treatments-xml/data/32/2A/02/322A023EFFA2345CBB803CD1FB4964A2.xml
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<document id="C608F067C09F2D0D8D02B6368378466E" ID-CLB-Dataset="55321" ID-DOI="10.11646/zootaxa.5222.6.2" ID-GBIF-Dataset="23334aaa-4031-4b4b-8c62-04704c0f9c01" ID-ISSN="1175-5326" ID-Zenodo-Dep="7477364" ID-ZooBank="6A24E78D-3474-4210-A35A-21A777C4E8CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671804985563" checkinUser="plazi" docAuthor="Abstract, Nantasak Pinkaew" docDate="2022" docId="322A023EFFA2345CBB803CD1FB4964A2" docLanguage="en" docName="zootaxa.5222.6.2.pdf" docOrigin="Zootaxa 5222 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thymioptila clavata Abstract 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="536" masterDocId="CE137A46FFA3345EBB173E36FFA4637F" masterDocTitle="A new species of genus Thymioptila Diakonoff (Lepidoptera: Tortricidae) from Thailand and a new record, Thymiotila oedalea (Meyrick, 1909), from Laos" masterLastPageNumber="544" masterPageNumber="534" pageNumber="535" updateTime="1699413927843" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="417AB1E3E37EB7532A653CD06BE28EB2">A new species of genus Thymioptila Diakonoff (Lepidoptera: Tortricidae) from Thailand and a new record, Thymiotila oedalea (Meyrick, 1909), from Laos</mods:title>
</mods:titleInfo>
<mods:name id="9D4588683DE29F2EB7115ACEED91D529" type="personal">
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<mods:namePart id="FFE218AC608BF16953A7B78845CD8A72">Abstract, Nantasak Pinkaew</mods:namePart>
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<mods:title id="4718EA7359813F360D07BAD289F9E865">Zootaxa</mods:title>
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<mods:part id="FF3D17171DF5238D26E6050107DFBEC7">
<mods:date id="5F367F2618CB29EEE357C14A58AE76F5">2022</mods:date>
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<mods:number id="9F3E1AA264CC90C9668F898FDBF64FC8">2022-12-23</mods:number>
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<mods:identifier id="98AC4DF82288729D4877727347681635" type="DOI">10.11646/zootaxa.5222.6.2</mods:identifier>
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<treatment id="322A023EFFA2345CBB803CD1FB4964A2" ID-DOI="http://doi.org/10.5281/zenodo.7477372" ID-GBIF-Taxon="207932767" ID-Zenodo-Dep="7477372" LSID="urn:lsid:plazi:treatment:322A023EFFA2345CBB803CD1FB4964A2" httpUri="http://treatment.plazi.org/id/322A023EFFA2345CBB803CD1FB4964A2" lastPageId="2" lastPageNumber="536" pageId="1" pageNumber="535">
<subSubSection id="F299E0A3FFA2345FBB803CD1FCD1605F" pageId="1" pageNumber="535" type="nomenclature">
<paragraph id="BA3CB328FFA2345FBB803CD1FE48607E" blockId="1.[151,885,743,837]" box="[151,492,743,770]" pageId="1" pageNumber="535">
<heading id="E1740444FFA2345FBB803CD1FE48607E" bold="true" box="[151,492,743,770]" fontSize="11" level="1" pageId="1" pageNumber="535" reason="1">
<emphasis id="88F76F3AFFA2345FBB803CD1FE48607E" bold="true" box="[151,492,743,770]" pageId="1" pageNumber="535">
<taxonomicName id="7D83C8ABFFA2345FBB803CD1FE23607D" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[151,391,743,770]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA2345FBB803CD1FE23607D" bold="true" box="[151,391,743,770]" italics="true" pageId="1" pageNumber="535">Thymioptila clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93C4D241FFA2345FBA983CD1FE48607E" box="[399,492,743,769]" pageId="1" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid: zoobank.org:act:
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<subSubSection id="F299E0A3FFA2345FBB803D1CFE55603A" box="[151,497,810,837]" pageId="1" pageNumber="535" type="description">
<paragraph id="BA3CB328FFA2345FBB803D1CFE55603A" blockId="1.[151,885,743,837]" box="[151,497,810,837]" pageId="1" pageNumber="535">
(
<figureCitation id="22B8AFADFFA2345FBBB63D1CFEAF603B" box="[161,267,810,836]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 24" captionStartId-0="1.[151,250,1788,1812]" captionStartId-1="3.[151,264,1915,1939]" captionTargetBox-0="[151,1435,1112,1764]" captionTargetBox-1="[354,1236,207,1850]" captionTargetId-0="figure-292@1.[151,1435,1112,1764]" captionTargetId-1="figure-26@3.[321,1266,181,1891]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURE 1. Living male specimen of Thymioptila clavata sp. nov. (paratype, np 11987) with natural resting posture." captionText-1="FIGURES 24. Wing pattern of Thymioptila spp. 2. T. clavata sp. nov. (holotype). 3. T. oedalea (male). 4. T. oedalea (female). Scale bars = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7477368" figureDoi-1="http://doi.org/10.5281/zenodo.7477370" httpUri-0="https://zenodo.org/record/7477368/files/figure.png" httpUri-1="https://zenodo.org/record/7477370/files/figure.png" pageId="1" pageNumber="535">Figs. 12</figureCitation>
,
<figureCitation id="22B8AFADFFA2345FBA003D1CFE83603A" box="[279,295,810,837]" captionStart="FIGURES 57" captionStartId="4.[151,264,1993,2017]" captionTargetBox="[164,1423,709,1970]" captionTargetId="figure-188@4.[164,1423,709,1970]" captionTargetPageId="4" captionText="FIGURES 57. Head of Thymioptila spp. 5. T. clavata sp. nov. (holotype). 6. T. oedalea (male). 7. T. oedalea (female)." figureDoi="http://doi.org/10.5281/zenodo.7477374" httpUri="https://zenodo.org/record/7477374/files/figure.png" pageId="1" pageNumber="535">5</figureCitation>
,
