. Chaleponcus krai sp. nov., gonopods. Specimens from New Dabaga-Ulangambi (A) and Kalimbazi Mtn (B-D). A. Left gonopod, anterior view. B–D. Right gonopod. B. Mesal-dorsal view. C. Telopodite, mesal view. D. Posterior view. al = anterior lobe of telomere, cu = cucullus, il = intermediate lobe of telomere, mfp = metaplical flange process, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860492" httpUri="https://zenodo.org/record/3860492/files/figure.png" pageId="67" pageNumber="68">Fig. 19C</figureCitation>
). Setation of walking leg tarsi normal (as 
<figureCitation id="23DD8AFAC12EFFCB1A3CF0A5FB62FE0E" box="[1131,1218,402,429]" captionStart="Fig" captionStartId="12.[187,230,760,786]" captionTargetBox="[314,1268,265,748]" captionTargetId="figure@12.[314,1268,265,748]" captionTargetPageId="12" captionText="Fig. 5. Chaleponcus spp., tip of tarsus and claw of walking legs. — A. C. mwanihanensis sp. nov. as an example of normal setation. B–C. C. circumvallatus sp. nov. with the claw surrounded by a ’palisade’ of setae. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860462" httpUri="https://zenodo.org/record/3860462/files/figure.png" pageId="67" pageNumber="68">Fig.5A</figureCitation>
) … 
<taxonomicName id="7CE6EDFCC12EFFCB1ABBF0A5FABBFE0F" ID-CoL="TH6R" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1260,1307,402,428]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="krai" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1ABBF0A5FABBFE0F" box="[1260,1307,402,428]" italics="true" pageId="67" pageNumber="68">krai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B77F0A5FAD7FE0F" box="[1312,1399,402,428]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF081FAD7FE51" blockId="67.[187,1399,367,499]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF081FAD7FE51" pageId="67" pageNumber="68">
– Anterior lamella of telomere different. Setation of walking legs unusual: claws surrounded by dense ‘palisade’ of setae (
<figureCitation id="23DD8AFAC12EFFCB1C41F0EFFDFCFE50" box="[534,604,472,499]" captionStart="Fig" captionStartId="12.[187,230,760,786]" captionTargetBox="[314,1268,265,748]" captionTargetId="figure@12.[314,1268,265,748]" captionTargetPageId="12" captionText="Fig. 5. Chaleponcus spp., tip of tarsus and claw of walking legs. — A. C. mwanihanensis sp. nov. as an example of normal setation. B–C. C. circumvallatus sp. nov. with the claw surrounded by a ’palisade’ of setae. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860462" httpUri="https://zenodo.org/record/3860462/files/figure.png" pageId="67" pageNumber="68">Fig. 5</figureCitation>
B–C) ………………………………………………………………4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF32DFAD7FD3D" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF32DFAD7FDF4" blockId="67.[187,1399,538,670]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF32DFAD7FDF4" pageId="67" pageNumber="68">
4. Posterior lamella of telomere ending in two long spines. Anterior lamella of telomere with small accessory lamella, otherwise with smooth edges (
<figureCitation id="23DD8AFAC12EFFCB1D30F30AFC7BFDF4" box="[871,987,573,599]" captionStart="Fig" captionStartId="42.[187,229,1892,1918]" captionTargetBox="[189,1398,550,1833]" captionTargetId="figure@42.[189,1398,550,1833]" captionTargetPageId="42" captionText="Fig. 21. Chaleponcus circumvallatus sp. nov., left gonopod. A. Anterior view. B. Mesal-anterior view. C. Posterior view. D. Mesal-posterior view. al = anterior lobe of telomere, al’ = accessory lamella, cu = cucullus, mfp = metaplical flange process, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860496" httpUri="https://zenodo.org/record/3860496/files/figure.png" pageId="67" pageNumber="68">Fig. 21B</figureCitation>
) ……… 
<taxonomicName id="7CE6EDFCC12EFFCB1A18F30AFB5CFDF4" ID-CoL="TH62" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1103,1276,573,599]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1A18F30AFB5CFDF4" box="[1103,1276,573,599]" italics="true" pageId="67" pageNumber="68">circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B47F30AFAD7FDF4" box="[1296,1399,573,599]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF356FAD7FD3D" blockId="67.[187,1399,538,670]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF356FAD7FD3D" pageId="67" pageNumber="68">
– Posterior lamella of telomere ending in two short processes. Anterior lamella of telomere with a coarsely dentate basal part (
<figureCitation id="23DD8AFAC12EFFCB1C72F3B4FD2EFD3E" box="[549,654,643,669]" captionStart="Fig" captionStartId="44.[189,232,1893,1919]" captionTargetBox="[190,1398,501,1853]" captionTargetId="figure@44.[189,1398,501,1853]" captionTargetPageId="44" captionText="Fig. 22. Chaleponcus ibis sp. nov., left gonopod. A. Posterior view. B. Anterior view. C. Telopodite, mesal-anterior view. al = anterior lobe of telomere, al’ = accessory lamella?, cu = cucullus, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860498" httpUri="https://zenodo.org/record/3860498/files/figure.png" pageId="67" pageNumber="68">Fig. 22C</figureCitation>
) …………………………………………………… 
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<emphasis id="89924A6DC12EFFCB1ABEF3B4FAB3FD3E" box="[1257,1299,643,669]" italics="true" pageId="67" pageNumber="68">ibis</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B4CF3B3FAD7FD3D" box="[1307,1399,644,670]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF3F3FAD6FC86" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF3F3FAD7FCA1" blockId="67.[187,1399,708,806]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF3F3FAD7FCA1" pageId="67" pageNumber="68">
5 45–49 podous rings. Body diameter 
<quantity id="7C1E3B9AC12EFFCB1CDDF3F3FCB8FD7C" box="[650,792,708,735]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="67" pageNumber="68" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
(
<figureCitation id="23DD8AFAC12EFFCB1D7EF3F2FCD2FD7C" box="[809,882,709,735]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[204,1366,1100,1853]" captionTargetId="figure@8.[189,1398,1047,1869]" captionTargetPageId="8" captionText="Fig. 2. Body size of males of 13 species of the Chaleponcus dabagaensis group: Number of podous rings and vertical midbody diameter in large, small and ‘long, thin’ species (cf. text), as well as medium sized species in which the gonopod coxa has an obvious lateral process." figureDoi="http://doi.org/10.5281/zenodo.3860456" httpUri="https://zenodo.org/record/3860456/files/figure.png" pageId="67" pageNumber="68">Fig. 2</figureCitation>
). Cucullus of gonopod coxa subrectangular (
<figureCitation id="23DD8AFAC12EFFCB1EB5F3DFFE9DFCA1" box="[226,317,744,770]" captionStart="Fig" captionStartId="58.[189,232,1787,1813]" captionTargetBox="[277,1307,266,1752]" captionTargetId="figure@58.[276,1308,265,1752]" captionTargetPageId="58" captionText="Fig. 28. Chaleponcus gracilior sp. nov., gonopods. A. Gonopods in situ in body ring 7, anterior view. B. Right gonopod, detail of base of solenomere, showing basal side branch consisting of a tiny, spinelike part and a larger, irregularly lamellate one. C. Right gonopod, anterior-mesal view. D. Left gonopod, posterior view. E. Left telopodite, posterior-lateral view. F. Right telopodite, anterior view. al = anterior telomeral lamella, cu = cucullus, dp = distal telomeral process, mf = metaplical flange, ms = metaplical shelf, mss = metaplical shelf-spine, pb = proximal side branch of solenomere, pl = posterior telomeral lamella, prl = proplical lobe. Scales: 0.1 mm (A, C–F), 0.05 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.3860510" httpUri="https://zenodo.org/record/3860510/files/figure.png" pageId="67" pageNumber="68">Fig. 28</figureCitation>
). Limbus with spatulate lobes (
