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<document ID-CLB-Dataset="3837" ID-DOI="http://dx.doi.org/10.3897/asp.79.e62332" ID-GBIF-Dataset="c6cd0951-6d64-4b2f-b920-bb7ba439eb39" ID-Pensoft-Pub="1864-8312-79-509" ID-Pensoft-UUID="F1742697BB0C564B9BA5036415F063D4" ID-ZooBank="951E28C6EAC0494EBC255DEF1991583B" ModsDocID="1864-8312-79-509" checkinTime="1633953041321" checkinUser="pensoft" docAuthor="Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W. &amp; Harms, Danilo" docDate="2021" docId="32CE03EE0BAB5C5091ACC132BD79FB5F" docLanguage="en" docName="ArthropodSystPhyl 79: 509-533" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 79" docPubDate="2021-10-11" docSource="http://dx.doi.org/10.3897/asp.79.e62332" docTitle="Missulena iugum Greenberg &amp; Huey &amp; Framenau &amp; Harms 2021, sp. nov." docType="treatment" docUuid="D846D8BD-F6FB-4E8E-8C16-BA6574CF0F11" docUuidSource="ZooBank" docVersion="5" id="F1742697BB0C564B9BA5036415F063D4" lastPageNumber="509" masterDocId="F1742697BB0C564B9BA5036415F063D4" masterDocTitle="Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome" masterLastPageNumber="533" masterPageNumber="509" pageNumber="509" updateTime="1732804601173" updateUser="ExternalLinkService">
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<mods:title id="15D24C6C42D799206BE64C86E1234D4D">Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome</mods:title>
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<mods:namePart id="F69D0A7E0F532EEEF237DC40AAAFC03C">Greenberg, Marleen R.</mods:namePart>
<mods:affiliation id="B97F7929371662FA4F88670A23188E08">Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; Fachbereich Biologie, Universitaet Hamburg, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="33DBAEDD70DB3A961E982105B37F4950">Huey, Joel A.</mods:namePart>
<mods:affiliation id="1FB841B9954F81FBDF6D14B9440A2D3A">Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia</mods:affiliation>
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<mods:namePart id="5BDAFEC3095DB6D988C65F4AAEF0EA4B">Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier id="1D1B7C8B05199BEC2A0DE3F11F0EA60F" type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
<mods:affiliation id="CD076919203AE4CDEF2D9B937AE16685">Harry Butler Institute, Murdoch University, 90 South Street, Murdoch 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia &amp; Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="7A9D72DFD238D97112D836B93ECF8161">Harms, Danilo</mods:namePart>
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<mods:affiliation id="C9214E69775A0DC157A060F842D73D7E">Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:title id="DB748109421667F5BC9168B9E7BDBB8C">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="9D512A785276DDE04CDDC76AB1B6874E">2021</mods:date>
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<mods:number id="FBCD4F94BAB2A6DF8921E659A91D37C6">79</mods:number>
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<subSection id="74A75EE0202E87E6F012B42E87ACDBE6" pageId="0" pageNumber="509" type="3. taxonomy">
<treatment id="32CE03EE0BAB5C5091ACC132BD79FB5F" ID-GBIF-Taxon="189469058" LSID="urn:lsid:zoobank.org:act:D846D8BD-F6FB-4E8E-8C16-BA6574CF0F11" httpUri="http://treatment.plazi.org/id/32CE03EE0BAB5C5091ACC132BD79FB5F" lastPageNumber="509" pageId="0" pageNumber="509" scope_family="Actinopodidae" scope_order="Araneae">
<subSubSection id="BD3C791F4E57109F87831587416CC6AF" pageId="0" pageNumber="509" type="nomenclature">
<paragraph id="64679E569F7EE2CE20ABA88DF57C04BA" pageId="0" pageNumber="509">
3.2.