<figureCitation id="22B8AFADFFA2345FBA243D1CFEC9603B" box="[307,365,810,836]" captionStart-0="FIGURE 8" captionStart-1="FIGURES 910" captionStartId-0="5.[151,250,881,905]" captionStartId-1="5.[151,264,1951,1975]" captionTargetBox-0="[151,1435,181,857]" captionTargetBox-1="[151,1435,929,1927]" captionTargetId-0="figure-26@5.[151,1435,181,857]" captionTargetId-1="figure-44@5.[151,1435,929,1927]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 8. A tuft of raised scales on forewing of Thymioptila clavata sp. nov. (holotype)." captionText-1="FIGURES 910. Modified scales on anal area of hindwing of Thymioptila clavata sp. nov. (holotype). 9. A tuft of hair pencils tuft in anal fold and dense of long scales along 3A (upper side). 10. A row of long hair pencils along 3A (underside)." figureDoi-0="http://doi.org/10.5281/zenodo.7477376" figureDoi-1="http://doi.org/10.5281/zenodo.7477380" httpUri-0="https://zenodo.org/record/7477376/files/figure.png" httpUri-1="https://zenodo.org/record/7477380/files/figure.png" pageId="1" pageNumber="535">810</figureCitation>
,
<figureCitation id="22B8AFADFFA2345FBA6E3D1CFE32603B" box="[377,406,810,836]" captionStart="FIGURES 1112" captionStartId="6.[151,264,1157,1181]" captionTargetBox="[151,1435,181,1133]" captionTargetId="figure-15@6.[151,1435,181,1133]" captionTargetPageId="6" captionText="FIGURES 1112. Sex scales on hind leg of Thymioptila spp. 11. T. clavata sp. nov. (holotype). 12. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477384" httpUri="https://zenodo.org/record/7477384/files/figure.png" pageId="1" pageNumber="535">11</figureCitation>
,
<figureCitation id="22B8AFADFFA2345FBAB53D1CFE64603B" box="[418,448,810,836]" captionStart="FIGURES 1314" captionStartId="6.[151,264,1951,1975]" captionTargetBox="[167,1429,1206,1909]" captionTargetId="figure-37@6.[151,1435,1206,1928]" captionTargetPageId="6" captionText="FIGURES 1314. Sclerotized plates on 8th tergum (left) and sternum (right) of Thymioptila spp. 13. T. clavata sp. nov. (holotype). 14. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477388" httpUri="https://zenodo.org/record/7477388/files/figure.png" pageId="1" pageNumber="535">13</figureCitation>
,
<figureCitation id="22B8AFADFFA2345FBADB3D1CFE4D603A" box="[460,489,810,837]" captionStart="FIGURES 1516" captionStartId="7.[151,264,2006,2030]" captionTargetBox="[179,1406,195,1969]" captionTargetId="figure-26@7.[151,1435,181,1982]" captionTargetPageId="7" captionText="FIGURES 1516. Male genitalia of Thymioptila spp. 15. T. clavata sp. nov. (holotype). 16. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477392" httpUri="https://zenodo.org/record/7477392/files/figure.png" pageId="1" pageNumber="535">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F299E0A3FFA2345FBB803D44FED6673F" pageId="1" pageNumber="535" type="materials_examined">
<paragraph id="BA3CB328FFA2345FBB803D44FC1E60CF" blockId="1.[151,1436,881,1088]" pageId="1" pageNumber="535">
<emphasis id="88F76F3AFFA2345FBB803D44FEE360F4" bold="true" box="[151,327,881,908]" pageId="1" pageNumber="535">Type Material.</emphasis>
<materialsCitation id="0AEBB975FFA2345FBA5A3D44FC1E60CF" ID-GBIF-Occurrence="4003333305" collectingDate="2017-02-27" collectionCode="KKIC" collectorName="Pang Sida N. P." country="Thailand" elevation="260" latitude="14.015" location="Sa Kaeo Prov." longLatPrecision="21" longitude="102.19583" pageId="1" pageNumber="535" specimenCount="1" stateProvince="Sa Kaeo" typeStatus="holotype">
<typeStatus id="65380D8AFFA2345FBA5A3D44FE1C60F3" box="[333,440,882,908]" pageId="1" pageNumber="535" type="holotype">Holotype</typeStatus>
, ♁.
<collectingCountry id="C294F3B8FFA2345FBAF33D44FDDC60F3" box="[484,632,882,908]" name="Thailand" pageId="1" pageNumber="535">THAILAND</collectingCountry>
;
<collectingRegion id="78477DCAFFA2345FB9953D44FC8760F3" box="[642,803,882,908]" country="Thailand" name="Sa Kaeo" pageId="1" pageNumber="535">Sa Kaeo Prov.</collectingRegion>
,
<collectorName id="1776D6FEFFA2345FB83A3D44FC7060F3" box="[813,980,882,908]" pageId="1" pageNumber="535">Pang Sida N.P.</collectorName>
,
<geoCoordinate id="DFB7D5EFFFA2345FB8C83D44FBC660F1" box="[991,1122,882,910]" degrees="14" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="535" precision="15" seconds="54" value="14.015">
14
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00ʹ54ʺN
</geoCoordinate>
,
<geoCoordinate id="DFB7D5EFFFA2345FBF7A3D44FB5E60F1" box="[1133,1274,882,910]" degrees="102" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="535" precision="15" seconds="45" value="102.19583">
102
<superScript id="4DF61E60FFA2345FBF803D44FB3B60FF" attach="none" box="[1175,1183,882,896]" fontSize="6" pageId="1" pageNumber="535">o</superScript>
11ʹ45ʺE
</geoCoordinate>
, alt.
<quantity id="7D7B1ECDFFA2345FBE3A3D44FAD160F3" box="[1325,1397,882,908]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="535" unit="m" value="260.0">
<elevation id="31AE541BFFA2345FBE3A3D44FAD160F3" box="[1325,1397,882,908]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="535" unit="m" value="260.0">260 m</elevation>
</quantity>
,
<date id="CE3D95E8FFA2345FBE973D44FEAE60CF" pageId="1" pageNumber="535" value="2017-02-27">
<collectingDate id="DE796C00FFA2345FBE973D44FEAE60CF" pageId="1" pageNumber="535" value="2017-02-27">27 Feb. 2017</collectingDate>
</date>
, Pinkaew
<emphasis id="88F76F3AFFA2345FBA6A3DA1FE0A60CF" box="[381,430,918,944]" italics="true" pageId="1" pageNumber="535">et al</emphasis>
. leg., np9443 (genitalia slide NP3962);
<collectionCode id="DC922BEDFFA2345FB8793DA0FC1260CF" box="[878,950,918,944]" pageId="1" pageNumber="535">KKIC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="BA3CB328FFA2345FBBD03D8CFED6673F" blockId="1.[151,1436,881,1088]" pageId="1" pageNumber="535">
<materialsCitation id="0AEBB975FFA2345FBBD03D8CFD136086" ID-GBIF-Occurrence="4003333302" collectingDate="2008-03-17" collectorName="Khao Soi Dao N. P." country="Thailand" elevation="166" latitude="13.096944" location="Chanthaburi Prov." longLatPrecision="21" longitude="102.1725" pageId="1" pageNumber="535" specimenCount="1" stateProvince="Chanthaburi" typeStatus="paratype">
<typeStatus id="65380D8AFFA2345FBBD03D8CFE9C60AB" box="[199,312,954,980]" pageId="1" pageNumber="535" type="paratype">Paratypes</typeStatus>
: 2♁.
