473 lines
56 KiB
XML
473 lines
56 KiB
XML
<document id="F9DE2D6D779631D9678343B46589536C" ID-DOI="10.4202/app.00554.2018" ID-ISSN="1732-2421" ID-Zenodo-Dep="10980900" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1713284217429" checkinUser="felipe" docAuthor="Hryniewicz, Krzysztof, Amano, Kazutaka, Bitner, Maria Aleksandra, Hagström, Jonas, Kiel, Steffen, Klompmaker, Adiël A., Mörs, Thomas, Robins, Cristina M. & Kaim, Andrzej" docDate="2019" docId="320C87F44658FFC9FCD3FDDDFB0FD9E6" docLanguage="en" docName="ActaPalaeontolPol.64.1.101-141.pdf" docOrigin="Acta Palaeontologica Polonica 64 (1)" docSource="http://dx.doi.org/10.4202/app.00554.2018" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.2:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="2" docTitle="Neoliothyrina nakremi Bitner 2019, sp. nov." docType="treatment" docVersion="3" lastPageNumber="105" masterDocId="CE35FF8C465AFFCDFF8DFFFEFFA8DF58" masterDocTitle="A late Paleocene fauna from shallow-water chemosynthesis-based ecosystems, Spitsbergen, Svalbard" masterLastPageNumber="141" masterPageNumber="101" pageNumber="103" updateTime="1713358443740" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B43D68AACF02E01370B821CFE951D53C">A late Paleocene fauna from shallow-water chemosynthesis-based ecosystems, Spitsbergen, Svalbard</mods:title>
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<mods:namePart id="BDBFEF4FEE73157E0959D0F9E03DCA98">Hryniewicz, Krzysztof</mods:namePart>
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<mods:namePart id="620BE488DA5C80DEC3659C82DCA16AC4">Amano, Kazutaka</mods:namePart>
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<mods:namePart id="6118C22CAC248D08F7DA910E23C9BB96">Bitner, Maria Aleksandra</mods:namePart>
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<mods:namePart id="C14CBD0457673F1638A323733982F96F">Hagström, Jonas</mods:namePart>
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<mods:namePart id="4765DC04A9AE012F249E4192E8ED049E">Kiel, Steffen</mods:namePart>
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<mods:namePart id="875434850B1F9A50649DD8341E4ECBCC">Klompmaker, Adiël A.</mods:namePart>
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<mods:namePart id="0E25CB81802FD31A516B4DE411E3336A">Mörs, Thomas</mods:namePart>
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<mods:namePart id="24DB9BCA6C514799C2C872C643B0FA13">Robins, Cristina M.</mods:namePart>
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<mods:namePart id="AC57414495AFE60C87706571C8551A00">Kaim, Andrzej</mods:namePart>
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<mods:title id="9239C923E9D39AB4061BECCD66E12768">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="C4A4CCBA007302628A8ABFBD9E28ABF8">2019</mods:date>
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<mods:number id="5EBE27208362A89B8781C6EA07A02BF2">2019-02-13</mods:number>
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<mods:number id="6BB9ED40123A316489E555F8589C2485">64</mods:number>
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<mods:identifier id="ED8AF6C532E0D287BF4471A9EFF2F1E4" type="ISSN">1732-2421</mods:identifier>
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<treatment id="320C87F44658FFC9FCD3FDDDFB0FD9E6" LSID="urn:lsid:zoobank.org:act:C889EAF7-1909-4819-9B0E-9709A1EBA6ED" httpUri="http://treatment.plazi.org/id/320C87F44658FFC9FCD3FDDDFB0FD9E6" lastPageId="4" lastPageNumber="105" pageId="2" pageNumber="103">
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<subSubSection id="F2BF65694658FFCFFCD3FDDDFAEADD18" box="[862,1346,547,576]" pageId="2" pageNumber="103" type="nomenclature">
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<paragraph id="BA1A36E24658FFCFFCD3FDDDFAEADD18" blockId="2.[862,1346,547,576]" box="[862,1346,547,576]" pageId="2" pageNumber="103">
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<heading id="E152818E4658FFCFFCD3FDDDFAEADD18" box="[862,1346,547,576]" fontSize="12" level="2" pageId="2" pageNumber="103" reason="2">
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<taxonomicName id="7DA54D614658FFCFFCD3FDDDFB7FDD18" authority="Bitner" authorityName="Bitner" authorityYear="2019" box="[862,1239,547,576]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="2" pageNumber="103" phylum="Brachiopoda" rank="species" species="nakremi" status="sp. nov.">
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<emphasis id="88D1EAF04658FFCFFCD3FDDDFB28DD18" box="[862,1152,547,576]" italics="true" pageId="2" pageNumber="103">Neoliothyrina nakremi</emphasis>
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Bitner
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</taxonomicName>
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<taxonomicNameLabel id="93E2578B4658FFCFFB52FDDAFAEADD18" box="[1247,1346,548,576]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="F2BF65694658FFC9FCD3FDB1FB0FD9E6" lastPageId="4" lastPageNumber="105" pageId="2" pageNumber="103" type="description">
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<paragraph id="BA1A36E24658FFCFFCD3FDB1FC15DD3C" blockId="2.[862,957,591,613]" box="[862,957,591,613]" pageId="2" pageNumber="103">
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<figureCitation id="229E2A674658FFCFFCD3FDB1FC09DD3C" box="[862,929,591,613]" captionStart="Fig" captionStartId="3.[98,131,1199,1221]" captionTargetBox="[99,1481,163,1175]" captionTargetId="graphics-361@3.[99,1480,163,1175]" captionTargetPageId="3" captionText="Fig. 2. Transverse serial sections of the sellithyrid brachiopod Neoliothyrina nakremi Bitner sp. nov. paratype (ZPAL V.48/9-4) from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. L>20.2 mm. Numbers indicate distance in mm from the tip of the ventral umbo." figureDoi="http://doi.org/10.5281/zenodo.10980904" httpUri="https://zenodo.org/record/10980904/files/figure.png" pageId="2" pageNumber="103">Figs. 2</figureCitation>
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,
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<figureCitation id="229E2A674658FFCFFC26FDB1FC11DD3C" box="[939,953,591,612]" captionStart="Fig" captionStartId="4.[151,184,1498,1520]" captionTargetBox="[152,1532,161,1473]" captionTargetId="figure-236@4.[152,1532,161,1473]" captionTargetPageId="4" captionText="Fig. 3. Sellithyrid brachiopod Neoliothyrina nakremi Bitner sp. nov. from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. A. Holotype, ZPAL V.48/9-1, decorticated shell in ventral (A1), dorsal (A2), and posterior (A4) views; left-lateral view of both valves (A3). B. Paratype, ZPAL V.48/9-2; decorticated shell in ventral (B1), dorsal (B2), and posterior (B4) views; left-lateral view of both valves (B3). C. Paratype, ZPAL V.48/9-3, decorticated shell in ventral (C1), dorsal (C2), and posterior (C4) views; left-lateral view of both valves (C3)." figureDoi="http://doi.org/10.5281/zenodo.10980906" httpUri="https://zenodo.org/record/10980906/files/figure.png" pageId="2" pageNumber="103">3</figureCitation>
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.