<taxonomicName id="4464DF2E2D3EA81E8E0B19F795EE5913" ID-CoL="98YMT" LSID="http://zoobank.org/D846D8BD-F6FB-4E8E-8C16-BA6574CF0F11" authority="Greenberg &amp; Huey &amp; Framenau &amp; Harms, 2021" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum" status="sp. nov.">Missulena iugum</taxonomicName>
<taxonomicNameLabel id="AF302DBA48AB64793EAEC6ACF9A55DAF" pageId="0" pageNumber="509">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="E395FFE38B8433F3E75FCC4F4F8559A9" pageId="0" pageNumber="509" type="description">
<paragraph id="01E3C5C8FDB251FA2332E677DEF433C2" pageId="0" pageNumber="509">
<figureCitation id="F63B94119AA8AD4A6A28CD773C6A2C29" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">Figs 8</figureCitation>
<figureCitation id="1A62805D5FB5C58C753E70E619D904BF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">, 9</figureCitation>
<figureCitation id="32C3DCE835407C953614F8A559A16F6E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Missulena manningensis sp. nov. and Missulena iugum sp. nov. in Western Australia." figureDoi="10.3897/asp.79.e62332.figure10" httpUri="https://binary.pensoft.net/fig/597116" pageId="0" pageNumber="509">, 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B09632149D5C8E567D07C96F8D283FEA" pageId="0" pageNumber="509" type="type material">
<paragraph id="E3DB52842927BD7B24C290EFDD8A87A5" pageId="0" pageNumber="509">Type material.</paragraph>
<paragraph id="7633D54187A3963063FD94FEF64454DC" pageId="0" pageNumber="509">
<emphasis id="03AA0F1BB1ABD50E5BF48AAA16F5DFC7" bold="true" pageId="0" pageNumber="509">Holotype</emphasis>
: AUSTRALIA - Western Australia • ♂; Mt Ida, 80 km NW of Menzies;
<geoCoordinate id="1927862DF6D3C536234912C5126F5C3A" degrees="29" direction="south" minutes="12.5" orientation="latitude" precision="92" value="-29.208334">29°12.5S</geoCoordinate>
<geoCoordinate id="8B0D86788E5AC929594CF86C3DFC29A7" degrees="120" direction="east" minutes="24.48" orientation="longitude" precision="9" value="120.408">120°24.48E</geoCoordinate>
; 29 Mar. 2012; V. Saffer leg.; pitfall trap; WAM T123110.
<emphasis id="27E0D8473EF7298428C514BFBF99910C" bold="true" pageId="0" pageNumber="509">Paratype</emphasis>
: AUSTRALIA - Western Australia • ♂; same data as for holotype;
<geoCoordinate id="3AA2DAA3BFE557BA09CC3F7F91F7AF70" degrees="29" direction="south" minutes="12.97" orientation="latitude" precision="9" value="-29.216167">29°12.97S</geoCoordinate>
<geoCoordinate id="C4ACDA6DBB160468225B519C08795313" degrees="120" direction="east" minutes="25.43" orientation="longitude" precision="9" value="120.423836">120°25.43E</geoCoordinate>
; 27 July 2008; M. Quinn and G. Murray leg.; pitfall trap; WAM T110243.