<figureCitation id="23DD8AFAC12EFFCB1CEBF3DFFCB6FCA1" box="[700,790,744,770]" captionStart="Fig" captionStartId="11.[189,231,1754,1780]" captionTargetBox="[189,1398,416,1713]" captionTargetId="figure@11.[189,1398,416,1713]" captionTargetPageId="11" captionText="Fig. 4. Limbus types in Chaleponcus species, SEM micrographs, external view, except I: internal view. — A. Lobes pointed, more than twice as long as broad (C. scopus sp. nov.). B. Lobes pointed, slightly less than twice as long as broad (C. howelli sp. nov.). C. Lobes pointed, almost equilateral triangles (C. netus sp. nov.), D. Lobes rounded, slightly longer than broad (C. dabagaensis Kraus, 1958). E. Lobes broadly rounded, slightly shorter than broad (C. hamerae sp. nov.). F. Lobes spatulate (C. gracilior sp. nov.). G. Lobes rounded, broader than long, spinose, ridged (C. krai sp. nov.). H. Lobes ca. 3 times as broad as high, rounded, spinose (C. teres sp. nov.). I. Lobes rounded, much broader than high, smooth (C. malleolus sp. nov.). J. No lobes (C. circumvallatus sp. nov.). Scales: 0.01 mm (A– F, J), 0.001 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.3860460" httpUri="https://zenodo.org/record/3860460/files/figure.png" pageId="67" pageNumber="68">Fig. 4F</figureCitation>
) ………………………………… 
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<emphasis id="89924A6DC12EFFCB1AF1F3DFFAAEFCA1" box="[1190,1294,744,770]" italics="true" pageId="67" pageNumber="68">gracilior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B4EF3DFFAD7FCA1" box="[1305,1399,744,770]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF23BFAD6FC86" blockId="67.[187,1399,708,806]" box="[187,1398,779,806]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF23BFAD6FC86" box="[187,1398,779,806]" pageId="67" pageNumber="68">
– &lt;45 body rings and/or body diameter ≥ 
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. Cucullus and limbus different …………………6
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</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF27BFAD7FC29" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF27BFAD7FCC4" blockId="67.[187,1399,844,906]" box="[187,1399,844,871]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF27BFAD7FCC4" box="[187,1399,844,871]" pageId="67" pageNumber="68">6 Gonopod coxa with a lateral process ………………………………………………………………7</keyLead>
</paragraph>
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<keyLead id="0017354AC12EFFCB1EECF247FAD7FC29" box="[187,1399,880,906]" pageId="67" pageNumber="68">– Gonopod coxa without a lateral process …………………………………………………………14</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF285FAD7FB96" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF285FAD7FC4C" blockId="67.[187,1399,945,1077]" pageId="67" pageNumber="68">
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7 &lt;40 body rings. Body diameter 
<quantity id="7C1E3B9AC12EFFCB1C3AF286FCA1FC6F" box="[621,769,945,972]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="67" pageNumber="68" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
. No dorsal spine on anal valves. Lateral process of gonopod coxa a strongly bent hook (
<figureCitation id="23DD8AFAC12EFFCB1CECF2E3FCB8FC4D" box="[699,792,980,1006]" captionStart="Fig" captionStartId="25.[187,230,1852,1878]" captionTargetBox="[242,1338,265,1799]" captionTargetId="figure@25.[242,1338,265,1799]" captionTargetPageId="25" captionText="Fig. 14. Chaleponcus malleolus sp. nov., right gonopod. A. Anterior view. B. Mesal view, focus on coxa. C. Posterior view. D. Anterior-mesal view. E. Mesal view, focus on telopodite. F. Telopodite, ventral-posterior view. al = anterior lamella of telomere, cu = cucullus, mfp = metaplical flange process, mp = mesal metaplical process, mss; metaplical shelf-spine, pl1, pl2, pl3 = lobes of posterior lamella of telomere, st = main stem of telomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860482" httpUri="https://zenodo.org/record/3860482/files/figure.png" pageId="67" pageNumber="68">Fig. 14</figureCitation>
) ……………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1AC5F2E3FAA4FC4D" ID-CoL="5XQDB" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1170,1284,980,1006]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="malleolus" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1AC5F2E3FAA4FC4D" box="[1170,1284,980,1006]" italics="true" pageId="67" pageNumber="68">malleolus</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B43F2E2FAD7FC4C" box="[1300,1399,981,1007]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF2CFFAD7FB96" blockId="67.[187,1399,945,1077]" pageId="67" pageNumber="68">
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–&gt; 40 body rings. Body diameter ≥ 
<quantity id="7C1E3B9AC12EFFCB1CDEF2CFFD4AFBB1" box="[649,746,1016,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="67" pageNumber="68" unit="mm" value="1.8">1.8 mm</quantity>
. Each anal valve with a dorsal spine. Lateral process of gonopod coxa different …………………………………………………………………………8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF56BFAD6FB1E" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF56BFAD7FB39" blockId="67.[187,1399,1116,1213]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF56BFAD7FB39" pageId="67" pageNumber="68">
8 Metaplical shelf of gonopod coxa in two levels between which metaplical shelf-spine fits. Lateral process of gonopod coxa small to inconspicuous (
<figureCitation id="23DD8AFAC12EFFCB1D05F548FC0EFB39" box="[850,942,1151,1178]" captionStart="Fig" captionStartId="48.[189,232,1823,1849]" captionTargetBox="[189,1398,386,1781]" captionTargetId="figure@48.[189,1398,386,1781]" captionTargetPageId="48" captionText="Fig. 24. Chaleponcus vilici sp. nov., left gonopod. A. Posterior view. B. Mesal view. C. Mesal-anterior view. D. Close-up of metaplical shelf-spine, mesal view. E. Anterior (and slightly apical/ventral) view. F. Telopodite, distal view, showing solenomere and the two distal telomere branches. al = anterior lamella of telomere, cu = cucullus, la = lateral angle of coxa, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, slm = solenomere. Scales: 0.1 mm (A–D), 0.01 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.3860502" httpUri="https://zenodo.org/record/3860502/files/figure.png" pageId="67" pageNumber="68">Fig. 24</figureCitation>
) ……………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1A9AF548FAA8FB3A" ID-CoL="TH7Q" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1229,1288,1151,1177]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="vilici" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1A9AF548FAA8FB3A" box="[1229,1288,1151,1177]" italics="true" pageId="67" pageNumber="68">vilici</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B41F5B7FAD7FB39" box="[1302,1399,1152,1178]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF594FAD6FB1E" blockId="67.[187,1399,1116,1213]" box="[187,1398,1187,1213]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF594FAD6FB1E" box="[187,1398,1187,1213]" pageId="67" pageNumber="68">– Metaplical shelf in one level. Metaplical shelf-spine and lateral process different …………9</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC12EFFCB1EECF5D3FAD6FAE6" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF5D3FAD7FA82" blockId="67.[187,1399,1252,1349]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF5D3FAD7FA82" pageId="67" pageNumber="68">
9 Lateral margin of gonopod coxa basal to lateral process pronouncedly ‘hunch-backed’ (
<figureCitation id="23DD8AFAC12EFFCB1AB0F5D3FACEFB5D" box="[1255,1390,1252,1278]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[187,230,1858,1884]" captionStartId-1="23.[189,232,1896,1922]" captionTargetBox-0="[189,1398,437,1805]" captionTargetBox-1="[189,1398,289,1853]" captionTargetId-0="figure@21.[189,1398,437,1805]" captionTargetId-1="figure@23.[189,1398,289,1853]" captionTargetPageId-0="21" captionTargetPageId-1="23" captionText-0="Fig. 12. Chaleponcus dabagaensis Kraus, 1958, left gonopod. — A. Anterior view. B. Posterior view. C. Mesal-ventral view. D. Mesal-dorsal view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere, st = main telomeral stem, th = thumblike process of telomere main stem. Scales 0.1 mm." captionText-1="Fig. 13. Chaleponcus quasimodo sp. nov., left gonopod. A. Anterior view. B. Anterior-mesal view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3860478" figureDoi-1="http://doi.org/10.5281/zenodo.3860480" httpUri-0="https://zenodo.org/record/3860478/files/figure.png" httpUri-1="https://zenodo.org/record/3860480/files/figure.png" pageId="67" pageNumber="68">Figs 12–13</figureCitation>