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</paragraph>
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<paragraph id="BA1A36E24658FFCFFCD3FD88FACFDDD3" blockId="2.[862,1383,630,680]" box="[862,1383,630,652]" pageId="2" pageNumber="103">
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||
?1927
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||
<taxonomicName id="7DA54D614658FFCFFC13FD88FBEDDDD4" box="[926,1093,630,652]" class="Rhynchonellata" family="Cancellothyrididae" genus="Terebratulina" kingdom="Animalia" order="Terebratulida" pageId="2" pageNumber="103" phylum="Brachiopoda" rank="species" species="undetermined">
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<emphasis id="88D1EAF04658FFCFFC13FD88FB88DDD3" box="[926,1056,630,651]" italics="true" pageId="2" pageNumber="103">Terebratulina</emphasis>
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sp.
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||
</taxonomicName>
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||
;
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||
<bibRefCitation id="DE344B134658FFCFFBC2FD88FB4BDDD3" author="Gripp, K." box="[1103,1251,630,652]" pageId="2" pageNumber="103" pagination="3 - 38" refId="ref28977" refString="Gripp, K. 1927. Beitrage zur Geologie von Spitzbergen. Abhandlungen des Naturwissenschaftlichen Vereins zu Hamburg 21: 3 - 38." type="journal article" year="1927">Gripp 1927: 30</bibRefCitation>
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||
, pl. 6: 13–14.
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||
</paragraph>
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||
<paragraph id="BA1A36E24658FFCFFCD3FD6CFACFDDF0" blockId="2.[862,1383,630,680]" box="[862,1383,658,680]" pageId="2" pageNumber="103">
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||
2016
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||
<taxonomicName id="7DA54D614658FFCFFC19FD6CFBA9DDFF" authorityName="Roy" authorityYear="1980" box="[916,1025,658,679]" class="Rhynchonellata" family="Terebratulidae" genus="Pliothyrina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="2" pageNumber="103" phylum="Brachiopoda" rank="genus">
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||
<emphasis id="88D1EAF04658FFCFFC19FD6CFBA9DDFF" box="[916,1025,658,679]" italics="true" pageId="2" pageNumber="103">Pliothyrina</emphasis>
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||
</taxonomicName>
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||
? sp.;
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||
<bibRefCitation id="DE344B134658FFCFFBB7FD6CFABEDDF0" author="Hryniewicz, K. & Bitner, M. A. & Durska, E. & Hagstrom, J. & Hjalmarsdottir H. R. & Jenkins, R. G. & Little, C. T. S. & Miyajima, Y. & Nakrem, H. A. & Kaim, A." box="[1082,1302,658,680]" pageId="2" pageNumber="103" pagination="41 - 56" refId="ref29314" refString="Hryniewicz, K., Bitner, M. A., Durska, E., Hagstrom, J., Hjalmarsdottir H. R., Jenkins, R. G., Little, C. T. S., Miyajima, Y., Nakrem, H. A., and Kaim, A. 2016. Paleocene methane seep and wood-fall marine environments from Spitsbergen, Svalbard. Palaeogeography, Palaeoclimatology, Palaeoecology 462: 41 - 56." type="journal article" year="2016">Hryniewicz et al. 2016</bibRefCitation>
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, table 2.
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</paragraph>
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<paragraph id="BA1A36E24658FFCFFCD3FD4AFBADDDBE" blockId="2.[862,1535,692,742]" pageId="2" pageNumber="103">
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<emphasis id="88D1EAF04658FFCFFCD3FD4AFC42DD91" box="[862,1002,692,713]" italics="true" pageId="2" pageNumber="103">ZooBank LSID</emphasis>
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:
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<uri id="CE343AE04658FFCFFC7BFD4AFBADDDBE" pageId="2" pageNumber="103">
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urn:lsid:zoobank.org:act:
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<uuid id="CE030C374658FFCFFB53FD4AFBADDDBE" pageId="2" pageNumber="103">C889EAF7-1909-4819-9B0E-9709A1EBA6ED</uuid>
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</uri>
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</paragraph>
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<paragraph id="BA1A36E24658FFCFFCD3FD0CFBE7DC18" blockId="2.[862,1535,754,832]" pageId="2" pageNumber="103">
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<emphasis id="88D1EAF04658FFCFFCD3FD0CFC6DDC5F" box="[862,965,754,775]" italics="true" pageId="2" pageNumber="103">Etymology</emphasis>
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||
: In honour of Norwegian palaeontologist Hans Arne Nakrem, in recognition of his studies on fossil chemosynthesis-based faunas from Spitsbergen.
|
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</paragraph>
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<paragraph id="BA1A36E24658FFCFFCD3FCB2FA44DCEF" blockId="2.[862,1535,844,951]" pageId="2" pageNumber="103">
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<emphasis id="88D1EAF04658FFCFFCD3FCB2FC4ADC39" box="[862,994,844,865]" italics="true" pageId="2" pageNumber="103">Type material</emphasis>
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:
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<materialsCitation id="0ACD3CBF4658FFCFFC7DFCB2FB99DC26" collectionCode="ZPAL" pageId="2" pageNumber="103" specimenCount="1" typeStatus="holotype">
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<typeStatus id="651E88404658FFCFFC7DFCB2FBE5DC3A" box="[1008,1101,844,866]" pageId="2" pageNumber="103" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="DCB4AE274658FFCFFBDAFCB2FB38DC39" box="[1111,1168,844,865]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
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V.48/9-1, decorticated specimen with broken beak (
|
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<figureCitation id="229E2A674658FFCFFC50FC96FB81DC26" box="[989,1065,872,894]" captionStart="Fig" captionStartId="4.[151,184,1498,1520]" captionTargetBox="[152,1532,161,1473]" captionTargetId="figure-236@4.[152,1532,161,1473]" captionTargetPageId="4" captionText="Fig. 3. Sellithyrid brachiopod Neoliothyrina nakremi Bitner sp. nov. from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. A. Holotype, ZPAL V.48/9-1, decorticated shell in ventral (A1), dorsal (A2), and posterior (A4) views; left-lateral view of both valves (A3). B. Paratype, ZPAL V.48/9-2; decorticated shell in ventral (B1), dorsal (B2), and posterior (B4) views; left-lateral view of both valves (B3). C. Paratype, ZPAL V.48/9-3, decorticated shell in ventral (C1), dorsal (C2), and posterior (C4) views; left-lateral view of both valves (C3)." figureDoi="http://doi.org/10.5281/zenodo.10980906" httpUri="https://zenodo.org/record/10980906/files/figure.png" pageId="2" pageNumber="103">Fig. 3A</figureCitation>
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)
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</materialsCitation>
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.