</paragraph>
</subSubSection>
<subSubSection id="9E715494915992DFD8AC9CF6AF72B0B1" pageId="0" pageNumber="509" type="diagnosis">
<paragraph id="18E97182005151060DDB8A2AA1FD7545" pageId="0" pageNumber="509">Diagnosis.</paragraph>
<paragraph id="4B6BDC2C9EA9BB7B28049BC05121D677" pageId="0" pageNumber="509">
Males of
<taxonomicName id="223B07C8EAF9C7031A9B3524A03B4823" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum">
<emphasis id="5BB0B595C14BB1B2F5CBF7957357A0B4" italics="true" pageId="0" pageNumber="509">Missulena iugum</emphasis>
</taxonomicName>
sp. nov. share the red chelicerae and pars cephalica with
<taxonomicName id="2F794F9213ABCC2B62E078A158C5EFFC" lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis id="26CB1794F066D6E11887FBFB70912107" italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="91256167B3F177E1D46296D2FE46E936" lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis id="D90DC95C13FDED09ACE66DB1E7EABD43" italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="A88EB01CEB6F1F1626829047E9456E42" lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis id="9723D69730A137A9FCC72006B105F936" italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
,
<taxonomicName id="0879856145DA4D2D905FADD7B7A890AD" lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
<emphasis id="878EC44EB7B140BCAEA2E199D714CD4E" italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
</taxonomicName>
and
<taxonomicName id="4944E03D10EAC8047D158CB7CFCB33C9" lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis id="042DBC4F4D3E07E4D9BBBD6B2855B2B2" italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
that are morphologically most similar. They differ from
<taxonomicName id="96081888934C610289043F95D48EA45D" lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis id="5491C7C64103FDE7538E2433C414871B" italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
by a longer carapace (&gt;3.00 mm;
<taxonomicName id="72B9C762A4941319480C04B28A6B4D49" lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis id="D352BA377B1E90DAE0395E238871B0DC" italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
up to 2.8 mm) and a rastellum with strong, conical spines (simple in
<taxonomicName id="3B4D74AF2631EDDB54D310FBAD820932" lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis id="C9B1BE11487077A4ACDDB9986083B9F5" italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
). Rastellum and cuspules on labium and maxillae stronger than in
<taxonomicName id="DBD78991DD5C4D782905F02738B0912D" lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis id="9E208C8045CC5DB5A028475F3A91EF15" italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
(rastellum: 8-9 spines;
<taxonomicName id="41AF07E32E0ADFD33E99F9CC9DA18868" lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis id="2B8F3A44A4630A0E9A8A9982DD7E6DEB" italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
2-5; cuspules:
<taxonomicName id="059186B036E1F683FA0F5B7B6DDFA4D3" lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis id="38982BA3251B9F10B4CEE5D6EB9C742E" italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
none). Pars cephalica lower than in
<taxonomicName id="CEBFF3B1CE28A5E3DDB537DAF4E574CF" lsidName="M. occatoria" pageId="0" pageNumber="509" rank="species" species="occatoria">
<emphasis id="0EE131CD73AD1A9CE90A21B52087AF31" italics="true" pageId="0" pageNumber="509">M. occatoria</emphasis>
</taxonomicName>
(up to 1.69;
<taxonomicName id="EF2499C2DFB9311B85E75AC0433241CF" lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
<emphasis id="244F0C39FDC2D06F572B28963EC1C1FB" italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
</taxonomicName>
approx. 3.0) and carapace shorter (3.87 long, 4.98 wide;
<taxonomicName id="B876EC60ABF44325EE51024A2A5C900B" lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
<emphasis id="A0E388D4FD7432EA8391C37B311D3625" italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
</taxonomicName>
approx. 5.0 long, 7.0 wide). Differs from
<taxonomicName id="0A58982467AF74E080353CFBAA1FBB24" lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis id="4D60461931DCE08E836C735D9B4A920A" italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="997A5838BD2AA01169A75CD14E702FD2" lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis id="2010AA55AB9896E307F0351673FF5CFA" italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov. by the presence of a ridge in the cheliceral groove (Fig.
<figureCitation id="2E3AFBB8266F5DA29E71116D4A977082" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9G, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6FBE10D6AE007F73BDD3FF63FE92C28C" pageId="0" pageNumber="509" type="description">
<paragraph id="423BA502A48AF7494908303E89291406" pageId="0" pageNumber="509">Description.</paragraph>
<paragraph id="B73A9ACE62D66676C86451ED5B5B0623" pageId="0" pageNumber="509">
<emphasis id="84F285EDC99621AD8CC7069DE189C5B5" bold="true" pageId="0" pageNumber="509">MALE</emphasis>
(based on holotype; WAM T123110). Total length 9.89.