) ……………………………………………………………………………………………………10
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF41CFAD6FAE6" blockId="67.[187,1399,1252,1349]" box="[187,1398,1322,1349]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF41CFAD6FAE6" box="[187,1398,1322,1349]" pageId="67" pageNumber="68">– Lateral margin of gonopod coxa basal to lateral process not pronouncedly ‘hunch-backed’ ………11</keyLead>
</paragraph>
</keyStep>
<paragraph id="BB59967FC12EFFCB1EECF45BFAD7FA09" blockId="67.[187,1399,1388,1450]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF45BFAD7FA09" pageId="67" pageNumber="68">
<keyStep id="00128EDAC12EFFCB1EECF45BFAD7FA09" pageId="67" pageNumber="68">
10 Main stem of telomere with thumblike process (
<figureCitation id="23DD8AFAC12EFFCB1D69F45BFC0BFA25" box="[830,939,1388,1414]" captionStart="Fig" captionStartId="21.[187,230,1858,1884]" captionTargetBox="[189,1398,437,1805]" captionTargetId="figure@21.[189,1398,437,1805]" captionTargetPageId="21" captionText="Fig. 12. Chaleponcus dabagaensis Kraus, 1958, left gonopod. — A. Anterior view. B. Posterior view. C. Mesal-ventral view. D. Mesal-dorsal view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere, st = main telomeral stem, th = thumblike process of telomere main stem. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860478" httpUri="https://zenodo.org/record/3860478/files/figure.png" pageId="67" pageNumber="68">Fig. 12D</figureCitation>
) ………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1AB3F45BFAD7FA25" ID-CoL="TH64" authorityName="Kraus" authorityYear="1958" box="[1252,1399,1388,1414]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC12EFFCB1AB3F45BFAD7FA25" box="[1252,1399,1388,1414]" italics="true" pageId="67" pageNumber="68">dabagaensis</emphasis>
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– Main stem of telomere without thumblike process (
<figureCitation id="23DD8AFAC12EFFCB1D10F4B8FC3AFA0A" box="[839,922,1423,1449]" captionStart="Fig" captionStartId="23.[189,232,1896,1922]" captionTargetBox="[189,1398,289,1853]" captionTargetId="figure@23.[189,1398,289,1853]" captionTargetPageId="23" captionText="Fig. 13. Chaleponcus quasimodo sp. nov., left gonopod. A. Anterior view. B. Anterior-mesal view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860480" httpUri="https://zenodo.org/record/3860480/files/figure.png" pageId="67" pageNumber="68">Fig. 13</figureCitation>
) …………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1AC2F4B8FAB6FA0A" ID-CoL="TH7B" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1173,1302,1423,1449]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1AC2F4B8FAB6FA0A" box="[1173,1302,1423,1449]" italics="true" pageId="67" pageNumber="68">quasimodo</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B4AF4A7FAD7FA09" box="[1309,1399,1424,1450]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF4E6FAD7F9AD" blockId="67.[187,1399,1488,1550]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF4E6FAD7F9AD" pageId="67" pageNumber="68">
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11 Proximal lobe of telomere with long, fluted spine (
<figureCitation id="23DD8AFAC12EFFCB1D60F4E7FC06FA48" box="[823,934,1488,1515]" captionStart="Fig" captionStartId="17.[189,231,1928,1954]" captionTargetBox="[189,1398,266,1895]" captionTargetId="figure@17.[189,1398,266,1895]" captionTargetPageId="17" captionText="Fig. 10. Chaleponcus netus sp. nov., left gonopod, coloured. — A. Posterior view. B. Ventral view. Orange = coxite, blue = torsotope, yellow = solenomere (slm), green = proximal spine of solenomere (ps), pink = telomere. pxs = spine on proximal telomeral lobe, st = main stem of telomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860474" httpUri="https://zenodo.org/record/3860474/files/figure.png" pageId="67" pageNumber="68">Figs 10A</figureCitation>
, 
<figureCitation id="23DD8AFAC12EFFCB1DE6F4E6FC40FA48" box="[945,992,1489,1515]" captionStart="Fig" captionStartId="18.[189,232,1751,1777]" captionTargetBox="[265,1320,265,1716]" captionTargetId="figure@18.[265,1320,265,1716]" captionTargetPageId="18" captionText="Fig. 11. Chaleponcus netus sp. nov., gonopods. — A. Gonopods, anterior view. B. Left coxite, mesal view. C. Base of proximal solenomere spine (ps) with spinose socket. D. Right gonopod, anterior (and slightly mesal) view. E. Left gonopod telopodite, posterior view. F. Base of proximal solenomere spine (ps) in a different individual, with denticle (ps2). al = anterior lamella of telomere, il = intermediate lamella of telomere, mfp = metaplical flange process, ms = metaplical shelf, mss = metaplical shelfspine, pl = posterior lamella of telomere, ps = proximal spine of solenomere, ps2 = additional proximal spine of solenomere, pxl = proximal lobe of telomere, pxs = spine on proximal lobe of telomere. Scales: 0.1 mm (A–B, D–E), 0.05 mm (C, F)." figureDoi="http://doi.org/10.5281/zenodo.3860476" httpUri="https://zenodo.org/record/3860476/files/figure.png" pageId="67" pageNumber="68">11E</figureCitation>
) …………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1A8DF4E6FAB8FA49" ID-CoL="TH74" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1242,1304,1489,1514]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="netus" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1A8DF4E6FAB8FA49" box="[1242,1304,1489,1514]" italics="true" pageId="67" pageNumber="68">netus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12EFFCB1B49F4E6FAD7FA48" box="[1310,1399,1489,1515]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
– Proximal lobe of telomere without a spine ………………………………………………………12
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</paragraph>
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<paragraph id="BB59967FC12EFFCB1EECF702FAD7F9D0" blockId="67.[187,1399,1589,1686]" pageId="67" pageNumber="68">
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12 Lateral process of gonopod coxa small. Cucullus without a large mesal extension (
<figureCitation id="23DD8AFAC12EFFCB1B46F702FACDF9EC" box="[1297,1389,1589,1615]" captionStart="Fig" captionStartId="32.[189,232,1897,1923]" captionTargetBox="[242,1351,265,1860]" captionTargetId="figure@32.[242,1351,265,1860]" captionTargetPageId="32" captionText="Fig. 17. Chaleponcus mwanihanensis sp. nov., gonopods. A. Left gonopods, mesal-ventral view. B. Left gonopod, posterior view. C. Right gonopod, ventral view. D. Right gonopod, anterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860488" httpUri="https://zenodo.org/record/3860488/files/figure.png" pageId="67" pageNumber="68">Fig. 17</figureCitation>
) ……………………………………………………………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1A37F76FFAB5F9D1" ID-CoL="TH6Z" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1120,1301,1624,1650]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="mwanihanensis" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1A37F76FFAB5F9D1" box="[1120,1301,1624,1650]" italics="true" pageId="67" pageNumber="68">mwanihanensis</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B4BF76EFAD7F9D0" box="[1308,1399,1625,1651]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC12EFFCB1EECF74BFAD6F935" blockId="67.[187,1399,1589,1686]" box="[187,1398,1659,1686]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF74BFAD6F935" box="[187,1398,1659,1686]" pageId="67" pageNumber="68">
– Lateral process of gonopod coxa larger. Cucullus with a large mesal extension (