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<materialsCitation id="0ACD3CBF4658FFCFFBB5FC96FA40DCEF" collectionCode="ZPAL" pageId="2" pageNumber="103" specimenCount="1" typeStatus="paratype">
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<typeStatus id="651E88404658FFCFFBB5FC96FB31DC25" box="[1080,1177,872,893]" pageId="2" pageNumber="103" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="DCB4AE274658FFCFFB2DFC96FB71DC25" box="[1184,1241,872,893]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
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||
V.48/9-2–4, decorticated specimens with broken beak (
|
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<collectionCode id="DCB4AE274658FFCFFBECFC7AFB34DCC1" box="[1121,1180,900,921]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
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V.48/9-2) and anterior part (
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<collectionCode id="DCB4AE274658FFCFFA49FC7AFA57DCC1" box="[1476,1535,900,921]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
|
||
V.48/9-2–3); a set of acetate peels (
|
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<collectionCode id="DCB4AE274658FFCFFB22FC5FFB42DCEE" box="[1199,1258,929,950]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
|
||
V.48/9-4) (
|
||
<figureCitation id="229E2A674658FFCFFADAFC5EFA33DCEE" box="[1367,1435,928,950]" captionStart="Fig" captionStartId="3.[98,131,1199,1221]" captionTargetBox="[99,1481,163,1175]" captionTargetId="graphics-361@3.[99,1480,163,1175]" captionTargetPageId="3" captionText="Fig. 2. Transverse serial sections of the sellithyrid brachiopod Neoliothyrina nakremi Bitner sp. nov. paratype (ZPAL V.48/9-4) from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. L>20.2 mm. Numbers indicate distance in mm from the tip of the ventral umbo." figureDoi="http://doi.org/10.5281/zenodo.10980904" httpUri="https://zenodo.org/record/10980904/files/figure.png" pageId="2" pageNumber="103">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="229E2A674658FFCFFA28FC5FFA48DCEF" box="[1445,1504,929,951]" captionStart="Fig" captionStartId="4.[151,184,1498,1520]" captionTargetBox="[152,1532,161,1473]" captionTargetId="figure-236@4.[152,1532,161,1473]" captionTargetPageId="4" captionText="Fig. 3. Sellithyrid brachiopod Neoliothyrina nakremi Bitner sp. nov. from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. A. Holotype, ZPAL V.48/9-1, decorticated shell in ventral (A1), dorsal (A2), and posterior (A4) views; left-lateral view of both valves (A3). B. Paratype, ZPAL V.48/9-2; decorticated shell in ventral (B1), dorsal (B2), and posterior (B4) views; left-lateral view of both valves (B3). C. Paratype, ZPAL V.48/9-3, decorticated shell in ventral (C1), dorsal (C2), and posterior (C4) views; left-lateral view of both valves (C3)." figureDoi="http://doi.org/10.5281/zenodo.10980906" httpUri="https://zenodo.org/record/10980906/files/figure.png" pageId="2" pageNumber="103">3B, C</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FC3DFA90DC80" blockId="2.[862,1336,962,984]" box="[862,1336,962,984]" pageId="2" pageNumber="103">
|
||
<materialsCitation id="0ACD3CBF4658FFCFFCD3FC3DFA9BDC80" box="[862,1331,962,984]" country="Svalbard and Jan Mayen" location="Fossildalen" municipality="Spitsbergen" pageId="2" pageNumber="103" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FC3DFC71DC8F" box="[862,985,962,984]" italics="true" pageId="2" pageNumber="103">
|
||
<typeStatus id="651E88404658FFCFFCD3FC3DFC23DC80" box="[862,907,963,984]" pageId="2" pageNumber="103">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
:
|
||
<location id="BF7A60394658FFCFFC68FC3CFBFDDC80" LSID="urn:lsid:plazi:treatment:320C87F44658FFC9FCD3FDDDFB0FD9E6:BF7A60394658FFCFFC68FC3CFBFDDC80" box="[997,1109,962,984]" country="Svalbard and Jan Mayen" municipality="Spitsbergen" name="Fossildalen" pageId="2" pageNumber="103">Fossildalen</location>
|
||
,
|
||
<collectingMunicipality id="5A7EAC984658FFCFFBD2FC3CFB7ADC80" box="[1119,1234,962,984]" pageId="2" pageNumber="103">Spitsbergen</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="C2B276724658FFCFFB56FC3CFA9BDC80" box="[1243,1331,962,984]" name="Svalbard and Jan Mayen" pageId="2" pageNumber="103">Svalbard</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FC1AFC6EDB4E" blockId="2.[862,1535,996,1046]" pageId="2" pageNumber="103">
|
||
<materialsCitation id="0ACD3CBF4658FFCFFCD3FC1AFC6ADB4E" county="upper Paleocene" location="Cold" municipality="Basilika Formation" pageId="2" pageNumber="103" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FC1AFC70DCA1" box="[862,984,996,1017]" italics="true" pageId="2" pageNumber="103">
|
||
<typeStatus id="651E88404658FFCFFCD3FC1AFC23DCA1" box="[862,907,996,1017]" pageId="2" pageNumber="103">Type</typeStatus>
|
||
horizon
|
||
</emphasis>
|
||
:
|
||
<location id="BF7A60394658FFCFFC69FC1AFBBBDCA2" LSID="urn:lsid:plazi:treatment:320C87F44658FFC9FCD3FDDDFB0FD9E6:BF7A60394658FFCFFC69FC1AFBBBDCA2" box="[996,1043,996,1018]" county="upper Paleocene" municipality="Basilika Formation" name="Cold" pageId="2" pageNumber="103">Cold</location>
|
||
seep carbonates from the
|
||
<collectingMunicipality id="5A7EAC984658FFCFFA8BFC1AFA68DCA2" box="[1286,1472,996,1018]" pageId="2" pageNumber="103">Basilika Formation</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="537B4E6E4658FFCFFA45FC1AFC6ADB4E" pageId="2" pageNumber="103">upper Paleocene</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FBD9FAE0DBDA" blockId="2.[862,1534,1063,1154]" pageId="2" pageNumber="103">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FBD9FC62DB19" box="[862,970,1063,1089]" italics="true" pageId="2" pageNumber="103">Material.</emphasis>
|
||
—
|
||
<materialsCitation id="0ACD3CBF4658FFCFFC6AFBD9FAECDBDA" collectionCode="ZPAL" pageId="2" pageNumber="103" specimenCount="13" typeStatus="holotype">
|
||
<specimenCount id="ACA3FD6B4658FFCFFC6AFBD9FB2EDB19" box="[999,1158,1063,1089]" count="13" pageId="2" pageNumber="103" type="generic">13 specimens</specimenCount>
|
||
(
|
||
<collectionCode id="DCB4AE274658FFCFFB11FBD9FB4BDB19" box="[1180,1251,1063,1089]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
|
||
V.48/9-1–13), complete specimens, apart from the
|
||
<typeStatus id="651E88404658FFCFFB06FBB9FB46DB39" box="[1163,1262,1095,1121]" pageId="2" pageNumber="103" type="holotype">holotype</typeStatus>
|
||
all with broken anterior part, all from the type locality and horizon
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FB6DFBA0DBB6" blockId="2.[862,1534,1170,1262]" pageId="2" pageNumber="103">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FB6DFBAADBF4" box="[862,1026,1171,1196]" italics="true" pageId="2" pageNumber="103">Measurements</emphasis>
|
||
.—
|
||
<typeStatus id="651E88404658FFCFFBAEFB6CFB26DBF4" box="[1059,1166,1170,1196]" pageId="2" pageNumber="103" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectionCode id="DCB4AE274658FFCFFB2CFB6DFB40DBF5" box="[1185,1256,1171,1197]" country="Poland" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="2" pageNumber="103">ZPAL</collectionCode>
|
||
V.48/9-1): L,
|
||
<quantity id="7D5D9B074658FFCFFA1DFB6CFA52DBF5" box="[1424,1530,1170,1197]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="2" pageNumber="103" unit="mm" value="25.6">25.6 mm</quantity>
|
||
; W,
|
||
<quantity id="7D5D9B074658FFCFFC0EFB4DFC4FDB95" box="[899,999,1203,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="2" pageNumber="103" unit="mm" value="19.9">19.9 mm</quantity>
|
||
; H,
|
||
<quantity id="7D5D9B074658FFCFFB9EFB4DFBD1DB95" box="[1043,1145,1203,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="2" pageNumber="103" unit="mm" value="12.9">12.9 mm</quantity>
|
||
; other specimens inappropriate for measurements.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FB01FC49DAC1" blockId="2.[862,1535,1278,1433]" pageId="2" pageNumber="103">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FB01FC70DA40" box="[862,984,1279,1304]" italics="true" pageId="2" pageNumber="103">Diagnosis.</emphasis>
|
||
—Small
|
||
<taxonomicName id="7DA54D614658FFCFFBCFFB00FB76DA40" authorityName="Sahni" authorityYear="1925" box="[1090,1246,1278,1304]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="2" pageNumber="103" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04658FFCFFBCFFB00FB76DA40" box="[1090,1246,1278,1304]" italics="true" pageId="2" pageNumber="103">Neoliothyrina</emphasis>
|
||
</taxonomicName>
|
||
with rectimarginate anterior commissure. Outer hinge plates relatively wide, ventrally concave, inner hinge plates narrow and short, not contacting each other. Crural processes short, bluntly pointed. Loop short.