<emphasis id="9718CF3EAF2A616E496BB6CCFF012266" bold="true" pageId="0" pageNumber="509">Colour</emphasis>
: pars cephalica and chelicerae orange (Fig.
<figureCitation id="A53AE9F31EA3CEBC92C69559615F5B1E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8C</figureCitation>
); a slim, black ring surrounding the PME (Fig.
<figureCitation id="C1344142D69ABA6FC568DF215650D6C9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8G</figureCitation>
); pars thoracica brown with a light, metallic blue sheen (Fig.
<figureCitation id="BB9CFEE4E068F001B0716096A455D23C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8C</figureCitation>
); abdomen greyish with a very strong, metallic blue and green sheen on the dorsal side (Fig.
<figureCitation id="4EA977DEF2B928A923CF161BE2373439" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8D</figureCitation>
), ventrally more brownish with a faint hint of purple (Fig.
<figureCitation id="B98B6D7ECE101DF98FCAE002B84E16D0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8E</figureCitation>
); sternum orange, slightly fading into olive with 8 sigilla in different shades of orange (Fig.
<figureCitation id="173A789164BF3D47588FAF6A0DD87BA5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9C</figureCitation>
); labium and maxillae orange with a slightly darker reddish-orange spot on the base of labium (Fig.
<figureCitation id="4519E51F6B59079C5DB8456963E70EDE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8F</figureCitation>
); legs olive fading into light brown ventrally, dorsally brown (Fig.
<figureCitation id="9F15FD50E1BD3BFF7B0E9A3486C0D2AE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8A-B</figureCitation>
); spinnerets beige-coloured (Fig.
<figureCitation id="3002423D4021043D74F546424DA50BD5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8E</figureCitation>
).
<emphasis id="366759C336CA7E90D843D3865858CD81" bold="true" pageId="0" pageNumber="509">Carapace</emphasis>
: 3.87 long, 4.98 wide and 1.67 high; clypeus 0.34; pars cephalica covers 2.34 of its length, is highly elevated and faintly granulated with very few setae (Fig.
<figureCitation id="070988AD5A6728D4D283B14C98E6115D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9H</figureCitation>
); pars thoracica also granulated with bands of faint, radial fissures and with two notches close to the abdomen (Fig.
<figureCitation id="67C568C4AA1DE415ECCFA154FF825ED4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8C</figureCitation>
).
<emphasis id="76AE2A9B97CA81ADE0C3D929CD76A2BE" bold="true" pageId="0" pageNumber="509">Eyes</emphasis>
: OQ 3.7 times wider than long; outer width of each eye pair AME 0.53, ALE 2.54, PME 1.66 and PLE 2.2; diameter of AME 0.2, ALE 0.21, PME 0.11, PLE 0.19; anterior eyes slightly recurved; posterior eyes strongly recurved (Fig.
<figureCitation id="12671B231E1D0D497444BEEEEB76FF04" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8G</figureCitation>
).
<emphasis id="8233555A7B6A517866131CF91EE407F7" bold="true" pageId="0" pageNumber="509">Chelicerae</emphasis>
: 2.28 long and 1.55 wide on the base; edges rounded and curved with the widest point being 1.57 very close to the chelicerae base (Fig.
<figureCitation id="CD3C80B15B6B66AAFA68E40C20C71B5B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8C</figureCitation>
); few setae along the inner margin and slightly more evenly spread setae along the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 8 strong, conical spines (Fig.
<figureCitation id="C9F93CE9955276BCA409180468853B72" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8H</figureCitation>
); approx. 20 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth with each a ridge along the row and a general cheliceral teeth area in between those 2 clear rows (Fig.