<figureCitation id="23DD8AFAC12EFFCB1AF4F74CFA8AF935" box="[1187,1322,1659,1686]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="28.[189,232,1852,1878]" captionStartId-1="30.[189,232,1896,1922]" captionTargetBox-0="[189,1398,529,1808]" captionTargetBox-1="[189,1399,435,1852]" captionTargetId-0="figure@28.[189,1398,529,1808]" captionTargetId-1="figure@30.[189,1399,435,1852]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionText-0="Fig. 15. Chaleponcus scopus sp. nov., right gonopod. A. Posterior view. B. Ventral-posterior view. C. Mesal-posterior view. D. Anterior view. al = anterior lamella of telomere, al’ = thumblike process of al, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal solenomeral spine, slm = solenomere. Scales 0.1 mm." captionText-1="Fig. 16. Chaleponcus nikolajscharffi sp. nov., right gonopod. A. Mesal view. B. Anterior view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3860484" figureDoi-1="http://doi.org/10.5281/zenodo.3860486" httpUri-0="https://zenodo.org/record/3860484/files/figure.png" httpUri-1="https://zenodo.org/record/3860486/files/figure.png" pageId="67" pageNumber="68">Figs 15–16</figureCitation>
) …13
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<keyStep id="00128EDAC12EFFCB1EECF78AFAD7F8BD" pageId="67" pageNumber="68">
<paragraph id="BB59967FC12EFFCB1EECF78AFAD7F97B" blockId="67.[187,1399,1725,1822]" box="[187,1399,1725,1752]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EECF78AFAD7F97B" box="[187,1399,1725,1752]" pageId="67" pageNumber="68">
13 Lateral process of gonopod coxa pointed (
<figureCitation id="23DD8AFAC12EFFCB1D43F78AFC32F97B" box="[788,914,1725,1752]" captionStart="Fig" captionStartId="28.[189,232,1852,1878]" captionTargetBox="[189,1398,529,1808]" captionTargetId="figure@28.[189,1398,529,1808]" captionTargetPageId="28" captionText="Fig. 15. Chaleponcus scopus sp. nov., right gonopod. A. Posterior view. B. Ventral-posterior view. C. Mesal-posterior view. D. Anterior view. al = anterior lamella of telomere, al’ = thumblike process of al, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal solenomeral spine, slm = solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860484" httpUri="https://zenodo.org/record/3860484/files/figure.png" pageId="67" pageNumber="68">Figs. 15A</figureCitation>
) ……………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1AE4F789FAA3F974" ID-CoL="TH7D" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1203,1283,1726,1751]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="scopus" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1AE4F789FAA3F974" box="[1203,1283,1726,1751]" italics="true" pageId="67" pageNumber="68">scopus</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B44F789FAD7F97B" box="[1299,1399,1726,1752]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="BB59967FC12EFFCB1EEBF7D6FAD7F8BD" blockId="67.[187,1399,1725,1822]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EEBF7D6FAD7F8BD" pageId="67" pageNumber="68">
– Lateral process of gonopod coxa broadly rounded, irregularly dentate (
<figureCitation id="23DD8AFAC12EFFCB1AB8F7D7FACEF958" box="[1263,1390,1760,1787]" captionStart="Fig" captionStartId="30.[189,232,1896,1922]" captionTargetBox="[189,1399,435,1852]" captionTargetId="figure@30.[189,1399,435,1852]" captionTargetPageId="30" captionText="Fig. 16. Chaleponcus nikolajscharffi sp. nov., right gonopod. A. Mesal view. B. Anterior view. C. Ventral view. D. Posterior view. al = anterior lamella of telomere, cu = cucullus, lp = lateral coxal process, mf = metaplical flange, mp = mesal metaplical process, msl = metaplical shelf-forming lobe, mss = metaplical shelf-spine, pl = posterior lamella of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860486" httpUri="https://zenodo.org/record/3860486/files/figure.png" pageId="67" pageNumber="68">Fig. 16B</figureCitation>
) ……………………………………………………………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1A37F634FAA9F8BE" ID-CoL="TH76" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1120,1289,1795,1821]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nikolajscharffi" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1A37F634FAA9F8BE" box="[1120,1289,1795,1821]" italics="true" pageId="67" pageNumber="68">nikolajscharffi</emphasis>
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<paragraph id="BB59967FC12EFFCB1EEBF672FAD7F821" blockId="67.[188,1399,1861,1923]" pageId="67" pageNumber="68">
<keyLead id="0017354AC12EFFCB1EEBF672FAD7F821" pageId="67" pageNumber="68">
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14 Metaplical shelf-spine of gonopod coxa long, projecting ± mesad (
<figureCitation id="23DD8AFAC12EFFCB1DACF672FBF5F8FC" box="[1019,1109,1861,1887]" captionStart="Fig" captionStartId="39.[187,230,1861,1887]" captionTargetBox="[188,1398,321,1825]" captionTargetId="figure@39.[188,1398,321,1825]" captionTargetPageId="39" captionText="Fig. 20. Chaleponcus nectarinia sp. nov., gonopods. A. Right gonopod, posterior (-lateral) view. B. Right gonopod, anterior view. C. Tip of posterior lamella of telomere. D. Left gonopod, mesal (-posterior) view. E. Right gonopod, mesal view. al = anterior lobe of telomere, cu = cucullus, mfp = metaplical flange process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860494" httpUri="https://zenodo.org/record/3860494/files/figure.png" pageId="67" pageNumber="68">Figs 20</figureCitation>
A–B, 23A, D) ………15 – Metaplical shelf-spine not long, projecting ± mesad ……………………………………………16
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</paragraph>
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<paragraph id="BB59967FC12EFFCB1EEBF69EFAD7F844" blockId="67.[188,1399,1961,2023]" pageId="67" pageNumber="68">
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15 Posterior lamella of telomere divided into a horizontal lobe with spinose edges and a long, straight, distal spine-like part (
<figureCitation id="23DD8AFAC12EFFCB1C37F6FAFD4CF844" box="[608,748,1997,2023]" captionStart="Fig" captionStartId="39.[187,230,1861,1887]" captionTargetBox="[188,1398,321,1825]" captionTargetId="figure@39.[188,1398,321,1825]" captionTargetPageId="39" captionText="Fig. 20. Chaleponcus nectarinia sp. nov., gonopods. A. Right gonopod, posterior (-lateral) view. B. Right gonopod, anterior view. C. Tip of posterior lamella of telomere. D. Left gonopod, mesal (-posterior) view. E. Right gonopod, mesal view. al = anterior lobe of telomere, cu = cucullus, mfp = metaplical flange process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lobe of telomere, ps = proximal spine of solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860494" httpUri="https://zenodo.org/record/3860494/files/figure.png" pageId="67" pageNumber="68">Fig. 20C, E</figureCitation>
) …………………………………… 
<taxonomicName id="7CE6EDFCC12EFFCB1ACEF6F9FAB2F844" ID-CoL="TH73" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1177,1298,1998,2023]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="nectarinia" status="sp. nov.">
<emphasis id="89924A6DC12EFFCB1ACEF6F9FAB2F844" box="[1177,1298,1998,2023]" italics="true" pageId="67" pageNumber="68">nectarinia</emphasis>
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<taxonomicNameLabel id="92A1F716C12EFFCB1B4DF6FAFAD7F844" box="[1306,1399,1997,2023]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF03BFAD7FEEA" blockId="68.[189,1399,267,329]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF03BFAD7FEEA" pageId="68" pageNumber="69">
– Posterior lamella of telomere with strongly laciniate edges but without a long, straight, spine-like part ………………………………………………………………………… 
<taxonomicName id="7CE6EDFCC129FFCC1A08F018FAAFFEEA" ID-CoL="TH7P" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1119,1295,303,329]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vandenspiegeli" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1A08F018FAAFFEEA" box="[1119,1295,303,329]" italics="true" pageId="68" pageNumber="69">vandenspiegeli</emphasis>
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<taxonomicNameLabel id="92A1F716C129FFCC1B4EF018FAD7FEEA" box="[1305,1399,303,329]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="00128EDAC129FFCC1EEAF047FAD6FE73" pageId="68" pageNumber="69">
<paragraph id="BB59967FC129FFCC1EEAF047FAD7FE0D" blockId="68.[189,1399,368,465]" pageId="68" pageNumber="69">