<collectingCountry id="C294F3B8FFA2345FBA723D8DFDA060A9" box="[357,516,955,982]" name="Thailand" pageId="1" pageNumber="535">THAILAND</collectingCountry>
;
<collectingRegion id="78477DCAFFA2345FB9043D8CFD4160AB" box="[531,741,954,980]" country="Thailand" name="Chanthaburi" pageId="1" pageNumber="535">Chanthaburi Prov.</collectingRegion>
,
<collectorName id="1776D6FEFFA2345FB9E43D8CFC7060AB" box="[755,980,954,980]" pageId="1" pageNumber="535">Khao Soi Dao N.P.</collectorName>
,
<geoCoordinate id="DFB7D5EFFFA2345FB8F43D8DFBC860A9" box="[995,1132,955,982]" degrees="13" direction="north" minutes="05" orientation="latitude" pageId="1" pageNumber="535" precision="15" seconds="49" value="13.096944">13o05ʹ49ʺN</geoCoordinate>
,
<geoCoordinate id="DFB7D5EFFFA2345FBF6C3D8DFAAB60A9" box="[1147,1295,955,982]" degrees="102" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="535" precision="15" seconds="21" value="102.1725">102o10ʹ21ʺE</geoCoordinate>
, alt.
<quantity id="7D7B1ECDFFA2345FBE5C3D8CFA3360AB" box="[1355,1431,954,980]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="1" pageNumber="535" unit="m" value="166.0">
<elevation id="31AE541BFFA2345FBE5C3D8CFA3360AB" box="[1355,1431,954,980]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="1" pageNumber="535" unit="m" value="166.0">166 m</elevation>
</quantity>
,
<date id="CE3D95E8FFA2345FBB803DE8FE906087" box="[151,308,990,1016]" pageId="1" pageNumber="535" value="2008-03-17">
<collectingDate id="DE796C00FFA2345FBB803DE8FE906087" box="[151,308,990,1016]" pageId="1" pageNumber="535" value="2008-03-17">17 Mar. 2008</collectingDate>
</date>
, np2752 (genitalia slide NP1532)
</materialsCitation>
;
<materialsCitation id="0AEBB975FFA2345FB9D33DE8FECA673F" ID-GBIF-Occurrence="4003333301" collectingDate="2018-03-16" collectionCode="KKIC" collectorName="Phu Khieo W. S. &amp; Pinkaew" country="Thailand" elevation="770" latitude="16.506666" location="Chaiyaphum Prov." longLatPrecision="21" longitude="101.689445" pageId="1" pageNumber="535" specimenCount="1" stateProvince="Chaiyaphum" typeStatus="paratype">
<collectingRegion id="78477DCAFFA2345FB9D33DE8FC3E6087" box="[708,922,990,1016]" country="Thailand" name="Chaiyaphum" pageId="1" pageNumber="535">Chaiyaphum Prov.</collectingRegion>
,
<collectorName id="1776D6FEFFA2345FB8B13DE8FBC46087" box="[934,1120,990,1016]" pageId="1" pageNumber="535">Phu Khieo W.S.</collectorName>
,
<geoCoordinate id="DFB7D5EFFFA2345FBF7A3DE9FB536085" box="[1133,1271,991,1018]" degrees="16" direction="north" minutes="30" orientation="latitude" pageId="1" pageNumber="535" precision="15" seconds="24" value="16.506666">16o30ʹ24ʺN</geoCoordinate>
,
<geoCoordinate id="DFB7D5EFFFA2345FBE133DE9FA3C6085" box="[1284,1432,991,1018]" degrees="101" direction="east" minutes="41" orientation="longitude" pageId="1" pageNumber="535" precision="15" seconds="22" value="101.689445">101o41ʹ22ʺE</geoCoordinate>
, alt.
<quantity id="7D7B1ECDFFA2345FBBD43A34FEA96763" box="[195,269,1026,1052]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="535" unit="m" value="770.0">
<elevation id="31AE541BFFA2345FBBD43A34FEA96763" box="[195,269,1026,1052]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="535" unit="m" value="770.0">770 m</elevation>
</quantity>
,
<date id="CE3D95E8FFA2345FBA0D3A34FE126763" box="[282,438,1026,1052]" pageId="1" pageNumber="535" value="2018-03-16">
<collectingDate id="DE796C00FFA2345FBA0D3A34FE126763" box="[282,438,1026,1052]" pageId="1" pageNumber="535" value="2018-03-16">16 Mar. 2018</collectingDate>
</date>
, np11987 (genitalia slide NP3976). All specimens were collected by
<collectorName id="1776D6FEFFA2345FBFDE3A34FA8D6763" box="[1225,1321,1026,1052]" pageId="1" pageNumber="535">Pinkaew</collectorName>
<emphasis id="88F76F3AFFA2345FBE253A35FAC16763" box="[1330,1381,1026,1052]" italics="true" pageId="1" pageNumber="535">et al</emphasis>
. and deposited in
<collectionCode id="DC922BEDFFA2345FBA313A10FECA673F" box="[294,366,1062,1088]" pageId="1" pageNumber="535">KKIC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="EEFCE3A0FFA2345FBB8038CAFA98646B" ID-DOI="http://doi.org/10.5281/zenodo.7477368" ID-Zenodo-Dep="7477368" box="[151,1340,1788,1812]" httpUri="https://zenodo.org/record/7477368/files/figure.png" pageId="1" pageNumber="535" startId="1.[151,250,1788,1812]" targetBox="[151,1435,1112,1764]" targetPageId="1">
<paragraph id="BA3CB328FFA2345FBB8038CAFA98646B" blockId="1.[151,1340,1788,1812]" box="[151,1340,1788,1812]" pageId="1" pageNumber="535">
<emphasis id="88F76F3AFFA2345FBB8038CAFEB7646B" bold="true" box="[151,275,1788,1812]" pageId="1" pageNumber="535">FIGURE 1.</emphasis>
Living male specimen of
<taxonomicName id="7D83C8ABFFA2345FB90B38CAFD4C646B" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[540,744,1788,1812]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA2345FB90B38CAFD4C646B" box="[540,744,1788,1812]" italics="true" pageId="1" pageNumber="535">Thymioptila clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA2345FB9F938CAFC9A646B" bold="true" box="[750,830,1788,1812]" pageId="1" pageNumber="535">
<taxonomicNameLabel id="93C4D241FFA2345FB9F938CAFC9A646B" box="[750,830,1788,1812]" pageId="1" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype, np 11987) with natural resting posture.