<figureCitation id="6FCB648C26DA417C9B9E4F49F3A51E78" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9G, I</figureCitation>
); prolateral row with approx. 10 teeth; retrolateral row with 3 teeth (paratype has 4 teeth); intermediate area with 8 small teeth.
<emphasis id="5E299F43C04258B4F43C415A732AF162" bold="true" pageId="0" pageNumber="509">Maxillae</emphasis>
: 2.24 long and 1.52 wide; at least 85-100 weakly developed cuspules along entire anterior margin (Fig.
<figureCitation id="DDF1D5AC08577CB9E1593D350E4BA2D9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8F</figureCitation>
).
<emphasis id="DAC0F871BAFCFC886FA90488599069B8" bold="true" pageId="0" pageNumber="509">Labium</emphasis>
: 0.94 long and 0.91 wide on the base; conical; at least 35 weakly developed cuspules anteriorly (Fig.
<figureCitation id="736734BDE7D37EDFE7A574DF38AAA08C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8F</figureCitation>
); labiosternal junction visible (Fig.
<figureCitation id="C9DB2062E1FD23F26DA19C3F207EFDD1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9C</figureCitation>
).
<emphasis id="AC2BA034E510819F38C652E39640A704" bold="true" pageId="0" pageNumber="509">Sternum</emphasis>
: 2.84 long and 2.77 wide; ovoid (Fig.
<figureCitation id="896C4B840EAF963FE753380BD79A9AB2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9C</figureCitation>
); setae of various length somewhat densely but disordered along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair small and hardly visible, second pair smallest (roughly half the size of the first pair) and circular, third pair larger than first and second pair (roughly 2 times bigger than first pair) in the shape of a drop, and posterior pair biggest (roughly 2.5 times the size of the third pair) in the shape of an elongated drop, all sigilla slightly depressed.
<emphasis id="E6ABD7AD0F7170790BEBFC094F2A7413" bold="true" pageId="0" pageNumber="509">Abdomen</emphasis>
: 3.7 long and 3.41 wide; shape of a rounded trapezoid (Fig.
<figureCitation id="3133A725E106F4BB0CEA360DD7F198F7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8D</figureCitation>
); 4 spinnerets, PLS 0.88 long, 0.55 wide; PMS 0.45 long, 0.19 wide (Fig.
<figureCitation id="5CD8EF8AE2B3DB36D3F2A053D96178FE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Missulena iugum sp. nov. Male holotype (WAMT 123110): A habitus, dorsal view; B ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G eye region, dorsal view; H rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509">8E</figureCitation>
).
<emphasis id="36100A91F8A1BB7F993979B1F533A360" bold="true" pageId="0" pageNumber="509">Pedipalp</emphasis>
: length of trochanter 1.37, femur 4.17, patella 1.88, tibia 4.22, tarsus 0.75; all segments with setae, tibia ventrally densely covered with comparably long setae (Fig.
<figureCitation id="B0FB6DD9ADA0BD434B41F7A657351EC8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9D-F</figureCitation>
); tibia rather thin and slightly recurved from lateral view, 1.00 wide on the widest point from prolateral and 1.08 from dorsal view (Fig.
<figureCitation id="C7AF75FEE24B4F9DB1012873B6A79D05" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9D-F</figureCitation>
); bulb roughly pyriform, two strongly sclerotized sections connected by a velar median structure (
<normalizedToken id="3B5EF765AEA4B1EEB45B01997C3A8543" originalValue="“haematodocha”">&quot;haematodocha&quot;</normalizedToken>
); embolus rather short with an intumescence in proximal region; tip of embolus triangular with a small lamella and a tooth best visible from prolateral view (paratype, Fig.
<figureCitation id="AB6AA97D47B31AFB69B660E1BFF4DA8E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9J, K</figureCitation>
).
<emphasis id="257E5833BF023BBA5ACB55A0ECC7B827" bold="true" pageId="0" pageNumber="509">Legs</emphasis>
: brown setae of various sizes on all sides of the legs and bent towards the exterior; some comparably long setae dorsally on most segments double the length of the other setae; ventral preening comb on tarsi and metatarsi III and IV.