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16 Metaplical shelf of gonopod coxa in two levels between which metaplical shelf-spine fits. Gonopod coxa with a small to inconspicuous lateral process (
<figureCitation id="23DD8AFAC129FFCC1DBAF0A4FBE7FE0E" box="[1005,1095,403,429]" captionStart="Fig" captionStartId="48.[189,232,1823,1849]" captionTargetBox="[189,1398,386,1781]" captionTargetId="figure@48.[189,1398,386,1781]" captionTargetPageId="48" captionText="Fig. 24. Chaleponcus vilici sp. nov., left gonopod. A. Posterior view. B. Mesal view. C. Mesal-anterior view. D. Close-up of metaplical shelf-spine, mesal view. E. Anterior (and slightly apical/ventral) view. F. Telopodite, distal view, showing solenomere and the two distal telomere branches. al = anterior lamella of telomere, cu = cucullus, la = lateral angle of coxa, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior lamella of telomere, slm = solenomere. Scales: 0.1 mm (A–D), 0.01 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.3860502" httpUri="https://zenodo.org/record/3860502/files/figure.png" pageId="68" pageNumber="69">Fig. 24</figureCitation>
) ………… 
<taxonomicName id="7CE6EDFCC129FFCC1A86F0A4FAACFE0E" ID-CoL="TH7Q" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1233,1292,403,429]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vilici" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1A86F0A4FAACFE0E" box="[1233,1292,403,429]" italics="true" pageId="68" pageNumber="69">vilici</emphasis>
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<taxonomicNameLabel id="92A1F716C129FFCC1B40F0A3FAD7FE0D" box="[1303,1399,404,430]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF080FAD6FE73" blockId="68.[189,1399,368,465]" box="[189,1398,438,465]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF080FAD6FE73" box="[189,1398,438,465]" pageId="68" pageNumber="69">– Metaplical shelf in one level. Metaplical shelf-spine different …………………………………17</keyLead>
</paragraph>
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<keyStep id="00128EDAC129FFCC1EEAF0CFFAD7FDDF" pageId="68" pageNumber="69">
<paragraph id="BB59967FC129FFCC1EEAF0CFFAD7FD95" blockId="68.[189,1399,504,636]" pageId="68" pageNumber="69">
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17 Gonopod coxa with distal hook-like extension (
<figureCitation id="23DD8AFAC129FFCC1D77F0CFFC2DFDB1" box="[800,909,504,530]" captionStart="Fig" captionStartId="65.[189,232,1894,1920]" captionTargetBox="[189,1398,325,1850]" captionTargetId="figure@65.[189,1398,325,1850]" captionTargetPageId="65" captionText="Fig. 31. Chaleponcus tintin sp. nov., left gonopod. A. Anterior view. B. Mesal-anterior view. C. Ventral (-anterior) view. D. Posterior (-mesal) view. al = anterior telomeral lamella, cu = cucullus, mfp = metaplical flange process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, plb = basal branch of pl, ps = proximal spine on solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860516" httpUri="https://zenodo.org/record/3860516/files/figure.png" pageId="68" pageNumber="69">Fig. 31A</figureCitation>
). 41 podous rings, body diameter 
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………………………………………………………………………………………… 
<taxonomicName id="7CE6EDFCC129FFCC1A80F32BFAB5FD96" ID-CoL="TH7L" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1239,1301,540,565]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="tintin" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1A80F32BFAB5FD96" box="[1239,1301,540,565]" italics="true" pageId="68" pageNumber="69">tintin</emphasis>
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– Gonopod coxa without distal hook-like extension. Mostly&gt;41 podous rings and/or body diameter&gt; 
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………………………………………………………………………………18
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18 Anal valves without dorsal spines (
<figureCitation id="23DD8AFAC129FFCC1CF0F394FCA6FD1D" box="[679,774,675,702]" captionStart="Fig" captionStartId="12.[189,231,1893,1919]" captionTargetBox="[317,1271,896,1861]" captionTargetId="figure@12.[317,1271,896,1861]" captionTargetPageId="12" captionText="Fig. 6. Telson types in Chaleponcus species. — A. Dorsal and ventral anal valve spines well developed (C. nectarinia sp.nov.).B.Spinesverylong(C.circumvallatussp.nov.).C.Dorsal spinewell developed, ventral spinebarelyindicated(C.mwabvuisp.nov.).D.Nospines(C.teressp.nov.).E.Setiferoustuberclesonravelins (C. nectarinia sp. nov.). F. With setiferous tubercles, not on ravelins (C. mwabvui sp. nov.). Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860464" httpUri="https://zenodo.org/record/3860464/files/figure.png" pageId="68" pageNumber="69">Fig. 6D</figureCitation>
) ……………………………………… 
<taxonomicName id="7CE6EDFCC129FFCC1A84F393FAABFD1E" ID-CoL="TH7H" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1235,1291,676,701]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="teres" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1A84F393FAABFD1E" box="[1235,1291,676,701]" italics="true" pageId="68" pageNumber="69">teres</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B40F394FAD7FD1E" box="[1303,1399,675,701]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF3F0FAD6FD43" blockId="68.[189,1399,675,737]" box="[189,1398,710,737]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF3F0FAD6FD43" box="[189,1398,710,737]" pageId="68" pageNumber="69">– Each anal valve with a dorsal spine ……………………………………………………………19</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC129FFCC1EEAF23FFAD6FCE6" pageId="68" pageNumber="69">
<paragraph id="BB59967FC129FFCC1EEAF23FFAD7FC81" blockId="68.[189,1399,775,837]" box="[189,1399,775,802]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF23FFAD7FC81" box="[189,1399,775,802]" pageId="68" pageNumber="69">
19 Cucullus of gonopod coxa subrectangular in outline (
<figureCitation id="23DD8AFAC129FFCC1DDFF230FC42FC81" box="[904,994,775,802]" captionStart="Fig" captionStartId="54.[187,229,1891,1917]" captionTargetBox="[189,1398,454,1832]" captionTargetId="figure@54.[189,1398,454,1832]" captionTargetPageId="54" captionText="Fig. 26. Chaleponcus hamerae sp. nov., left gonopod. A. Mesal-anterior view. B. Posterior view. C.Anterior view. D. Ventral view. al, al’ = anterior telomeral lamella, cu = cucullus, mf = metaplical flange, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, prl = proplical lobe, pls = spine on pl. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860506" httpUri="https://zenodo.org/record/3860506/files/figure.png" pageId="68" pageNumber="69">Fig. 26</figureCitation>
B–C) ………… 
<taxonomicName id="7CE6EDFCC129FFCC1AF4F230FAAAFC82" ID-CoL="TH6J" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1187,1290,775,801]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="hamerae" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1AF4F230FAAAFC82" box="[1187,1290,775,801]" italics="true" pageId="68" pageNumber="69">hamerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B40F23FFAD7FC81" box="[1303,1399,776,802]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF21CFAD6FCE6" blockId="68.[189,1399,775,837]" box="[189,1398,811,837]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF21CFAD6FCE6" box="[189,1398,811,837]" pageId="68" pageNumber="69">– Outline of cucullus more rounded ……………………………………………………………20</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC129FFCC1EEAF25BFAD7FC0A" pageId="68" pageNumber="69">
<paragraph id="BB59967FC129FFCC1EEAF25BFAD7FC24" blockId="68.[189,1399,876,938]" box="[189,1399,876,903]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF25BFAD7FC24" box="[189,1399,876,903]" pageId="68" pageNumber="69">
20 Anterior lamella of gonopod telomere with long, smooth spine (
<figureCitation id="23DD8AFAC129FFCC1DB1F25BFB99FC25" box="[998,1081,876,902]" captionStart="Fig" captionStartId="63.[189,232,1860,1886]" captionTargetBox="[206,1385,265,1818]" captionTargetId="figure@63.[206,1385,265,1818]" captionTargetPageId="63" captionText="Fig. 30. Chaleponcus howelli sp. nov., left gonopod. A. Mesal-ventral (-anterior) view. B. Posterior view. C. Anterior view. D. Ventral (-anterior) view. al = anterior telomeral lamella, als = spine on al, cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere, sts = spine on telomeral main stem. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860514" httpUri="https://zenodo.org/record/3860514/files/figure.png" pageId="68" pageNumber="69">Fig. 30</figureCitation>