|
||
</paragraph>
|
||
<caption id="EEDA666A4658FFCFFF1AFA60FD1DDA88" ID-DOI="http://doi.org/10.5281/zenodo.10980902" ID-Zenodo-Dep="10980902" httpUri="https://zenodo.org/record/10980902/files/figure.png" pageId="2" pageNumber="103" startId="2.[151,184,1438,1460]" targetBox="[152,823,162,1413]" targetPageId="2" targetType="figure">
|
||
<paragraph id="BA1A36E24658FFCFFF1AFA60FD1DDA88" blockId="2.[151,824,1437,1488]" pageId="2" pageNumber="103">
|
||
Fig. 1.
|
||
<emphasis id="88D1EAF04658FFCFFF5AFA63FF4FDAEA" bold="true" box="[215,231,1437,1458]" pageId="2" pageNumber="103">A</emphasis>
|
||
. Map of Svalbard showing location of the study area.
|
||
<emphasis id="88D1EAF04658FFCFFD57FA60FD41DAEB" bold="true" box="[730,745,1438,1459]" pageId="2" pageNumber="103">B</emphasis>
|
||
. Map of the study area with the fossil localities indicated (asterisks).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA1A36E24658FFCFFCD3FA54FBE7D9DD" blockId="2.[862,1535,1450,1959]" pageId="2" pageNumber="103">
|
||
<emphasis id="88D1EAF04658FFCFFCD3FA54FC44DA9B" box="[862,1004,1450,1475]" italics="true" pageId="2" pageNumber="103">Description.</emphasis>
|
||
—Shell of medium size, subpentagonal to elongate oval in outline, biconvex with ventral valve deeper. Shell surface smooth with numerous growth lines. Lateral commissure slightly ventrally convex, anterior commissure rectimarginate. Beak suberect with rounded beak ridges. Foramen large, circular, mesothyrid. Symphytium small, only partially visible.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24658FFCFFC09F975FC7CD8FF" blockId="2.[862,1535,1450,1959]" pageId="2" pageNumber="103">Internal characters investigated in transverse serial sections. Cardinal process distinct, flat, surface with numerous grooves. Inner socket ridges thick, parallel to valve margin. Dental sockets moderately deep. Outer hinge plates relatively wide, ventrally concave, separated from inner hinge plates by distinct crural bases. Inner hinge plates short, narrow, not connecting. Crural processes short, tapering, slightly inwardly curving. Loop short, transverse band not preserved.</paragraph>
|
||
<caption id="EEDA666A4659FFCEFFEFFB51FA93DBB9" ID-DOI="http://doi.org/10.5281/zenodo.10980904" ID-Zenodo-Dep="10980904" httpUri="https://zenodo.org/record/10980904/files/figure.png" pageId="3" pageNumber="104" startId="3.[98,131,1199,1221]" targetBox="[99,1481,163,1175]" targetPageId="3" targetType="figure">
|
||
<paragraph id="BA1A36E24659FFCEFFEFFB51FA93DBB9" blockId="3.[98,1482,1199,1249]" pageId="3" pageNumber="104">
|
||
Fig. 2. Transverse serial sections of the sellithyrid brachiopod
|
||
<taxonomicName id="7DA54D614659FFCEFD1FFB51FC35DB9D" authority="Bitner" authorityName="Bitner" authorityYear="2019" box="[658,925,1199,1221]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="nakremi" status="sp. nov.">
|
||
<emphasis id="88D1EAF04659FFCEFD1FFB51FCF6DB9C" box="[658,862,1199,1220]" italics="true" pageId="3" pageNumber="104">Neoliothyrina nakremi</emphasis>
|
||
Bitner
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="93E2578B4659FFCEFC2FFB4EFC40DB9D" box="[930,1000,1200,1221]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype (ZPAL V.48/9-4) from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard. L>20.2 mm. Numbers indicate distance in mm from the tip of the ventral umbo.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA1A36E24659FFCEFFEFFAFFFDE5D8FF" blockId="3.[98,771,1281,1959]" pageId="3" pageNumber="104">
|
||
<emphasis id="88D1EAF04659FFCEFFEFFAFFFF67DA43" box="[98,207,1281,1307]" italics="true" pageId="3" pageNumber="104">Remarks.</emphasis>
|
||
—Probably the first record of this species is
|
||
<taxonomicName id="7DA54D614659FFCEFFEFFADCFE81DA64" box="[98,297,1314,1340]" class="Rhynchonellata" family="Cancellothyrididae" genus="Terebratulina" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="undetermined">
|
||
<emphasis id="88D1EAF04659FFCEFFEFFADCFF57DA64" box="[98,255,1314,1340]" italics="true" pageId="3" pageNumber="104">Terebratulina</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
described and figured by
|
||
<bibRefCitation id="DE344B134659FFCEFDE8FADCFD56DA64" author="Gripp, K." box="[613,766,1314,1340]" pageId="3" pageNumber="104" pagination="3 - 38" refId="ref28977" refString="Gripp, K. 1927. Beitrage zur Geologie von Spitzbergen. Abhandlungen des Naturwissenschaftlichen Vereins zu Hamburg 21: 3 - 38." type="journal article" year="1927">Gripp (1927)</bibRefCitation>
|
||
, but the material is missing and the precise identification from the figure alone is impossible. The material studied herein was initially questionably assigned to
|
||
<taxonomicName id="7DA54D614659FFCEFD0CFA7AFF7FDAE6" authority="Roy, 1980" authorityName="Roy" authorityYear="1980" class="Rhynchonellata" family="Terebratulidae" genus="Pliothyrina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFD0CFA7AFCAADAC6" box="[641,770,1412,1438]" italics="true" pageId="3" pageNumber="104">Pliothyrina</emphasis>
|
||