<emphasis id="060C8498BAB5B5C9F1A84C8AC7CB6312" bold="true" pageId="0" pageNumber="509">Leg spination</emphasis>
: leg I: tibia rl1, v13, pl0, d0; metatarsus rl2, v10, pl1, d0; tarsus rl2, v7, pl5, d0; leg II: tibia rl1, v11, pl1, d0; metatarsus rl2, v11, pl0, d0; tarsus rl5, v9, pl1, d0; leg III: tibia rl6, v5, pl5, d6; metatarsus rl5, v7, pl5, d9; tarsus rl10, v9, pl2, d6; leg IV: tibia rl1, v10, pl1, d1; metatarsus rl1, v9, pl3, d2; tarsus rl14, v13, pl5, d5; patella I with 9 spines spread out prolaterally and three spine ventrally in a vertical row; patella II two spines ventrally in a vertical row; patella III with 27 spines spread out dorsally and prolaterally, 4 spines retrolaterally (Fig.
<figureCitation id="67FCD3F7E967FC06DC4D661D38A12EAF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9A</figureCitation>
) and 3 spines ventrally in a vertical row; patella IV with approx. 7 spines on the dorsal side (Fig.
<figureCitation id="62B0DB40C545583380FEDBF5BBD69868" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Missulena iugum sp. nov. Male holotype (WAMT 123110): A patella III, dorsal view; B patella IV, dorsal view; C sternum, ventral view; D right pedipalp, retrolateral view; E same, ventral view; F same, prolateral view; G cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove; H carapace, lateral view; I pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT 110243), right pedipalp (left pedipalp not complete): J embolus with embolar tooth, prolateral-dorsal view; K embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40 µm; K 50 µm." figureDoi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509">9B</figureCitation>
), approx. 6 spines on the prolateral side, all very small, and 3 spines ventrally in a vertical row.
<emphasis id="7B617E451EA5FA988A73353259DF4D4B" bold="true" pageId="0" pageNumber="509">Leg measurement</emphasis>
: Leg I: femur 4.2, patella 1.63, tibia 3.06, metatarsus 2.76, tarsus 1.58, total 13.23. Leg II: femur 3.47, patella 1.53, tibia 2.66, metatarsus 2.24, tarsus 1.48, total 11.38. Leg III: femur 3.07, patella 1.35, tibia 1.88, metatarsus 2.37, tarsus 1.52, total 10.19. Leg IV: femur 3.91, patella 1.59, tibia 2.99, metatarsus 2.78, tarsus 1.64, total 12.91. Formula: 1&gt;4&gt;2&gt;3.