C–D) …… 
<taxonomicName id="7CE6EDFCC129FFCC1AEAF25BFAB3FC25" ID-CoL="TH6N" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1213,1299,876,902]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="howelli" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1AEAF25BFAB3FC25" box="[1213,1299,876,902]" italics="true" pageId="68" pageNumber="69">howelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B4BF25AFAD7FC24" box="[1308,1399,877,903]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF2A7FAD7FC0A" blockId="68.[189,1399,876,938]" box="[189,1399,911,938]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF2A7FAD7FC0A" box="[189,1399,911,938]" pageId="68" pageNumber="69">– No such spine ……………………………………………………………………………………21</keyLead>
</paragraph>
</keyStep>
<keyStep id="00128EDAC129FFCC1EEAF2E6FAD7FBF6" pageId="68" pageNumber="69">
<paragraph id="BB59967FC129FFCC1EEAF2E6FAD7FBAC" blockId="68.[189,1399,977,1109]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF2E6FAD7FBAC" pageId="68" pageNumber="69">
21 Anterior lamella of gonopod telomere divided into two triangular lobes with serrate margins (
<figureCitation id="23DD8AFAC129FFCC1EABF2C3FE2AFBAC" box="[252,394,1012,1039]" captionStart="Fig" captionStartId="56.[187,230,1857,1883]" captionTargetBox="[189,1398,724,1812]" captionTargetId="figure@56.[189,1398,724,1812]" captionTargetPageId="56" captionText="Fig. 27. Chaleponcus termini sp. nov., gonopods. A. Right gonopod, posterior view. B. Right gonopod, anterior view. C. Left telopodite, mesal-ventral view. D. Left coxite, mesal view. E. Right telopodite, anterior-ventral view. al1, al2 = anterior telomeral lobe, cu = cucullus, mf = metaplical flange, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, ps = proximal spine on solenomere. Scales 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3860508" httpUri="https://zenodo.org/record/3860508/files/figure.png" pageId="68" pageNumber="69">Fig. 27B, E</figureCitation>
). Posterior lamella simple …………………………………………… 
<taxonomicName id="7CE6EDFCC129FFCC1AEAF2C2FAB2FBAD" ID-CoL="TH7J" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1213,1298,1013,1038]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="termini" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1AEAF2C2FAB2FBAD" box="[1213,1298,1013,1038]" italics="true" pageId="68" pageNumber="69">termini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B4CF2C2FAD7FBAC" box="[1307,1399,1013,1039]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF52FFAD7FBF6" blockId="68.[189,1399,977,1109]" pageId="68" pageNumber="69">
<keyLead id="0017354AC129FFCC1EEAF52FFAD7FBF6" pageId="68" pageNumber="69">
– Anterior lamella of gonopod telomere simple. Posterior lamella with a slender, spinose branch with a long, thin, bifurcate side branch (
<figureCitation id="23DD8AFAC129FFCC1CF1F50DFCB1FBF7" box="[678,785,1082,1108]" captionStart="Fig" captionStartId="61.[189,233,1824,1850]" captionTargetBox="[190,1398,265,1806]" captionTargetId="figure@61.[189,1398,265,1806]" captionTargetPageId="61" captionText="Fig. 29. Chaleponcus mwabvui sp. nov., right gonopod. A. Posterior view. B. Anterior-mesal view. C. Coxa, tip, anterior view. D. Telopodite, apical-dorsal view. E. Process plp of telomere. al = anterior telomeral lamella, cu = cucullus, mf = metaplical flange, mp = mesal metaplical process, ms = metaplical shelf, mss = metaplical shelf-spine, pl = posterior telomeral lamella, plp1, plp2, posterior telomeral lamella processes, ps = proximal spine on solenomere, slm = solenomere. Scales: 0.1 mm (A–D), 0.05 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.3860512" httpUri="https://zenodo.org/record/3860512/files/figure.png" pageId="68" pageNumber="69">Fig. 29E</figureCitation>
) ………………………………… 
<taxonomicName id="7CE6EDFCC129FFCC1AF5F50DFAACFBF7" ID-CoL="TH6Y" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1186,1292,1082,1108]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="mwabvui" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1AF5F50DFAACFBF7" box="[1186,1292,1082,1108]" italics="true" pageId="68" pageNumber="69">mwabvui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B4FF50CFAD7FBF6" box="[1304,1399,1083,1109]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="F3FCC5F4C129FFCC1EEAF54AFCECFA95" pageId="68" pageNumber="69" type="distribution">
<paragraph id="BB59967FC129FFCC1EEAF54AFE5BFB39" blockId="68.[189,507,1149,1178]" box="[189,507,1149,1178]" pageId="68" pageNumber="69">
<emphasis id="89924A6DC129FFCC1EEAF54AFE5BFB39" bold="true" box="[189,507,1149,1178]" italics="true" pageId="68" pageNumber="69">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF586FCECFA95" blockId="68.[189,1399,1201,1334]" pageId="68" pageNumber="69">
<figureCitation id="23DD8AFAC129FFCC1EEAF586FEB4FB68" box="[189,276,1201,1227]" captionStart="Fig" captionStartId="69.[187,230,1685,1711]" captionTargetBox="[215,1377,822,1616]" captionTargetId="figure@69.[189,1398,752,1635]" captionTargetPageId="69" captionText="Fig. 32. Distribution of species of the Chaleponcus dabagaensis group in Forest Reserves of the Udzungwa Mts. The position of the Forest Reserves in relation to each other approximately corresponds to their true relative positions, cf. Fig. 1. Underlined species acronyms denote species known only from one Forest Reserve. Shared species are shown on lines connecting the circles. – Forest Reserve acronyms: KIG = Kigogo, KK = Kiranzi/Kitunguli, KR = Kisinga/Rugaro, MWA = Mwanihana, NDU = New Dabaga/Ulangambi, US = Udzungwa Scarp, WKS = West Kilombero Scarp. – Species acronyms: BAS = basiliscus, CIR = circumvallatus, DAB = dabagaensis, GRA = gracilior, HAM = hamerae, HOW = howelli, IBI = ibis, KRA = krai, MAL = malleolus, MWB = mwabvui, MWN = mwanihanensis, NEC = nectarinia, NET = netus, NIK = nikolajscharffi, QUA = quasimodo, SCO = scopus, TES = teres, TEM = termini, TIN = tintin, VAN = vandenspiegeli, VIL = vilici." figureDoi="http://doi.org/10.5281/zenodo.3860518" httpUri="https://zenodo.org/record/3860518/files/figure.png" pageId="68" pageNumber="69">Fig. 32</figureCitation>
shows the distribution of the 21 species of the 
<taxonomicName id="7CE6EDFCC129FFCC1D3EF586FB00FB68" authorityName="Kraus" authorityYear="1958" box="[873,1184,1201,1227]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC129FFCC1D3EF586FB00FB68" box="[873,1184,1201,1227]" italics="true" pageId="68" pageNumber="69">Chaleponcus dabagaensis</emphasis>
</taxonomicName>
group plotted on a semi-diagrammatic map of the Forest Reserves (FR) from which species of the group have been collected. Fifteen of the species have been collected in only one FR, with New Dabaga/Ulangambi FR being the richest (6 exclusive species, 11 species in all).
</paragraph>
</subSubSection>
<subSubSection id="F3FCC5F4C129FFC01EEAF46BFED8FEEB" lastPageId="72" lastPageNumber="73" pageId="68" pageNumber="69" type="discussion">
<paragraph id="BB59967FC129FFCC1EEAF46BFEFEFA1E" blockId="68.[189,1399,1372,1469]" pageId="68" pageNumber="69">
Five species are shared between two FR, and 
<taxonomicName id="7CE6EDFCC129FFCC1C85F46AFCFCFAD5" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[722,860,1372,1399]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="gracilior" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1C85F46AFCFCFAD5" box="[722,860,1372,1399]" italics="true" pageId="68" pageNumber="69">C. gracilior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1D34F46AFC1EFAD4" box="[867,958,1373,1399]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
even occurs in four FR, being clearly the most widespread of the species. The easternmost FR, Mwanihana, does not share any species with any other FR.