</paragraph>
</caption>
<subSubSection id="F299E0A3FFA2345CBBD03974FAD86166" lastPageId="2" lastPageNumber="536" pageId="1" pageNumber="535" type="diagnosis">
<paragraph id="BA3CB328FFA2345CBBD03974FAD86166" blockId="1.[151,1437,1858,2029]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="536" pageId="1" pageNumber="535">
<emphasis id="88F76F3AFFA2345FBBD03974FEE66423" bold="true" box="[199,322,1858,1884]" pageId="1" pageNumber="535">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="7D83C8ABFFA2345FB9B93974FBB36423" authority="(Meyrick)" baseAuthorityName="Meyrick" box="[686,1047,1858,1884]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA2345FB9B93974FC3C6423" box="[686,920,1858,1884]" italics="true" pageId="1" pageNumber="535">Thymioptila oedalea</emphasis>
(Meyrick)
</taxonomicName>
. Forewing of both species have a tuft of the longitudinal raised scales at posterior end of closed cell. These scales are quite distinct and more raised up in resting posture of living specimens (
<figureCitation id="22B8AFADFFA2345FB99639BCFD6064DA" box="[641,708,1930,1957]" captionStart="FIGURE 1" captionStartId="1.[151,250,1788,1812]" captionTargetBox="[151,1435,1112,1764]" captionTargetId="figure-292@1.[151,1435,1112,1764]" captionTargetPageId="1" captionText="FIGURE 1. Living male specimen of Thymioptila clavata sp. nov. (paratype, np 11987) with natural resting posture." figureDoi="http://doi.org/10.5281/zenodo.7477368" httpUri="https://zenodo.org/record/7477368/files/figure.png" pageId="1" pageNumber="535">Fig. 1</figureCitation>
). Labial palpus of both species has similar pattern (
<figureCitation id="22B8AFADFFA2345FBE3639BCFA3464DA" box="[1313,1424,1930,1957]" captionStart="FIGURES 57" captionStartId="4.[151,264,1993,2017]" captionTargetBox="[164,1423,709,1970]" captionTargetId="figure-188@4.[164,1423,709,1970]" captionTargetPageId="4" captionText="FIGURES 57. Head of Thymioptila spp. 5. T. clavata sp. nov. (holotype). 6. T. oedalea (male). 7. T. oedalea (female)." figureDoi="http://doi.org/10.5281/zenodo.7477374" httpUri="https://zenodo.org/record/7477374/files/figure.png" pageId="1" pageNumber="535">Figs. 57</figureCitation>
). Marking at the middle of costa is slightly different by a moderately large subtriangular mark in
<taxonomicName id="7D83C8ABFFA2345FBFCA3999FAF764B7" authority="(Meyrick)" box="[1245,1363,1966,1992]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA2345FBFCA3999FAF764B7" box="[1245,1363,1966,1992]" italics="true" pageId="1" pageNumber="535">T. oedalea</emphasis>
</taxonomicName>
(
<figureCitation id="22B8AFADFFA2345FBE723998FF676492" captionStart="FIGURES 24" captionStartId="3.[151,264,1915,1939]" captionTargetBox="[354,1236,207,1850]" captionTargetId="figure-26@3.[321,1266,181,1891]" captionTargetPageId="3" captionText="FIGURES 24. Wing pattern of Thymioptila spp. 2. T. clavata sp. nov. (holotype). 3. T. oedalea (male). 4. T. oedalea (female). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7477370" httpUri="https://zenodo.org/record/7477370/files/figure.png" pageId="1" pageNumber="535">Figs. 34</figureCitation>
) but a rather small spot in
<taxonomicName id="7D83C8ABFFA2345FBAE539E4FD776493" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[498,723,2002,2028]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA2345FBAE539E4FD776493" box="[498,723,2002,2028]" italics="true" pageId="1" pageNumber="535">Thymioptila clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA2345FB9CC39E5FC916492" bold="true" box="[731,821,2003,2029]" pageId="1" pageNumber="535">
<taxonomicNameLabel id="93C4D241FFA2345FB9CC39E5FC916492" box="[731,821,2003,2029]" pageId="1" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="22B8AFADFFA2345FB85139E4FC2F6492" box="[838,907,2002,2029]" captionStart="FIGURES 24" captionStartId="3.[151,264,1915,1939]" captionTargetBox="[354,1236,207,1850]" captionTargetId="figure-26@3.[321,1266,181,1891]" captionTargetPageId="3" captionText="FIGURES 24. Wing pattern of Thymioptila spp. 2. T. clavata sp. nov. (holotype). 3. T. oedalea (male). 4. T. oedalea (female). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7477370" httpUri="https://zenodo.org/record/7477370/files/figure.png" pageId="1" pageNumber="535">Fig. 2</figureCitation>
). Hind wing of
<taxonomicName id="7D83C8ABFFA2345FBF2939E5FB086493" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[1086,1196,2002,2028]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="535" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA2345FBF2939E5FB086493" box="[1086,1196,2002,2028]" italics="true" pageId="1" pageNumber="535">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA2345FBFA339E5FAAA6492" bold="true" box="[1204,1294,2003,2029]" pageId="1" pageNumber="535">
<taxonomicNameLabel id="93C4D241FFA2345FBFA339E5FAAA6492" box="[1204,1294,2003,2029]" pageId="1" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a tuft of long hair pencils on anal marginal groove on the upper side (
<figureCitation id="22B8AFADFFA1345CB8433EA1FC3C63CE" box="[852,920,151,177]" captionStart="FIGURES 910" captionStartId="5.[151,264,1951,1975]" captionTargetBox="[151,1435,929,1927]" captionTargetId="figure-44@5.[151,1435,929,1927]" captionTargetPageId="5" captionText="FIGURES 910. Modified scales on anal area of hindwing of Thymioptila clavata sp. nov. (holotype). 9. A tuft of hair pencils tuft in anal fold and dense of long scales along 3A (upper side). 10. A row of long hair pencils along 3A (underside)." figureDoi="http://doi.org/10.5281/zenodo.7477380" httpUri="https://zenodo.org/record/7477380/files/figure.png" pageId="2" pageNumber="536">Fig. 9</figureCitation>