<bibRefCitation id="DE344B134659FFCEFFEFFA5BFF7FDAE6" author="Roy, A. van" box="[98,215,1444,1471]" pageId="3" pageNumber="104" pagination="1 - 9" refId="ref34777" refString="Roy, A. van 1980. Pliothyrina, genre Terebratulide (Brachiopode) nouveau du Neogene. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 52: 1 - 9." type="journal article" year="1980">Roy, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="DE344B134659FFCEFE97FA5AFD84DAE7" author="Hryniewicz, K. & Bitner, M. A. & Durska, E. & Hagstrom, J. & Hjalmarsdottir H. R. & Jenkins, R. G. & Little, C. T. S. & Miyajima, Y. & Nakrem, H. A. & Kaim, A." box="[282,556,1444,1471]" pageId="3" pageNumber="104" pagination="41 - 56" refId="ref29314" refString="Hryniewicz, K., Bitner, M. A., Durska, E., Hagstrom, J., Hjalmarsdottir H. R., Jenkins, R. G., Little, C. T. S., Miyajima, Y., Nakrem, H. A., and Kaim, A. 2016. Paleocene methane seep and wood-fall marine environments from Spitsbergen, Svalbard. Palaeogeography, Palaeoclimatology, Palaeoecology 462: 41 - 56." type="journal article" year="2016">Hryniewicz et al. 2016</bibRefCitation>
|
||
); however, the investigations of internal structures suggest attribution to
|
||
<taxonomicName id="7DA54D614659FFCEFFEFFA1BFEA8DAA7" authorityName="Sahni" authorityYear="1925" box="[98,256,1509,1535]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFFEFFA1BFEA8DAA7" box="[98,256,1509,1535]" italics="true" pageId="3" pageNumber="104">Neoliothyrina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="7DA54D614659FFCEFE9EFA1BFE3CDAA7" authorityName="Roy" authorityYear="1980" box="[275,404,1509,1535]" class="Rhynchonellata" family="Terebratulidae" genus="Pliothyrina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFE9EFA1BFE3CDAA7" box="[275,404,1509,1535]" italics="true" pageId="3" pageNumber="104">Pliothyrina</emphasis>
|
||
</taxonomicName>
|
||
is known from the Oligocene to Pliocene in northern Europe (
|
||
<bibRefCitation id="DE344B134659FFCEFD88F9F8FD0ED978" author="Cooper, A. G." box="[517,678,1542,1568]" pageId="3" pageNumber="104" pagination="1 - 445" refId="ref26750" refString="Cooper, A. G. 1983. The Terebratulacea (Brachiopoda) Triassic to Recent: a study of the brachidia (loops). Smithsonian Contributions to Paleobiology 50: 1 - 445." type="journal article" year="1983">Cooper 1983</bibRefCitation>
|
||
), while
|
||
<taxonomicName id="7DA54D614659FFCEFFEFF9D9FEA8D919" authorityName="Sahni" authorityYear="1925" box="[98,256,1575,1601]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFFEFF9D9FEA8D919" box="[98,256,1575,1601]" italics="true" pageId="3" pageNumber="104">Neoliothyrina</emphasis>
|
||
</taxonomicName>
|
||
is known from the Upper Cretaceous (Santonian–Maastrichtian) of
|
||
<collectingCountry id="C2B276724659FFCEFE11F9B9FDEAD939" box="[412,578,1607,1633]" name="United Kingdom" pageId="3" pageNumber="104">Great Britain</collectingCountry>
|
||
,
|
||
<collectingCountry id="C2B276724659FFCEFDDBF9B9FD6CD939" box="[598,708,1607,1633]" name="Germany" pageId="3" pageNumber="104">Germany</collectingCountry>
|
||
, and
|
||
<collectingCountry id="C2B276724659FFCEFFEFF996FF19D9DA" box="[98,177,1640,1666]" name="Poland" pageId="3" pageNumber="104">Poland</collectingCountry>
|
||
(
|
||
<bibRefCitation id="DE344B134659FFCEFF41F996FED3D9DB" author="Steinich, G." box="[204,379,1640,1667]" pageId="3" pageNumber="104" pagination="1 - 220" refId="ref36507" refString="Steinich, G. 1965. Die artikulaten Brachiopoden der Rugener Schreibkreide (Unter-Maastricht). Palaontologische Abhandlungen 2 (1): 1 - 220." type="journal article" year="1965">Steinich 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B134659FFCEFE1EF996FD3DD9DA" author="Popiel-Barczyk, E." box="[403,661,1640,1666]" pageId="3" pageNumber="104" pagination="3 - 86" refId="ref34248" refString="Popiel-Barczyk, E. 1968. Upper Cretaceous terebratulids (Brachiopoda) from the Middle Vistula Gorge. Prace Muzeum Ziemi 12: 3 - 86." type="journal article" year="1968">Popiel-Barczyk 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B134659FFCEFD20F996FF33D9FB" author="Cooper, A. G." pageId="3" pageNumber="104" pagination="1 - 445" refId="ref26750" refString="Cooper, A. G. 1983. The Terebratulacea (Brachiopoda) Triassic to Recent: a study of the brachidia (loops). Smithsonian Contributions to Paleobiology 50: 1 - 445." type="journal article" year="1983">Cooper 1983</bibRefCitation>
|
||
). Both genera are characterized by the presence of inner hinge plates, a feature rarely present in terebratuloids (
|
||
<bibRefCitation id="DE344B134659FFCEFF2AF934FE74D9BD" author="MacKinnon, D. I. & Lee, D. E." box="[167,476,1738,1765]" pageId="3" pageNumber="104" pagination="1974 - 1993" refId="ref32102" refString="MacKinnon, D. I. and Lee, D. E. 2006. Loop morphology and terminology in Terebratulida. In: R. L. Kaesler (ed.), Treatise of Invertebrate Paleontology. Part H Brachiopoda revised, Vol. 5, 1974 - 1993. Geological Society of America, Boulder and University of Kansas, Lawrence." type="journal article" year="2006">MacKinnon and Lee 2006</bibRefCitation>
|
||
).