</paragraph>
<caption id="488E4E41EFCE9D2ED8E9435F5889C22F" doi="10.3897/asp.79.e62332.figure8" httpUri="https://binary.pensoft.net/fig/597114" pageId="0" pageNumber="509" start="Figure 8" startId="F8">
<paragraph id="C93B3F6F5B5D7ACC5354DDF2C76AEEC3" pageId="0" pageNumber="509">
<emphasis id="4AACDB3D89C74305224191FD57896D6C" bold="true" pageId="0" pageNumber="509">Figure 8.</emphasis>
<taxonomicName id="20D9FE26BC34A4F664CB11272E0E7BC6" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum">
<emphasis id="495DB7F480AA8556B64C7827DF76607A" italics="true" pageId="0" pageNumber="509">Missulena iugum</emphasis>
</taxonomicName>
sp. nov. Male holotype (WAMT123110):
<emphasis id="A5EB925F61841CE6D0A0DAF5919792D9" bold="true" pageId="0" pageNumber="509">A</emphasis>
habitus, dorsal view;
<emphasis id="053C2AC9D8F863F481DF3FF05C79C944" bold="true" pageId="0" pageNumber="509">B</emphasis>
ventral view;
<emphasis id="0B76BBE3D02C70A5CDFCCA561239AEE8" bold="true" pageId="0" pageNumber="509">C</emphasis>
carapace, dorsal view;
<emphasis id="33226CD04D28012D27DE0ED7BB39AB46" bold="true" pageId="0" pageNumber="509">D</emphasis>
abdomen, dorsal view;
<emphasis id="5CDA0D398A3F35295F09ACF6696A8443" bold="true" pageId="0" pageNumber="509">E</emphasis>
same, ventral view;
<emphasis id="7DD88BC6C38659BF31629195CFC68A34" bold="true" pageId="0" pageNumber="509">F</emphasis>
maxillae, labium, and chelicerae, ventral view;
<emphasis id="850B8F7A6C68606D382D024508C841DE" bold="true" pageId="0" pageNumber="509">G</emphasis>
eye region, dorsal view;
<emphasis id="278C5CDEDAD8F2AED4D71561C5D93EAC" bold="true" pageId="0" pageNumber="509">H</emphasis>
rastellum, anterior view. Scale bars: A, B 4.0 mm; C-H 2.0 mm.
</paragraph>
</caption>
<caption id="09CBC44F3772C4645409EFD829CD9FD3" doi="10.3897/asp.79.e62332.figure9" httpUri="https://binary.pensoft.net/fig/597115" pageId="0" pageNumber="509" start="Figure 9" startId="F9">
<paragraph id="C680B7F2F8B976C5172A0717991225A4" pageId="0" pageNumber="509">
<emphasis id="1C31FCF5D4D5BA0848CBBCE6EBD85940" bold="true" pageId="0" pageNumber="509">Figure 9.</emphasis>
<taxonomicName id="D1539E36E62E450D5F80DE8B1DE66329" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum">
<emphasis id="AEB83DAB7BBD5E2934C5EFE5604E0794" italics="true" pageId="0" pageNumber="509">Missulena iugum</emphasis>
</taxonomicName>
sp. nov. Male holotype (WAMT123110):
<emphasis id="0A647FF16E17B9A547B596A9FE65DA0F" bold="true" pageId="0" pageNumber="509">A</emphasis>
patella III, dorsal view;
<emphasis id="80AD5DEDB1342307210A20B5A1B7A27F" bold="true" pageId="0" pageNumber="509">B</emphasis>
patella IV, dorsal view;
<emphasis id="7C4BA5C628A1448EE140341DC2D67D35" bold="true" pageId="0" pageNumber="509">C</emphasis>
sternum, ventral view;
<emphasis id="4D20485D26D1C10D7BD267ABCE28BC52" bold="true" pageId="0" pageNumber="509">D</emphasis>
right pedipalp, retrolateral view;
<emphasis id="8B7AB664BE69FE158D6DD4A0A46960D1" bold="true" pageId="0" pageNumber="509">E</emphasis>
same, ventral view;
<emphasis id="97E34F2909F143FBF76FC653AF4A7A23" bold="true" pageId="0" pageNumber="509">F</emphasis>
same, prolateral view;
<emphasis id="4E33DCACB33616A6E6E66A89FC015B5D" bold="true" pageId="0" pageNumber="509">G</emphasis>
cheliceral teeth in cheliceral groove, arrows pointing to the ridge on both sides of the groove;
<emphasis id="7BB638C83EAC644BBA7A49607CD2B32D" bold="true" pageId="0" pageNumber="509">H</emphasis>
carapace, lateral view;
<emphasis id="097D1F91028FE964701E7805E9DB3A2C" bold="true" pageId="0" pageNumber="509">I</emphasis>