) and a row of long setaform scales along 3A on the underside (
<figureCitation id="22B8AFADFFA1345CBA753E8DFE1063AA" box="[354,436,187,213]" captionStart="FIGURES 910" captionStartId="5.[151,264,1951,1975]" captionTargetBox="[151,1435,929,1927]" captionTargetId="figure-44@5.[151,1435,929,1927]" captionTargetPageId="5" captionText="FIGURES 910. Modified scales on anal area of hindwing of Thymioptila clavata sp. nov. (holotype). 9. A tuft of hair pencils tuft in anal fold and dense of long scales along 3A (upper side). 10. A row of long hair pencils along 3A (underside)." figureDoi="http://doi.org/10.5281/zenodo.7477380" httpUri="https://zenodo.org/record/7477380/files/figure.png" pageId="2" pageNumber="536">Fig. 10</figureCitation>
), but both characters are absent in
<emphasis id="88F76F3AFFA1345CB82A3E8DFC1D63AA" box="[829,953,187,213]" italics="true" pageId="2" pageNumber="536">
<taxonomicName id="7D83C8ABFFA1345CB82A3E8DFC1163AA" authority="(Meyrick)" box="[829,949,187,213]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">T. oedalea</taxonomicName>
.
</emphasis>
Hind tibia in male of
<taxonomicName id="7D83C8ABFFA1345CBFA53E8DFA8463AA" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[1202,1312,187,213]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA1345CBFA53E8DFA8463AA" box="[1202,1312,187,213]" italics="true" pageId="2" pageNumber="536">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA1345CBE3F3E8DFA2663AA" bold="true" box="[1320,1410,187,213]" pageId="2" pageNumber="536">
<taxonomicNameLabel id="93C4D241FFA1345CBE3F3E8DFA2663AA" box="[1320,1410,187,213]" pageId="2" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is enlarged by covered bushy scales (
<figureCitation id="22B8AFADFFA1345CB9093EE9FDC86386" box="[542,620,223,249]" captionStart="FIGURES 1112" captionStartId="6.[151,264,1157,1181]" captionTargetBox="[151,1435,181,1133]" captionTargetId="figure-15@6.[151,1435,181,1133]" captionTargetPageId="6" captionText="FIGURES 1112. Sex scales on hind leg of Thymioptila spp. 11. T. clavata sp. nov. (holotype). 12. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477384" httpUri="https://zenodo.org/record/7477384/files/figure.png" pageId="2" pageNumber="536">Fig. 11</figureCitation>
) but hind tibia of
<taxonomicName id="7D83C8ABFFA1345CB8203EE9FC0F6386" authority="(Meyrick)" box="[823,939,223,249]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA1345CB8203EE9FC0F6386" box="[823,939,223,249]" italics="true" pageId="2" pageNumber="536">T. oedalea</emphasis>
</taxonomicName>
is simple and covered with long and narrow setaform scales (
<figureCitation id="22B8AFADFFA1345CBA4F3F35FE096262" box="[344,429,259,285]" captionStart="FIGURES 1112" captionStartId="6.[151,264,1157,1181]" captionTargetBox="[151,1435,181,1133]" captionTargetId="figure-15@6.[151,1435,181,1133]" captionTargetPageId="6" captionText="FIGURES 1112. Sex scales on hind leg of Thymioptila spp. 11. T. clavata sp. nov. (holotype). 12. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477384" httpUri="https://zenodo.org/record/7477384/files/figure.png" pageId="2" pageNumber="536">Fig. 12</figureCitation>
). Pregenital abdomen of both species can be separated by shape of sclerotized plate on tergum VIII (broad subtriangular in
<taxonomicName id="7D83C8ABFFA1345CB9273F11FD3A623E" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[560,670,295,321]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA1345CB9273F11FD3A623E" box="[560,670,295,321]" italics="true" pageId="2" pageNumber="536">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA1345CB9B13F11FCA5623E" bold="true" box="[678,769,295,321]" pageId="2" pageNumber="536">
<taxonomicNameLabel id="93C4D241FFA1345CB9B13F11FCA5623E" box="[678,769,295,321]" pageId="2" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but elongate subtriangular in
<taxonomicName id="7D83C8ABFFA1345CBF433F11FB6D623E" authority="(Meyrick)" box="[1108,1225,295,321]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA1345CBF433F11FB6D623E" box="[1108,1225,295,321]" italics="true" pageId="2" pageNumber="536">T. oedalea</emphasis>
</taxonomicName>
) and sternum VIII (narrow band connected with slightly sinuate band in
<taxonomicName id="7D83C8ABFFA1345CB9E13F7DFCC0621A" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[758,868,331,357]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA1345CB9E13F7DFCC0621A" box="[758,868,331,357]" italics="true" pageId="2" pageNumber="536">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA1345CB87B3F7DFC62621A" bold="true" box="[876,966,331,357]" pageId="2" pageNumber="536">
<taxonomicNameLabel id="93C4D241FFA1345CB87B3F7DFC62621A" box="[876,966,331,357]" pageId="2" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but narrow band connected with strongly curve band in
<taxonomicName id="7D83C8ABFFA1345CBA213F59FE0E62F6" authority="(Meyrick)" box="[310,426,367,393]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA1345CBA213F59FE0E62F6" box="[310,426,367,393]" italics="true" pageId="2" pageNumber="536">T. oedalea</emphasis>
</taxonomicName>