|
||
<taxonomicName id="7DA54D614659FFCEFE62F934FDD8D9BC" authorityName="Roy" authorityYear="1980" box="[495,624,1738,1764]" class="Rhynchonellata" family="Terebratulidae" genus="Pliothyrina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFE62F934FDD8D9BC" box="[495,624,1738,1764]" italics="true" pageId="3" pageNumber="104">Pliothyrina</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="7DA54D614659FFCEFFEFF914FEA8D85C" authorityName="Sahni" authorityYear="1925" box="[98,256,1770,1796]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFFEFF914FEA8D85C" box="[98,256,1770,1796]" italics="true" pageId="3" pageNumber="104">Neoliothyrina</emphasis>
|
||
</taxonomicName>
|
||
in having very narrow or absent outer hinge plates and its crural processes are unusually long and curved anteromedially (
|
||
<bibRefCitation id="DE344B134659FFCEFEF1F8D2FDBED81E" author="Cooper, A. G." box="[380,534,1836,1862]" pageId="3" pageNumber="104" pagination="1 - 445" refId="ref26750" refString="Cooper, A. G. 1983. The Terebratulacea (Brachiopoda) Triassic to Recent: a study of the brachidia (loops). Smithsonian Contributions to Paleobiology 50: 1 - 445." type="journal article" year="1983">Cooper 1983</bibRefCitation>
|
||
), whereas the outer hinge plates are well-developed and the crural processes are bluntly pointed in
|
||
<taxonomicName id="7DA54D614659FFCEFEE8F893FDABD8DF" authorityName="Sahni" authorityYear="1925" box="[357,515,1901,1927]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF04659FFCEFEE8F893FDABD8DF" box="[357,515,1901,1927]" italics="true" pageId="3" pageNumber="104">Neoliothyrina</emphasis>
|
||
</taxonomicName>
|
||
. These characters are all observed in the material from
|
||
<collectingCountry id="C2B276724659FFCEFE6DF873FDE1D8FF" box="[480,585,1933,1959]" name="Svalbard and Jan Mayen" pageId="3" pageNumber="104">Svalbard</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24659FFCEFCC3FAFCFB7DD958" blockId="3.[808,1481,1281,1960]" pageId="3" pageNumber="104">
|
||
Due to the rectimarginate anterior commissure, very narrow and short inner hinge plate, and small crural processes,
|
||
<taxonomicName id="7DA54D614659FFCEFCA5FABCFBDDDA04" authority="Bitner" authorityName="Bitner" authorityYear="2019" box="[808,1141,1346,1372]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="nakremi" status="sp. nov.">
|
||
<emphasis id="88D1EAF04659FFCEFCA5FABCFB80DA04" box="[808,1064,1346,1372]" italics="true" pageId="3" pageNumber="104">Neoliothyrina nakremi</emphasis>
|
||
Bitner
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="93E2578B4659FFCEFBF0FABDFB7BDA05" box="[1149,1235,1347,1373]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
|
||
closely resembles the Maastrichtian
|
||
<taxonomicName id="7DA54D614659FFCEFC5FFA9DFA6CDA25" authority="Popiel-Barczyk, 1968" authorityName="Popiel-Barczyk" authorityYear="1968" box="[978,1476,1379,1405]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="plana">
|
||
<emphasis id="88D1EAF04659FFCEFC5FFA9DFB14DA25" box="[978,1212,1379,1405]" italics="true" pageId="3" pageNumber="104">Neoliothyrina plana</emphasis>
|
||
<bibRefCitation id="DE344B134659FFCEFB44FA9DFA6CDA25" author="Popiel-Barczyk, E." box="[1225,1476,1379,1405]" pageId="3" pageNumber="104" pagination="3 - 86" refId="ref34248" refString="Popiel-Barczyk, E. 1968. Upper Cretaceous terebratulids (Brachiopoda) from the Middle Vistula Gorge. Prace Muzeum Ziemi 12: 3 - 86." type="journal article" year="1968">Popiel-Barczyk, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="C2B276724659FFCEFCEBFA7AFC1EDAC6" box="[870,950,1412,1438]" name="Poland" pageId="3" pageNumber="104">Poland</collectingCountry>
|
||
.
|
||
<taxonomicName id="7DA54D614659FFCEFC4DFA7AFB16DAC6" authorityName="Bitner" authorityYear="2019" box="[960,1214,1412,1438]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="nakremi">
|
||
<emphasis id="88D1EAF04659FFCEFC4DFA7AFB16DAC6" box="[960,1214,1412,1438]" italics="true" pageId="3" pageNumber="104">Neoliothyrina nakremi</emphasis>
|
||
</taxonomicName>
|
||
is, however, more elongate and more pentagonal compared with
|
||
<taxonomicName id="7DA54D614659FFCEFB7BFA5BFAFCDAE6" authorityName="Popiel-Barczyk" authorityYear="1968" box="[1270,1364,1444,1470]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="plana">
|
||
<emphasis id="88D1EAF04659FFCEFB7BFA5BFAFCDAE6" box="[1270,1364,1444,1470]" italics="true" pageId="3" pageNumber="104">N. plana</emphasis>
|
||
</taxonomicName>
|
||
. Thus, the present finding extends the stratigraphical and geographical range of this genus into the Paleocene.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E24659FFC9FCC3F9F8FE79D9E7" blockId="3.[808,1481,1281,1960]" lastBlockId="4.[151,824,1634,1959]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
|
||
The investigated specimens differ strongly from the
|
||
<typeStatus id="651E88404659FFCEFA1AF9F9FA61D979" box="[1431,1481,1543,1569]" pageId="3" pageNumber="104">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="7DA54D614659FFCEFC0AF9D9FA8CD91A" authority="Davidson, 1852" authorityName="Davidson" authorityYear="1852" box="[903,1316,1575,1602]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="obesa">
|
||
<emphasis id="88D1EAF04659FFCEFC0AF9D9FBC2D919" box="[903,1130,1575,1601]" italics="true" pageId="3" pageNumber="104">Neoliothyrina obesa</emphasis>
|
||
<bibRefCitation id="DE344B134659FFCEFBFFF9D9FA8CD91A" author="Davidson, T." box="[1138,1316,1575,1602]" pageId="3" pageNumber="104" pagination="1 - 54" refId="ref27770" refString="Davidson, T. 1852. A Monograph of British Cretaceous Brachiopoda. Part II. Palaeontographical Society Monograph 6: 1 - 54." type="journal article" year="1852">Davidson, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="7DA54D614659FFCEFAA3F9D9FCC1D939" authorityName="Davidson" authorityYear="1852" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="obesa">
|
||
<emphasis id="88D1EAF04659FFCEFAA3F9D9FCC1D939" italics="true" pageId="3" pageNumber="104">Neoliothyrina obesa</emphasis>
|
||
</taxonomicName>
|
||
is much larger than
|
||
<taxonomicName id="7DA54D614659FFCEFBEFF9B6FB4BD939" authorityName="Bitner" authorityYear="2019" box="[1122,1251,1607,1633]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="nakremi">
|
||
<emphasis id="88D1EAF04659FFCEFBEFF9B6FB4BD939" box="[1122,1251,1607,1633]" italics="true" pageId="3" pageNumber="104">N. nakremi</emphasis>
|
||
</taxonomicName>
|
||
, having its anterior commissure distinctly biplicate and its inner hinge plates are well developed, contacting or even overlapping each other in many specimens (
|
||