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF4D3FDE2F956" blockId="68.[189,1399,1508,1781]" pageId="68" pageNumber="69">
As 
<figureCitation id="23DD8AFAC129FFCC1EB3F4D3FE88FA5D" box="[228,296,1508,1534]" captionStart="Fig" captionStartId="2.[189,232,1928,1954]" captionTargetBox="[189,1401,1042,1875]" captionTargetId="figure@2.[190,1399,1040,1873]" captionTargetPageId="2" captionText="Fig. 1. Map of the Udzungwa Mountains, showing the location of the forest reserves (green rings) where the material studied here was collected, as well as names of individual mountains in West Kilombero FR. Based on fig. 1 in Marshall et al. (2010)." figureDoi="http://doi.org/10.5281/zenodo.3860454" httpUri="https://zenodo.org/record/3860454/files/figure.png" pageId="68" pageNumber="69">Fig. 1</figureCitation>
clearly shows, there are several forested areas in the Udzungwa Mts where the 
<taxonomicName id="7CE6EDFCC129FFCC1A95F4D2FAD7FA5D" authorityName="Kraus" authorityYear="1958" box="[1218,1399,1508,1535]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC129FFCC1A95F4D2FAD7FA5D" box="[1218,1399,1508,1535]" italics="true" pageId="68" pageNumber="69">C. dabagaensis</emphasis>
</taxonomicName>
group has not been collected. This is probably due to a collecting bias, since the very comprehensive millipede material from the Udzungwa Mts kept in ZMUC largely derives from the FRs where the 
<taxonomicName id="7CE6EDFCC129FFCC1B0BF71CFEF0F9CB" authorityName="Kraus" authorityYear="1958" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC129FFCC1B0BF71CFEF0F9CB" italics="true" pageId="68" pageNumber="69">C. dabagaensis</emphasis>
</taxonomicName>
group is now known to occur. Additional collection effort, especially in the large southerncentral Matundu forest, is likely to reveal several additional species of the group. Even renewed collecting in the forest reserves already sampled is likely to reveal new species, since three (
<taxonomicName id="7CE6EDFCC129FFCC1B77F7A3FAEAF90D" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1312,1354,1684,1710]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="ibis" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1B77F7A3FAEAF90D" box="[1312,1354,1684,1710]" italics="true" pageId="68" pageNumber="69">ibis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1B02F7A2FF4FF971" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="7CE6EDFCC129FFCC1EABF780FE0CF972" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[252,428,1719,1745]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="vandenspiegeli" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1EABF780FE0CF972" box="[252,428,1719,1745]" italics="true" pageId="68" pageNumber="69">vandenspiegeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1FE5F78FFDABF971" box="[434,523,1720,1746]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="7CE6EDFCC129FFCC1C4FF78FFDF6F972" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[536,598,1720,1745]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="tintin" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1C4FF78FFDF6F972" box="[536,598,1720,1745]" italics="true" pageId="68" pageNumber="69">tintin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1C0BF78FFD17F971" box="[604,695,1720,1746]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
) out of the 21 species are singletons and one (
<taxonomicName id="7CE6EDFCC129FFCC1A9EF780FAD6F972" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1225,1398,1719,1745]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="circumvallatus" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1A9EF780FAD6F972" box="[1225,1398,1719,1745]" italics="true" pageId="68" pageNumber="69">circumvallatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1EEAF7ECFEB9F956" box="[189,281,1755,1781]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
) is a doubleton (
<figureCitation id="23DD8AFAC129FFCC1F89F7EDFD95F956" box="[478,565,1754,1781]" captionStart="Fig" captionStartId="70.[189,232,2000,2026]" captionTargetId="figure@70.[348,1217,1323,1957]" captionTargetPageId="70" captionText="Fig. 33. The number of studied specimens (males) per species." figureDoi="http://doi.org/10.5281/zenodo.3860520" httpUri="https://zenodo.org/record/3860520/files/figure.png" pageId="68" pageNumber="69">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="BB59967FC129FFCC1EEAF62BFAABF860" blockId="68.[189,1399,1820,1987]" pageId="68" pageNumber="69">
The 
<taxonomicName id="7CE6EDFCC129FFCC1EA7F62BFD82F895" authorityName="Kraus" authorityYear="1958" box="[240,546,1820,1846]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="dabagaensis">
<emphasis id="89924A6DC129FFCC1EA7F62BFD82F895" box="[240,546,1820,1846]" italics="true" pageId="68" pageNumber="69">Chaleponcus dabagaensis</emphasis>
</taxonomicName>
group consists of high-altitude species, all material having been collected at 
<quantity id="7C1E3B9AC129FFCC1E8EF608FEDAF8F9" box="[217,378,1855,1882]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" metricValueMax="2.1" metricValueMin="1.39" pageId="68" pageNumber="69" unit="m" value="1745.0" valueMax="2100.0" valueMin="1390.0">1390–2100 m</quantity>
asl, and with the highest species diversity above 
<quantity id="7C1E3B9AC129FFCC1DE2F608FBADF8F9" box="[949,1037,1855,1882]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="68" pageNumber="69" unit="m" value="1700.0">1700 m</quantity>
(
<tableCitation id="F664A3C4C129FFCC1A4CF608FBD6F8FA" box="[1051,1142,1855,1881]" captionStart="Table 2" captionStartId="70.[189,258,268,294]" captionTargetBox="[198,1369,382,1198]" captionTargetId="graphics@70.[189,1401,367,1203]" captionTargetPageId="70" captionText="Table 2. Altitudinal range of species of the Chaleponcus dabagaensis group, based on “maximal ranges”, see Material and methods. No altitude data are available for C. basiliscus and C. ibis." httpUri="http://table.plazi.org/id/EF99C6F7C12BFFCE1EEAF03BFB03FEEA" pageId="68" pageNumber="69" tableUuid="EF99C6F7C12BFFCE1EEAF03BFB03FEEA">Table 2</tableCitation>
). The vast majority of specimens was collected in montane forest, but some species were found in disturbed habitats as well, e.g., the relatively widespread 
<taxonomicName id="7CE6EDFCC129FFCC1C76F6B1FD0AF803" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[545,682,1926,1952]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="gracilior" status="sp. nov.">
<emphasis id="89924A6DC129FFCC1C76F6B1FD0AF803" box="[545,682,1926,1952]" italics="true" pageId="68" pageNumber="69">C. gracilior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C129FFCC1CE6F6B1FCABF803" box="[689,779,1926,1952]" pageId="68" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
Very often several (up to 5) species were represented in a sample, but whether such coexisting species occupy different microhabitats is not known.
</paragraph>
<paragraph id="BB59967FC128FFCD1EECF03AFDDCFE8A" blockId="69.[187,636,268,297]" box="[187,636,268,297]" pageId="69" pageNumber="70">
<emphasis id="89924A6DC128FFCD1EECF03AFDDCFE8A" bold="true" box="[187,636,268,297]" italics="true" pageId="69" pageNumber="70">Two notes on general morphology</emphasis>
</paragraph>
<paragraph id="BB59967FC128FFCD1EECF077FDFFFEF9" blockId="69.[187,607,320,347]" box="[187,607,320,347]" pageId="69" pageNumber="70">
<heading id="E0112113C128FFCD1EECF077FDFFFEF9" bold="true" box="[187,607,320,347]" fontSize="11" level="3" pageId="69" pageNumber="70" reason="3">
<emphasis id="89924A6DC128FFCD1EECF077FDFFFEF9" bold="true" box="[187,607,320,347]" pageId="69" pageNumber="70">Torsion of the gonopod telopodite</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC128FFCD1EECF058FC0DFD99" blockId="69.[187,1399,367,570]" pageId="69" pageNumber="70">
Torsion of the gonopod telopodite is characteristic of the majority of species of 
<taxonomicName id="7CE6EDFCC128FFCD1ADBF058FAE1FE2A" authorityName="Attems" authorityYear="1909" box="[1164,1345,367,393]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
and the related family 
<taxonomicName id="7CE6EDFCC128FFCD1FC4F0A5FDE5FE0F" box="[403,581,402,428]" class="Diplopoda" family="Spirostreptidae" kingdom="Animalia" order="Spirostreptida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
(
<bibRefCitation id="DF77EB8EC128FFCD1C02F0A5FD41FE0F" author="Kraus O." box="[597,737,402,428]" pageId="69" pageNumber="70" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">Kraus 1966</bibRefCitation>
, 
<bibRefCitation id="DF77EB8EC128FFCD1CB9F0A5FC3BFE0F" author="Hoffman R. L." box="[750,923,402,428]" pageId="69" pageNumber="70" pagination="167 - 186" refId="ref30677" refString="Hoffman R. L. 2008. Two new genera of spirostreptid millipeds from central Africa, with a new terminology for male genitalia in the family Spirostreptidae (Diplopoda Spirostreptida). Tropical Zoology 21: 167 - 186." type="journal article" year="2008">Hoffman 2008</bibRefCitation>
) and is important for the function of the gonopod (Barnett &amp; Telford 1996). In all odontopygids examined by me the torsion always begins with an anteriad bend, i.e., the torsion is clockwise when followed from coxa towards the gonopod tip and viewed from a mesal point of view on the RIGHT gonopod. Published drawings by, e.g., 
<bibRefCitation id="DF77EB8EC128FFCD1A8DF0CBFAD6FDB4" author="Kraus O." box="[1242,1398,508,535]" pageId="69" pageNumber="70" pagination="1 - 143" refId="ref30903" refString="Kraus O. 1966. Phylogenie, Chorologie und Systematik der Odontopygoideen (Diplopoda, Spirostreptomorpha). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 512: 1 - 143." type="journal article" year="1966">Kraus (1966)</bibRefCitation>
and 
<bibRefCitation id="DF77EB8EC128FFCD1EBAF328FE73FD99" author="Frederiksen S. B." box="[237,467,543,570]" pageId="69" pageNumber="70" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen (2013a</bibRefCitation>
,b) confirm the generality of this pattern.