pattern of cheliceral teeth in cheliceral groove. Male paratype (WAMT110243), right pedipalp (left pedipalp not complete):
<emphasis id="1532E3223C9FF3A5E056EDADB8ADD93F" bold="true" pageId="0" pageNumber="509">J</emphasis>
embolus with embolar tooth, prolateral-dorsal view;
<emphasis id="64182D866CA57FAD127B1E784978E00F" bold="true" pageId="0" pageNumber="509">K</emphasis>
embolus with lamella and tooth, retrolateral view. Scale bars: A, B, G, I 1.0 mm; C-F, H 2.0 mm; J 40
<normalizedToken id="9F9A3B5DDB5CC296766EA1BF3BE68CA7" originalValue="µm">µm</normalizedToken>
; K 50
<normalizedToken id="8DECDD6033776B9A630436CAFA335166" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="67DB895D8BA7274BA9DCF03A148D95A0" pageId="0" pageNumber="509" type="etymology">
<paragraph id="B4FD3D7AF52E87B46249043EB7E321D9" pageId="0" pageNumber="509">Etymology.</paragraph>
<paragraph id="7744132707263B82049A0966B4BD951C" pageId="0" pageNumber="509">
The specific epithet is a Latin noun (
<taxonomicName id="3AEA8DDB4E612D28FA316CEF54080ED6" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum">
<emphasis id="E9CD44C5C9A62EB1EE6767D1D197ADAD" italics="true" pageId="0" pageNumber="509">Missulena iugum</emphasis>
</taxonomicName>
= ridge) in apposition, referring to the strongly developed ridges along the cheliceral groove of the males.
</paragraph>
</subSubSection>
<subSubSection id="91F978DD3FC14CD752A4E00B95D9FBCF" pageId="0" pageNumber="509" type="distribution">
<paragraph id="DA553629F71029D104A31DD7CB42189D" pageId="0" pageNumber="509">Distribution.</paragraph>
<paragraph id="172358D3CC2CE661AF93CA474A614600" pageId="0" pageNumber="509">
Known only from the Mt Ida region approximately 16 km east of Ularring in the Goldfields region of Western Australia (Fig.
<figureCitation id="06232D92786EBDA513B3174A32DB461B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Missulena manningensis sp. nov. and Missulena iugum sp. nov. in Western Australia." figureDoi="10.3897/asp.79.e62332.figure10" httpUri="https://binary.pensoft.net/fig/597116" pageId="0" pageNumber="509">10</figureCitation>
). The habitat of the holotype comprises
<taxonomicName id="84AFA4A8E135BB124DBEEFB101015E3A" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" lsidName="Acacia" order="Fabales" pageId="0" pageNumber="509" phylum="Tracheophyta" rank="genus">
<emphasis id="ADC90B17EECE6F5570D0AAF24D2DEDAC" italics="true" pageId="0" pageNumber="509">Acacia</emphasis>
</taxonomicName>
shrubland.
</paragraph>
<caption id="1B19B313B3191411782EB16F8D11F6C2" doi="10.3897/asp.79.e62332.figure10" httpUri="https://binary.pensoft.net/fig/597116" pageId="0" pageNumber="509" start="Figure 10" startId="F10">
<paragraph id="AD4B502E9913F5391F5C7B2306555E97" pageId="0" pageNumber="509">
<emphasis id="E640124D4B3B29ED46338BA2A75C46DE" bold="true" pageId="0" pageNumber="509">Figure 10.</emphasis>
Distribution records of
<taxonomicName id="FEE6B54ED405BD2BE611979A2CF9F560" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena manningensis" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="manningensis">
<emphasis id="13FDAE5A11DD4E7416FE0357DEA68DDB" italics="true" pageId="0" pageNumber="509">Missulena manningensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="D7A5FA65CD7112FBC05736EBDF1E1FFE" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena iugum" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="iugum">
<emphasis id="443A8BC3B5300C24A479235D28A64C0A" italics="true" pageId="0" pageNumber="509">Missulena iugum</emphasis>
</taxonomicName>
sp. nov. in Western Australia.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>