<bibRefCitation id="DE344B134659FFCEFBADF957FB69D99C" author="Steinich, G." box="[1056,1217,1705,1732]" pageId="3" pageNumber="104" pagination="1 - 220" refId="ref36507" refString="Steinich, G. 1965. Die artikulaten Brachiopoden der Rugener Schreibkreide (Unter-Maastricht). Palaontologische Abhandlungen 2 (1): 1 - 220." type="journal article" year="1965">Steinich 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B134659FFCEFB5FF957FA6DD99B" author="Popiel-Barczyk, E." box="[1234,1477,1705,1731]" pageId="3" pageNumber="104" pagination="3 - 86" refId="ref34248" refString="Popiel-Barczyk, E. 1968. Upper Cretaceous terebratulids (Brachiopoda) from the Middle Vistula Gorge. Prace Muzeum Ziemi 12: 3 - 86." type="journal article" year="1968">Popiel-Barczyk 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B134659FFCEFCA5F934FC14D9BC" author="Cooper, A. G." box="[808,956,1738,1764]" pageId="3" pageNumber="104" pagination="1 - 445" refId="ref26750" refString="Cooper, A. G. 1983. The Terebratulacea (Brachiopoda) Triassic to Recent: a study of the brachidia (loops). Smithsonian Contributions to Paleobiology 50: 1 - 445." type="journal article" year="1983">Cooper 1983</bibRefCitation>
|
||
). Also,
|
||
<taxonomicName id="7DA54D614659FFCEFB9BF934FA61D9BD" authority="(Lundgren, 1885)" baseAuthorityName="Lundgren" baseAuthorityYear="1885" box="[1046,1481,1738,1765]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="fallax">
|
||
<emphasis id="88D1EAF04659FFCEFB9BF934FB51D9BC" box="[1046,1273,1738,1764]" italics="true" pageId="3" pageNumber="104">Neoliothyrina fallax</emphasis>
|
||
(
|
||
<bibRefCitation id="DE344B134659FFCEFA86F934FA6AD9BD" author="Lundgren, B." box="[1291,1474,1738,1765]" pageId="3" pageNumber="104" pagination="1 - 72" refId="ref32080" refString="Lundgren, B. 1885. Undersokningar ofver Brachiopoderna i Sveriges Kritsystem. Lunds Universitets Arskrift 20: 1 - 72." type="journal article" year="1885">Lundgren, 1885</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has a larger maximum size than the taxon from
|
||
<collectingCountry id="C2B276724659FFCEFAEBF915FA61D85D" box="[1382,1481,1771,1797]" name="Svalbard and Jan Mayen" pageId="3" pageNumber="104">Svalbard</collectingCountry>
|
||
(
|
||
<bibRefCitation id="DE344B134659FFCEFCBDF8F5FB91D87D" author="Brunnich Nielsen, K." box="[816,1081,1803,1829]" pageId="3" pageNumber="104" pagination="127 - 178" refId="ref26052" refString="Brunnich Nielsen, K. 1909. Brachiopoderne i Danmarks kridtaflejringer. Det Kongelige Danske Videnskabernes Selskabs Skrifter, Naturvidenskabelig og Mathematisk Afdeling 6: 127 - 178." type="journal article" year="1909">Brünnich Nielsen 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B134659FFCEFBCBF8F5FA98D87E" author="Popiel-Barczyk, E." box="[1094,1328,1803,1830]" pageId="3" pageNumber="104" pagination="3 - 86" refId="ref34248" refString="Popiel-Barczyk, E. 1968. Upper Cretaceous terebratulids (Brachiopoda) from the Middle Vistula Gorge. Prace Muzeum Ziemi 12: 3 - 86." type="journal article" year="1968">Popiel-Barczyk 1968</bibRefCitation>
|
||
) and exhibits a more curved beak and biplicate anterior commissure. The species
|
||
<taxonomicName id="7DA54D614659FFCEFC09F8B2FBC7D83F" baseAuthorityName="Hagenow" baseAuthorityYear="1842" box="[900,1135,1868,1895]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="fittoni">
|
||
<emphasis id="88D1EAF04659FFCEFC09F8B2FBC7D83F" box="[900,1135,1868,1895]" italics="true" pageId="3" pageNumber="104">Neoliothyrina fittoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="DE344B134659FFCEFB0FF8B3FA9FD83F" author="Hagenow, F. von" box="[1154,1335,1869,1895]" pageId="3" pageNumber="104" pagination="528 - 575" refId="ref29035" refString="Hagenow, F. von 1842. Monographie der Rug'schen Kreide-Versteinerungen. III. Abteilung: Mollusken. Neues Jahrbuch fur Mineralogie, Geognosie, Geologie und Petrefaktenkunde 5: 528 - 575." type="journal article" year="1842">Hagenow, 1842</bibRefCitation>
|
||
, assigned to this genus by
|
||
<bibRefCitation id="DE344B134659FFCEFC4FF893FBF6D8D0" author="Steinich, G." box="[962,1118,1901,1928]" pageId="3" pageNumber="104" pagination="1 - 220" refId="ref36507" refString="Steinich, G. 1965. Die artikulaten Brachiopoden der Rugener Schreibkreide (Unter-Maastricht). Palaontologische Abhandlungen 2 (1): 1 - 220." type="journal article" year="1965">Steinich 1965</bibRefCitation>
|
||
), is similar in size to
|
||
<taxonomicName id="7DA54D614659FFCEFAC1F890FA61D8DF" authorityName="Bitner" authorityYear="2019" box="[1356,1481,1901,1927]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="nakremi">
|
||
<emphasis id="88D1EAF04659FFCEFAC1F890FA61D8DF" box="[1356,1481,1901,1927]" italics="true" pageId="3" pageNumber="104">N. nakremi</emphasis>
|
||
</taxonomicName>
|
||
but differs in the character of cardinal process; in
|
||
<taxonomicName id="7DA54D614659FFCEFAD0F871FA6AD8F0" baseAuthorityName="Hagenow" baseAuthorityYear="1842" box="[1373,1474,1934,1960]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="3" pageNumber="104" phylum="Brachiopoda" rank="species" species="fittoni">
|
||
<emphasis id="88D1EAF04659FFCEFAD0F871FA6AD8F0" box="[1373,1474,1934,1960]" italics="true" pageId="3" pageNumber="104">N. fittoni</emphasis>
|
||
</taxonomicName>
|
||
, the cardinal process has a distinct cardinal knob that is very poorly developed in other
|
||
<taxonomicName id="7DA54D61465EFFC9FE5CF97DFD61D9C5" box="[465,713,1667,1693]" pageId="4" pageNumber="105">
|
||
<emphasis id="88D1EAF0465EFFC9FE5CF97DFDC4D9C5" box="[465,620,1667,1693]" italics="true" pageId="4" pageNumber="105">Neoliothyrina</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="DE344B13465EFFC9FD51F97DFF67D9E7" author="Steinich, G." pageId="4" pageNumber="105" pagination="1 - 220" refId="ref36507" refString="Steinich, G. 1965. Die artikulaten Brachiopoden der Rugener Schreibkreide (Unter-Maastricht). Palaontologische Abhandlungen 2 (1): 1 - 220." type="journal article" year="1965">Steinich 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DE344B13465EFFC9FF57F95AFE6DD9E7" author="Popiel-Barczyk, E." box="[218,453,1700,1727]" pageId="4" pageNumber="105" pagination="3 - 86" refId="ref34248" refString="Popiel-Barczyk, E. 1968. Upper Cretaceous terebratulids (Brachiopoda) from the Middle Vistula Gorge. Prace Muzeum Ziemi 12: 3 - 86." type="journal article" year="1968">Popiel-Barczyk 1968</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="EEDA666A465EFFC9FF1AFA24FBC1D91C" ID-DOI="http://doi.org/10.5281/zenodo.10980906" ID-Zenodo-Dep="10980906" httpUri="https://zenodo.org/record/10980906/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,184,1498,1520]" targetBox="[152,1532,161,1473]" targetPageId="4" targetType="figure">
|
||
<paragraph id="BA1A36E2465EFFC9FF1AFA24FBC1D91C" blockId="4.[151,1536,1498,1606]" pageId="4" pageNumber="105">
|
||
Fig. 3. Sellithyrid brachiopod
|
||
<taxonomicName id="7DA54D61465EFFC9FE21FA24FD11DAA8" authority="Bitner" authorityName="Bitner" authorityYear="2019" box="[428,697,1498,1520]" class="Rhynchonellata" family="Terebratulidae" genus="Neoliothyrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="105" phylum="Brachiopoda" rank="species" species="nakremi" status="sp. nov.">
|
||
<emphasis id="88D1EAF0465EFFC9FE21FA24FDD1DAB7" box="[428,633,1498,1519]" italics="true" pageId="4" pageNumber="105">Neoliothyrina nakremi</emphasis>
|
||
Bitner
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="93E2578B465EFFC9FD32FA25FCAFDAA8" box="[703,775,1499,1520]" pageId="4" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the upper Paleocene, Basilika Formation, Fossildalen, Spitsbergen, Svalbard.