), especially two large patches of dense scale sockets between tergum VII and sternum VII on posterior 1/
<quantity id="7D7B1ECDFFA1345CBA2D3FA5FECC62D2" box="[314,360,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="536" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="7D83C8ABFFA1345CBA783FA5FE7A62D2" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[367,478,403,429]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA1345CBA783FA5FE7A62D2" box="[367,478,403,429]" italics="true" pageId="2" pageNumber="536">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA1345CBAF13FA5FDE462D2" bold="true" box="[486,576,403,429]" pageId="2" pageNumber="536">
<taxonomicNameLabel id="93C4D241FFA1345CBAF13FA5FDE462D2" box="[486,576,403,429]" pageId="2" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but absent in
<taxonomicName id="7D83C8ABFFA1345CB9F73FA5FCF162D2" authority="(Meyrick)" box="[736,853,403,429]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA1345CB9F73FA5FCF162D2" box="[736,853,403,429]" italics="true" pageId="2" pageNumber="536">T. oedalea</emphasis>
</taxonomicName>
(
<figureCitation id="22B8AFADFFA1345CB8713FA5FC4862D2" box="[870,1004,403,429]" captionStart="FIGURES 1314" captionStartId="6.[151,264,1951,1975]" captionTargetBox="[167,1429,1206,1909]" captionTargetId="figure-37@6.[151,1435,1206,1928]" captionTargetPageId="6" captionText="FIGURES 1314. Sclerotized plates on 8th tergum (left) and sternum (right) of Thymioptila spp. 13. T. clavata sp. nov. (holotype). 14. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477388" httpUri="https://zenodo.org/record/7477388/files/figure.png" pageId="2" pageNumber="536">Figs. 1314</figureCitation>
). Male genitalia of
<taxonomicName id="7D83C8ABFFA1345CBFDB3FA5FA9E62D2" authority="Abstract, 2022" authorityName="Abstract" authorityYear="2022" box="[1228,1338,403,429]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata" status="sp. nov.">
<emphasis id="88F76F3AFFA1345CBFDB3FA5FA9E62D2" box="[1228,1338,403,429]" italics="true" pageId="2" pageNumber="536">T. clavata</emphasis>
</taxonomicName>
<emphasis id="88F76F3AFFA1345CBE553FA5FA3862D2" bold="true" box="[1346,1436,403,429]" pageId="2" pageNumber="536">
<taxonomicNameLabel id="93C4D241FFA1345CBE553FA5FA3862D2" box="[1346,1436,403,429]" pageId="2" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished by subtriangular uncus, subtriangular cucullus with a large curved spine on large subtriangular lobe at apex and a large claviform cucullus process (
<figureCitation id="22B8AFADFFA1345CB9F43FEDFC97628A" box="[739,819,474,501]" captionStart="FIGURES 1516" captionStartId="7.[151,264,2006,2030]" captionTargetBox="[179,1406,195,1969]" captionTargetId="figure-26@7.[151,1435,181,1982]" captionTargetPageId="7" captionText="FIGURES 1516. Male genitalia of Thymioptila spp. 15. T. clavata sp. nov. (holotype). 16. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477392" httpUri="https://zenodo.org/record/7477392/files/figure.png" pageId="2" pageNumber="536">Fig. 15</figureCitation>
) whereas uncus is sub-semicircular, cucullus is narrow and curve with a large curved spine at apex and quite narrow claviform cucullus process in
<taxonomicName id="7D83C8ABFFA1345CBF803FC9FAA86166" authority="(Meyrick)" box="[1175,1292,511,537]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="oedalea">
<emphasis id="88F76F3AFFA1345CBF803FC9FAA86166" box="[1175,1292,511,537]" italics="true" pageId="2" pageNumber="536">T. oedalea</emphasis>
</taxonomicName>
(
<figureCitation id="22B8AFADFFA1345CBE0B3FC9FACA6166" box="[1308,1390,511,537]" captionStart="FIGURES 1516" captionStartId="7.[151,264,2006,2030]" captionTargetBox="[179,1406,195,1969]" captionTargetId="figure-26@7.[151,1435,181,1982]" captionTargetPageId="7" captionText="FIGURES 1516. Male genitalia of Thymioptila spp. 15. T. clavata sp. nov. (holotype). 16. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477392" httpUri="https://zenodo.org/record/7477392/files/figure.png" pageId="2" pageNumber="536">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F299E0A3FFA1345CBBD03C14FE0664EA" pageId="2" pageNumber="536" type="description">
<paragraph id="BA3CB328FFA1345CBBD03C14FE6961B2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="536">
<emphasis id="88F76F3AFFA1345CBBD03C14FEFD6143" bold="true" box="[199,345,546,572]" pageId="2" pageNumber="536">Description.</emphasis>
Head (
<figureCitation id="22B8AFADFFA1345CBAA33C15FE5C6142" box="[436,504,546,573]" captionStart="FIGURES 57" captionStartId="4.[151,264,1993,2017]" captionTargetBox="[164,1423,709,1970]" captionTargetId="figure-188@4.[164,1423,709,1970]" captionTargetPageId="4" captionText="FIGURES 57. Head of Thymioptila spp. 5. T. clavata sp. nov. (holotype). 6. T. oedalea (male). 7. T. oedalea (female)." figureDoi="http://doi.org/10.5281/zenodo.7477374" httpUri="https://zenodo.org/record/7477374/files/figure.png" pageId="2" pageNumber="536">Fig. 5</figureCitation>
). Lower frons yellowish white; upper frons light brown; vertex light brown mixed with brown near chaetosema, posterior margin of compound eyes edged with brown alternated with white scales; antenna light brown; labial palpus with first segment white, second segment enlarged, parallel margins except base, basal half white, apical half light brown with a large transverse brown patch apically, from dorsal to ventral margin, third segment short, brown.
</paragraph>
<paragraph id="BA3CB328FFA1345CBBD03CE1FEC8662A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="536">
Thorax: pronotal collar light brown mixed with some purple-brown scales, tegulae light brown ground with large patch inner half, brown mixed with dark brown; mesonotum light brown mixed with greyish brown, without posterior crest; Hind legs (
<figureCitation id="22B8AFADFFA1345CBADF3D29FDBD6046" box="[456,537,799,825]" captionStart="FIGURES 1112" captionStartId="6.[151,264,1157,1181]" captionTargetBox="[151,1435,181,1133]" captionTargetId="figure-15@6.[151,1435,181,1133]" captionTargetPageId="6" captionText="FIGURES 1112. Sex scales on hind leg of Thymioptila spp. 11. T. clavata sp. nov. (holotype). 12. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477384" httpUri="https://zenodo.org/record/7477384/files/figure.png" pageId="2" pageNumber="536">Fig. 11</figureCitation>
) strongly modified, inner side of femur with patch of dense, long white scales along ventral margin, tibia enlarged by dense scales, slightly narrowed at posterior end, mostly white mixed with dark brown beneath. Forewing subrectangular, wingspan 19.520.0 mm in males (
<figureCitation id="22B8AFADFFA1345CBF5E3D51FB2B60FE" box="[1097,1167,871,897]" captionStart="FIGURES 24" captionStartId="3.[151,264,1915,1939]" captionTargetBox="[354,1236,207,1850]" captionTargetId="figure-26@3.[321,1266,181,1891]" captionTargetPageId="3" captionText="FIGURES 24. Wing pattern of Thymioptila spp. 2. T. clavata sp. nov. (holotype). 3. T. oedalea (male). 4. T. oedalea (female). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7477370" httpUri="https://zenodo.org/record/7477370/files/figure.png" pageId="2" pageNumber="536">Fig. 2</figureCitation>
) (n = 3); costal margin rather straight, gently curved at apical 1/4, termen truncate, rather straight from wing apex to M
<subScript id="2607B16DFFA1345CBFF03DABFB4B60D4" attach="left" box="[1255,1263,925,939]" fontSize="6" pageId="2" pageNumber="536">3</subScript>
then curved to tornus; costal strigulae well developed, creamy white separated by small dark brown streaks, 1
<superScript id="4DF61E60FFA1345CBFFC3D99FB5260C2" attach="none" box="[1259,1270,943,957]" fontSize="6" pageId="2" pageNumber="536">st</superScript>
2
<superScript id="4DF61E60FFA1345CBE053D98FA8660C3" attach="left" box="[1298,1314,942,956]" fontSize="6" pageId="2" pageNumber="536">nd</superScript>
indistinct, 3
<superScript id="4DF61E60FFA1345CBBB23DE4FF17609F" attach="none" box="[165,179,978,992]" fontSize="6" pageId="2" pageNumber="536">rd</superScript>
4
<superScript id="4DF61E60FFA1345CBBD83DE4FF78609F" attach="left" box="[207,220,978,992]" fontSize="6" pageId="2" pageNumber="536">th</superScript>
double pairs, 5
<superScript id="4DF61E60FFA1345CBA9E3DE4FE32609F" attach="none" box="[393,406,978,992]" fontSize="6" pageId="2" pageNumber="536">th</superScript>
9
<superScript id="4DF61E60FFA1345CBAA53DE4FE1B609F" attach="left" box="[434,447,978,992]" fontSize="6" pageId="2" pageNumber="536">th</superScript>
more distinct pair on apical half; ground color creamy white mixed with scattered spots and short, light brown longitudinal lines throughout, basal 1/5 with rather small indistinct patch, brown mixed with dark brown, middle of costa with a small oblique mark, brown mixed with dark brown extending from costa to R
<subScript id="2607B16DFFA1345CBBBD3A67FF166720" attach="left" box="[170,178,1105,1119]" fontSize="6" pageId="2" pageNumber="536">1</subScript>
, apical 1/3 with a large indistinct transverse patch, brown mixed with dark brown, extending from R
<subScript id="2607B16DFFA1345CBE3F3A67FA946720" attach="left" box="[1320,1328,1105,1119]" fontSize="6" pageId="2" pageNumber="536">4</subScript>
to tornus broken by a narrow, longitudinal raised scale patch at the posterior end of closed cell between base of R
<subScript id="2607B16DFFA1345CBE2D3A43FAE667FC" attach="left" box="[1338,1346,1141,1155]" fontSize="6" pageId="2" pageNumber="536">5</subScript>
and M
<subScript id="2607B16DFFA1345CBE833A43FA3867FC" attach="left" box="[1428,1436,1141,1155]" fontSize="6" pageId="2" pageNumber="536">2</subScript>
(
<figureCitation id="22B8AFADFFA1345CBB883AB1FF4667DE" box="[159,226,1159,1185]" captionStart="FIGURE 8" captionStartId="5.[151,250,881,905]" captionTargetBox="[151,1435,181,857]" captionTargetId="figure-26@5.[151,1435,181,857]" captionTargetPageId="5" captionText="FIGURE 8. A tuft of raised scales on forewing of Thymioptila clavata sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7477376" httpUri="https://zenodo.org/record/7477376/files/figure.png" pageId="2" pageNumber="536">Fig. 8</figureCitation>
), more raised up in living specimens (
<figureCitation id="22B8AFADFFA1345CB9913AB1FD6267DE" box="[646,710,1159,1185]" captionStart="FIGURE 1" captionStartId="1.[151,250,1788,1812]" captionTargetBox="[151,1435,1112,1764]" captionTargetId="figure-292@1.[151,1435,1112,1764]" captionTargetPageId="1" captionText="FIGURE 1. Living male specimen of Thymioptila clavata sp. nov. (paratype, np 11987) with natural resting posture." figureDoi="http://doi.org/10.5281/zenodo.7477368" httpUri="https://zenodo.org/record/7477368/files/figure.png" pageId="2" pageNumber="536">Fig. 1</figureCitation>
); fringe scales light brown; underside brown with narrow creamy strip along costa apical 1/2. Hindwing (
<figureCitation id="22B8AFADFFA1345CB9593A9DFD6067BA" box="[590,708,1195,1221]" captionStart-0="FIGURE 8" captionStart-1="FIGURES 910" captionStartId-0="5.[151,250,881,905]" captionStartId-1="5.[151,264,1951,1975]" captionTargetBox-0="[151,1435,181,857]" captionTargetBox-1="[151,1435,929,1927]" captionTargetId-0="figure-26@5.[151,1435,181,857]" captionTargetId-1="figure-44@5.[151,1435,929,1927]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 8. A tuft of raised scales on forewing of Thymioptila clavata sp. nov. (holotype)." captionText-1="FIGURES 910. Modified scales on anal area of hindwing of Thymioptila clavata sp. nov. (holotype). 9. A tuft of hair pencils tuft in anal fold and dense of long scales along 3A (upper side). 10. A row of long hair pencils along 3A (underside)." figureDoi-0="http://doi.org/10.5281/zenodo.7477376" figureDoi-1="http://doi.org/10.5281/zenodo.7477380" httpUri-0="https://zenodo.org/record/7477376/files/figure.png" httpUri-1="https://zenodo.org/record/7477380/files/figure.png" pageId="2" pageNumber="536">Figs. 810</figureCitation>
) brown slight paler to base, anal area with long hairy scales, light brown, anal margin sclerotized, raised upward forming longitudinal groove, covered with white scales, bearing a tuft of long hair pencils, yellowish white; underside brown, slightly paler to base, except apical 1/2 of costa with a narrow creamy white strip and brownish yellow anal area with a distinct row of light brown long setaform scales along 3A medially.
</paragraph>
<paragraph id="BA3CB328FFA1345CBBD03B69FC27640E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="536">
Abdomen. Pregenital abdomen (
<figureCitation id="22B8AFADFFA1345CB9213B69FD236606" box="[566,647,1375,1401]" captionStart="FIGURES 1314" captionStartId="6.[151,264,1951,1975]" captionTargetBox="[167,1429,1206,1909]" captionTargetId="figure-37@6.[151,1435,1206,1928]" captionTargetPageId="6" captionText="FIGURES 1314. Sclerotized plates on 8th tergum (left) and sternum (right) of Thymioptila spp. 13. T. clavata sp. nov. (holotype). 14. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477388" httpUri="https://zenodo.org/record/7477388/files/figure.png" pageId="2" pageNumber="536">Fig. 13</figureCitation>
) with a narrowed sclerotized T-shaped band on sternum VIII; tergum VIII with a moderately sclerotized subtriangular plate, slightly tapered posteriorly to rounded apex, with two patch of dense scales socket on membrane between tergum VII and sternum VII on posterior 1/2. Male genitalia (
<figureCitation id="22B8AFADFFA1345CBE673B91FF10669A" captionStart="FIGURES 1516" captionStartId="7.[151,264,2006,2030]" captionTargetBox="[179,1406,195,1969]" captionTargetId="figure-26@7.[151,1435,181,1982]" captionTargetPageId="7" captionText="FIGURES 1516. Male genitalia of Thymioptila spp. 15. T. clavata sp. nov. (holotype). 16. T. oedalea." figureDoi="http://doi.org/10.5281/zenodo.7477392" httpUri="https://zenodo.org/record/7477392/files/figure.png" pageId="2" pageNumber="536">Fig. 15</figureCitation>
) with tegumen subtriangular, moderately wide, apicolateral 1/2 with dense scale sockets, basal 1/2 with sparsely scale sockets; uncus moderately large, subtriangular, rounded apex, with long, sparsely setose, posterior surface with two separated patch of dense scale sockets; socii narrow, membranous, densely setose; gnathos arising from near midlength of tegumen, forming an upcurved sclerotized narrow band; vinculum rather narrow; juxta subtriangular; caulis short; anellus a high hood-like surrounding basal half of phallus; phallus moderately long and wide, slightly sclerotized, margins rather parallel; tuba analis with wrinkled surface; cornuti absent; valva rather long and slender; sacculus with widened basal opening, basal half sparsely setose, apical half with moderately dense setae, costal margin with moderately dense setae at basal 1/2; neck distinct with deep incised ventrally; cucullus short forming moderately large subtriangular lobe, pointing ventrally with a large, pointed, curved spine at apex, covered with moderately dense scale sockets, with sparse long setae ventrally, dorsoapical corner with a distinct large and long clavate process, rounded apex, slightly curved upward and protruding outward, strongly widened apical 1/2, basal 1/2 with moderately dense scale sockets, apical 1/2 densely setose.
</paragraph>
<paragraph id="BA3CB328FFA1345CBBD0394DFE0664EA" blockId="2.[151,1437,151,2013]" box="[199,418,1915,1941]" pageId="2" pageNumber="536">
<emphasis id="88F76F3AFFA1345CBBD0394DFE8164EA" bold="true" box="[199,293,1915,1941]" pageId="2" pageNumber="536">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F299E0A3FFA1345CBBD039A8FBBD64C6" box="[199,1049,1950,1977]" pageId="2" pageNumber="536" type="distribution">
<paragraph id="BA3CB328FFA1345CBBD039A8FBBD64C6" blockId="2.[151,1437,151,2013]" box="[199,1049,1950,1977]" pageId="2" pageNumber="536">
<emphasis id="88F76F3AFFA1345CBBD039A8FEC564C7" bold="true" box="[199,353,1950,1976]" pageId="2" pageNumber="536">Distribution.</emphasis>
<collectingCountry id="C294F3B8FFA1345CBA7039A9FE6F64C6" box="[359,459,1951,1977]" name="Thailand" pageId="2" pageNumber="536">Thailand</collectingCountry>
(
<collectingRegion id="78477DCAFFA1345CBACD39A9FD9864C6" box="[474,572,1951,1977]" country="Thailand" name="Sa Kaeo" pageId="2" pageNumber="536">Sa Kaeo</collectingRegion>
,
<collectingRegion id="78477DCAFFA1345CB95039A9FD7064C6" box="[583,724,1951,1977]" country="Thailand" name="Chanthaburi" pageId="2" pageNumber="536">Chanthaburi</collectingRegion>
, and
<collectingRegion id="78477DCAFFA1345CB81839A9FBAA64C6" box="[783,1038,1951,1977]" country="Thailand" name="Chaiyaphum" pageId="2" pageNumber="536">Chaiyaphum Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="F299E0A3FFA1345CBBD039F5FB4964A2" box="[199,1261,1987,2013]" pageId="2" pageNumber="536" type="etymology">
<paragraph id="BA3CB328FFA1345CBBD039F5FB4964A2" blockId="2.[151,1437,151,2013]" box="[199,1261,1987,2013]" pageId="2" pageNumber="536">
<emphasis id="88F76F3AFFA1345CBBD039F5FEEA64A2" bold="true" box="[199,334,1987,2013]" pageId="2" pageNumber="536">Etymology.</emphasis>
The specific epithet (
<taxonomicName id="7D83C8ABFFA1345CB95539F5FD3064A2" authorityName="Abstract" authorityYear="2022" box="[578,660,1987,2013]" class="Insecta" family="Tortricidae" genus="Thymioptila" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="536" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="88F76F3AFFA1345CB95539F5FD3064A2" box="[578,660,1987,2013]" italics="true" pageId="2" pageNumber="536">clavata</emphasis>
</taxonomicName>
= clavate) refers to the shape of the cucullus process.
</paragraph>
</subSubSection>
</treatment>
</document>