</paragraph>
<paragraph id="BB59967FC128FFCD1EECF356FDFBFDD9" blockId="69.[187,603,608,635]" box="[187,603,608,635]" pageId="69" pageNumber="70">
<heading id="E0112113C128FFCD1EECF356FDFBFDD9" bold="true" box="[187,603,608,635]" fontSize="11" level="3" pageId="69" pageNumber="70" reason="3">
<emphasis id="89924A6DC128FFCD1EECF356FDFBFDD9" bold="true" box="[187,603,608,635]" pageId="69" pageNumber="70">Intercalary cuticular microscutes</emphasis>
</heading>
</paragraph>
<paragraph id="BB59967FC128FFC01EECF3A7FED8FEEB" blockId="69.[187,1405,655,717]" lastBlockId="72.[189,1398,267,329]" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
Very small (
<emphasis id="89924A6DC128FFCD1F1CF3A7FEC7FD0A" box="[331,359,656,681]" italics="true" pageId="69" pageNumber="70">ca</emphasis>
. 
<quantity id="7C1E3B9AC128FFCD1F22F3B8FE14FD09" box="[373,436,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="mm" value="1.0">1 µm</quantity>
) circular microstructures were frequently observed between cuticular microscutes on body rings (
<figureCitation id="23DD8AFAC128FFCD1F20F385FE72FD6F" box="[375,466,690,716]" captionStart="Fig" captionStartId="71.[189,232,1966,1992]" captionTargetBox="[189,1398,556,1921]" captionTargetId="figure@71.[189,1398,556,1921]" captionTargetPageId="71" captionText="Fig. 34. Intercalary cuticular microscutes in Chaleponcus spp. A. C. netus sp. nov. B. C. tintin sp. nov. C–D. C. nikolajscharffi sp. nov. Fig. D shows fungus-like outgrowths from scutes. Scales 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3860522" httpUri="https://zenodo.org/record/3860522/files/figure.png" pageId="69" pageNumber="70">Fig. 34</figureCitation>
). Similar structures were reported by 
<bibRefCitation id="DF77EB8EC128FFCD1DC8F385FB75FD6F" author="Akkari N. &amp; Enghoff H." box="[927,1237,690,716]" pageId="69" pageNumber="70" pagination="1 - 24" refId="ref29875" refString="Akkari N. &amp; Enghoff H. 2011. On some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda). ZooKeys 156: 1 - 24. http: // dx. doi. org / 10.3897 / zookeys. 156.2134" type="journal article" year="2011">Akkari &amp; Enghoff (2011)</bibRefCitation>
from several families of the order 
<taxonomicName id="7CE6EDFCC12AFFCF1F97F03CFDF8FE86" box="[448,600,267,293]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
and have also been observed on 
<taxonomicName id="7CE6EDFCC12AFFCF1DA5F03BFEA1FEEA" authority="(Verhoeff, 1893)" baseAuthorityName="Verhoeff" baseAuthorityYear="1893" class="Diplopoda" family="Dorypetalidae" genus="Lusitanipus" kingdom="Animalia" order="Callipodida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="89924A6DC12AFFCF1DA5F03BFB55FE86" box="[1010,1269,267,293]" italics="true" pageId="71" pageNumber="72">Lusitanipus alternans</emphasis>
(Verhoeff, 1893)
</taxonomicName>
(
<taxonomicName id="7CE6EDFCC12AFFCF1F42F019FE04FEEB" box="[277,420,302,328]" class="Diplopoda" kingdom="Animalia" order="Callipodida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="order">Callipodida</taxonomicName>
: 
<taxonomicName id="7CE6EDFCC12AFFCF1FE5F019FDFAFEEA" box="[434,602,302,329]" class="Diplopoda" family="Dorypetalidae" kingdom="Animalia" order="Callipodida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="family">Dorypetalidae</taxonomicName>
) (HE and S. Reboleira unpublished), but have not been recorded from other millipede orders. Such intercalary cuticular microscutes are, however, discernible on fig. 8 of 
<bibRefCitation id="DF77EB8EC12AFFCF1E88F042FE6DFE2C" author="Frederiksen S. B." box="[223,461,373,399]" pageId="71" pageNumber="72" pagination="597 - 600" refId="ref30328" refString="Frederiksen S. B. 2013 a. East African odontopygid millipedes 2: A new, geographically disjunct species of Chaleponcus (Attems 1914) from the Pare Mts, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3636: 597 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3636.4.7" type="journal article" year="2013">Frederiksen (2013a)</bibRefCitation>
(
<taxonomicName id="7CE6EDFCC12AFFCF1FB7F042FC68FE2C" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[480,968,373,399]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="parensis">
<emphasis id="89924A6DC12AFFCF1FB7F042FD45FE2C" box="[480,741,373,399]" italics="true" pageId="71" pageNumber="72">Chaleponcus parensis</emphasis>
Frederiksen, 2013
</taxonomicName>
), as well as on fig. 1 of 
<bibRefCitation id="DF77EB8EC12AFFCF1ABBF042FEBAFE10" author="Frederiksen S. B." pageId="71" pageNumber="72" pagination="59 - 66" refId="ref30389" refString="Frederiksen S. B. 2013 b. East African odontopygid millipedes 3: Two new genera; Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. http: // dx. doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen (2013b)</bibRefCitation>
(
<taxonomicName id="7CE6EDFCC12AFFCF1F7CF0AFFCC5FE11" authority="Frederiksen, 2013 b" authorityName="Frederiksen" authorityYear="2013" box="[299,869,408,434]" class="Diplopoda" family="Odontopygidae" genus="Lamelloramus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="rhombiformis">
<emphasis id="89924A6DC12AFFCF1F7CF0AFFDDBFE11" box="[299,635,408,434]" italics="true" pageId="71" pageNumber="72">Lamelloramus rhombiformis</emphasis>
<bibRefCitation id="DF77EB8EC12AFFCF1CD4F0AFFCC5FE11" author="Frederiksen S. B." box="[643,869,408,434]" pageId="71" pageNumber="72" pagination="59 - 66" refId="ref30389" refString="Frederiksen S. B. 2013 b. East African odontopygid millipedes 3: Two new genera; Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. http: // dx. doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013b</bibRefCitation>
</taxonomicName>
). They have also been observed on a species of the genus 
<taxonomicName id="7CE6EDFCC12AFFCF1F04F08BFD07FE76" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[339,679,443,470]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="89924A6DC12AFFCF1F04F08BFE6AFE75" box="[339,458,444,470]" italics="true" pageId="71" pageNumber="72">Aquattuor</emphasis>
Frederiksen, 2013
</taxonomicName>
, and may be widespread in 
<taxonomicName id="7CE6EDFCC12AFFCF1DBCF08CFB04FE76" authorityName="Attems" authorityYear="1909" box="[1003,1188,443,469]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
. The nature of the microscutes is unknown. In a preparation of 
<taxonomicName id="7CE6EDFCC12AFFCF1C8EF0E8FC06FE5A" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[729,934,479,505]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nikolajscharffi" status="sp. nov.">
<emphasis id="89924A6DC12AFFCF1C8EF0E8FC06FE5A" box="[729,934,479,505]" italics="true" pageId="71" pageNumber="72">C. nikolajscharffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92A1F716C12AFFCF1DE7F0E8FBADFE5A" box="[944,1037,479,505]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
some of the microscutes give rise to projecting, club-shaped outgrowths of a few µm length (
<figureCitation id="23DD8AFAC125FFC01DC8F03CFBA9FE86" box="[927,1033,267,293]" captionStart="Fig" captionStartId="71.[189,232,1966,1992]" captionTargetBox="[189,1398,556,1921]" captionTargetId="figure@71.[189,1398,556,1921]" captionTargetPageId="71" captionText="Fig. 34. Intercalary cuticular microscutes in Chaleponcus spp. A. C. netus sp. nov. B. C. tintin sp. nov. C–D. C. nikolajscharffi sp. nov. Fig. D shows fungus-like outgrowths from scutes. Scales 0.001 mm." figureDoi="http://doi.org/10.5281/zenodo.3860522" httpUri="https://zenodo.org/record/3860522/files/figure.png" pageId="72" pageNumber="73">Fig. 34D</figureCitation>
) which may represent parasitic fungi of a kind?
</paragraph>
</subSubSection>
</treatment>
</document>