|
||
<emphasis id="88D1EAF0465EFFC9FF1AFA08FF0FD953" bold="true" box="[151,167,1526,1547]" pageId="4" pageNumber="105">A</emphasis>
|
||
. Holotype, ZPAL V.48/9-1, decorticated shell in ventral (A
|
||
<subScript id="262134A7465EFFC9FD49F9FEFD64D956" attach="left" box="[708,716,1536,1550]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
|
||
), dorsal (A
|
||
<subScript id="262134A7465EFFC9FCB8F9FEFC95D956" attach="right" box="[821,829,1536,1550]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
|
||
), and posterior (A
|
||
<subScript id="262134A7465EFFC9FC6BF9FEFC46D956" attach="right" box="[998,1006,1536,1550]" fontSize="6" pageId="4" pageNumber="105">4</subScript>
|
||
) views; left-lateral view of both valves (A
|
||
<subScript id="262134A7465EFFC9FAFEF9FEFAD3D956" attach="left" box="[1395,1403,1536,1550]" fontSize="6" pageId="4" pageNumber="105">3</subScript>
|
||
).
|
||
<emphasis id="88D1EAF0465EFFC9FA02FA08FA36D953" bold="true" box="[1423,1438,1526,1547]" pageId="4" pageNumber="105">B</emphasis>
|
||
. Paratype, ZPAL V.48/9-2; decorticated shell in ventral (B
|
||
<subScript id="262134A7465EFFC9FDCCF9E2FDE1D972" attach="left" box="[577,585,1564,1578]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
|
||
), dorsal (B
|
||
<subScript id="262134A7465EFFC9FD22F9E2FD1FD972" attach="left" box="[687,695,1564,1578]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
|
||
), and posterior (B
|
||
<subScript id="262134A7465EFFC9FCD6F9E2FCCBD972" attach="left" box="[859,867,1564,1578]" fontSize="6" pageId="4" pageNumber="105">4</subScript>
|
||
) views; left-lateral view of both valves (B
|
||
<subScript id="262134A7465EFFC9FB6DF9E2FB40D972" attach="left" box="[1248,1256,1564,1578]" fontSize="6" pageId="4" pageNumber="105">3</subScript>
|
||
).
|
||
<emphasis id="88D1EAF0465EFFC9FB76F9ECFAA3D970" bold="true" box="[1275,1291,1554,1576]" pageId="4" pageNumber="105">C</emphasis>
|
||
. Paratype, ZPAL V.48/9-3, decorticated shell in ventral (C
|
||
<subScript id="262134A7465EFFC9FE22F9C6FE1FD91E" attach="left" box="[431,439,1592,1606]" fontSize="6" pageId="4" pageNumber="105">1</subScript>
|
||
), dorsal (C
|
||
<subScript id="262134A7465EFFC9FD90F9C6FD8DD91E" attach="left" box="[541,549,1592,1606]" fontSize="6" pageId="4" pageNumber="105">2</subScript>
|
||
), and posterior (C
|
||
<subScript id="262134A7465EFFC9FD46F9C6FD7BD91E" attach="left" box="[715,723,1592,1606]" fontSize="6" pageId="4" pageNumber="105">4</subScript>
|
||
) views; left-lateral view of both valves (C
|
||
<subScript id="262134A7465EFFC9FBD9F9C6FBF4D91E" attach="left" box="[1108,1116,1592,1606]" fontSize="6" pageId="4" pageNumber="105">3</subScript>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA1A36E2465EFFC9FF30F938FD43D8FF" blockId="4.[151,824,1634,1959]" pageId="4" pageNumber="105">
|
||
The specimens assigned by
|
||
<bibRefCitation id="DE344B13465EFFC9FE62F938FDD6D9B8" author="Gripp, K." box="[495,638,1734,1760]" pageId="4" pageNumber="105" pagination="3 - 38" refId="ref28977" refString="Gripp, K. 1927. Beitrage zur Geologie von Spitzbergen. Abhandlungen des Naturwissenschaftlichen Vereins zu Hamburg 21: 3 - 38." type="journal article" year="1927">Gripp (1927)</bibRefCitation>
|
||
to
|
||
<taxonomicName id="7DA54D61465EFFC9FD10F938FEF0D859" authority="d'Orbigny, 1847" authorityName="d'Orbigny" authorityYear="1847" class="Rhynchonellata" family="Cancellothyrididae" genus="Terebratulina" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="105" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF0465EFFC9FD10F938FC90D9B8" box="[669,824,1734,1760]" italics="true" pageId="4" pageNumber="105">Terebratulina</emphasis>
|
||
<bibRefCitation id="DE344B13465EFFC9FF1AF919FEF0D859" author="d'Orbigny, C. A." box="[151,344,1767,1793]" pageId="4" pageNumber="105" refId="ref28131" refString="d'Orbigny, C. A. 1847. Paleontologie Francaise. Description des Molluques et Rayonnes fossiles. Terrains Cretaces. 4. Brachiopodes. 390 pp. Victor Masson, Paris." type="book" year="1847">d’Orbigny, 1847</bibRefCitation>
|
||
</taxonomicName>
|
||
, undoubtedly belong to smooth, shortlooped terebratulides and most probably represent the new species described here. However, the genus
|
||
<taxonomicName id="7DA54D61465EFFC9FD10F8D7FC90D81B" authorityName="d'Orbigny" authorityYear="1847" box="[669,824,1833,1859]" class="Rhynchonellata" family="Cancellothyrididae" genus="Terebratulina" kingdom="Animalia" order="Terebratulida" pageId="4" pageNumber="105" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="88D1EAF0465EFFC9FD10F8D7FC90D81B" box="[669,824,1833,1859]" italics="true" pageId="4" pageNumber="105">Terebratulina</emphasis>
|
||
</taxonomicName>
|
||
is characterized by a ribbed surface, not seen on
|
||
<bibRefCitation id="DE344B13465EFFC9FD69F8B5FF77D8DE" author="Gripp, K." pageId="4" pageNumber="105" pagination="3 - 38" refId="ref28977" refString="Gripp, K. 1927. Beitrage zur Geologie von Spitzbergen. Abhandlungen des Naturwissenschaftlichen Vereins zu Hamburg 21: 3 - 38." type="journal article" year="1927">Gripp’s (1927)</bibRefCitation>
|
||
figures, and an incomplete foramen with disjunct deltidial plates, absent in the material studied herein.
|
||
</paragraph>
|
||
<paragraph id="BA1A36E2465EFFC9FCD3F99CFB0FD9E6" blockId="4.[862,1534,1634,1726]" pageId="4" pageNumber="105">
|
||
<emphasis id="88D1EAF0465EFFC9FCD3F99CFAB2D924" box="[862,1306,1634,1660]" italics="true" pageId="4" pageNumber="105">Stratigraphic and geographic range.</emphasis>
|
||
—Upper Paleocene cold seep carbonates from the Basilika Formation, Fossildalen, Spitsbergen,
|
||
<collectingCountry id="C2B27672465EFFC9FBB1F95AFB0BD9E6" box="[1084,1187,1700,1726]" name="Svalbard and Jan Mayen" pageId="4" pageNumber="105">Svalbard</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |