178 lines
16 KiB
XML
178 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="319C2A95EB0951588C6E881DEA4919AF" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia osaensis Kriebel 2016, comb. nov." docType="treatment" docVersion="3" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="201" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="201" updateTime="1668141747722" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kriebel, Ricardo</mods:namePart>
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<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-07-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>67</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>326</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
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<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182226950" LSID="urn:lsid:plazi:treatment:319C2A95EB0951588C6E881DEA4919AF" httpUri="http://treatment.plazi.org/id/319C2A95EB0951588C6E881DEA4919AF" lastPageNumber="201" pageId="200" pageNumber="201">
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<subSubSection pageId="200" pageNumber="201" type="nomenclature">
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<paragraph pageId="200" pageNumber="201">
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<taxonomicName LSID="319C2A95-EB09-5158-8C6E-881DEA4919AF" authority="(Aguilar, Kriebel & Almeda) Kriebel" authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis" status="comb. nov.">Conostegia osaensis (Aguilar, Kriebel & Almeda) Kriebel</taxonomicName>
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<taxonomicNameLabel pageId="200" pageNumber="201">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 178" captionStartId="F178" captionText="Figure 178. Conostegia osaensis. A Habit and detail of leaf abaxial surface B Flowering branch C Close up of the flower D Infructescence. Photographs taken Reinaldo Aguilar and vouchered R. Aguilar 10200." figureDoi="10.3897/phytokeys.67.6703.figure178" httpUri="https://binary.pensoft.net/fig/96536" pageId="200" pageNumber="201">Fig. 178</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="200" pageNumber="201" type="reference_group">
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<paragraph pageId="200" pageNumber="201">
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis" status="comb. nov.">Conostegia osaensis</taxonomicName>
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(Aguilar, Kriebel & Almeda) Kriebel. Basionym:
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<taxonomicName authorityName="Kriebel" authorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis" status="comb. nov.">Miconia osaensis</taxonomicName>
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Aguilar, Kriebel and Almeda, Proc. Calif. Acad. Sci. Series 4, 59(10): 490, f. 1A-J, 2A-D. 2008. Type: Costa Rica. Puntarenas:
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<normalizedToken originalValue="Cantón">Canton</normalizedToken>
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de Osa, Reserva Forestal Golfo Dulce, entrada a Chocuaco, por la casa de Moncho, 200-350 m, 28 May 1997, R. Aguilar 5145 (holotype: INB!; isotypes: CAS!, CR!, MO!, USJ).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="200" pageNumber="201" type="description">
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<paragraph pageId="200" pageNumber="201">Description.</paragraph>
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<paragraph pageId="200" pageNumber="201">
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Trees 9-25 m tall with the uppermost flattened and two sided branchlets completely covered with a stellulate-lepidote indumenta; the nodal line present. Leaves of a pair equal to subequal in size. Petioles 1.5-4.5 cm long. Leaf blades 7.5-21
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<normalizedToken originalValue="×">x</normalizedToken>
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3-7 cm, 5-plinerved, with the innermost pair of primary veins diverging ca. 0.5-1.5 cm above the blade base in opposite, alternate or subalternate fashion, elliptic to elliptic-oblong or slightly elliptic-lanceolate, the base acute, the apex acute to acuminate, the margin entire to inconspicuously crenulate-denticulate, the adaxial surface glabrous and inconspicuously glandular puncticulate, the abaxial surface completely covered by peltate scales. Inflorescence a terminal panicle 10.8-15 cm long branching above the base, accessory branches present, reddish stellulate-lepidote indument throughout; bracts and bracteoles 1-1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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0.25-0.5 mm, triangular, persistent. Pedicels 0.25-0.5 mm. Flowers 5-merous, calyx calyptrate, flower buds 3-4.5
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<normalizedToken originalValue="×">x</normalizedToken>
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2-2.75 mm, elliptic-ovate, calyx teeth, the base rounded, the apex acute, not constricted, the hypanthial and calyptrate portions differentiated in texture, color and indument different, the calyptra being white or translucent, very thin and more sparsely pubescent than the hypanthium, the latter campanulate to urceolate, 2.5-3
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<normalizedToken originalValue="×">x</normalizedToken>
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2.25-2.75 mm, densely covered with peltate scales that grade into stellate hairs, the torus also beset with scales. Petals 5-8
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<normalizedToken originalValue="×">x</normalizedToken>
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3-6 mm, white, obtriangular, spreading at anthesis, emarginate at the apex, glabrous. Stamens 10, Stamens ca. 5-8 mm long, androecium slightly bilaterally symmetric, the filaments 2.5-4.5 mm, white, 3.25-3.75
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<normalizedToken originalValue="×">x</normalizedToken>
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1-1.5 m, oblong, yellow, laterally compressed, the connective dorsally thickened, the pore ca. 0.2 mm wide, slightly ventrally inclined. Ovary 5-locular, inferior, apically glabrous and slightly ribbed, forming a low collar around the style base, with pronounced elevated lines. Style ca. 5 mm long, straight and abruptly curved near the apex, vertical distance from the anther to the stigma ca. 0-0.5 mm, stigma punctiform to truncate, ca. 0.8 mm wide. Berry ca. 6
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<normalizedToken originalValue="×">x</normalizedToken>
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6 mm, purple black. Seeds 1.3-1.9 mm long, broadly pyramidate, rounded to bluntly angled on the convex face, the testa smooth.
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</paragraph>
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<caption ID-Zenodo-Dep="974011" doi="10.3897/phytokeys.67.6703.figure178" httpUri="https://binary.pensoft.net/fig/96536" pageId="200" pageNumber="201" start="Figure 178" startId="F178">
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<paragraph pageId="200" pageNumber="201">
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<emphasis bold="true" pageId="200" pageNumber="201">Figure 178.</emphasis>
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="200" pageNumber="201">A</emphasis>
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Habit and detail of leaf abaxial surface
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<emphasis bold="true" pageId="200" pageNumber="201">B</emphasis>
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Flowering branch
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<emphasis bold="true" pageId="200" pageNumber="201">C</emphasis>
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Close up of the flower
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<emphasis bold="true" pageId="200" pageNumber="201">D</emphasis>
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Infructescence. Photographs taken Reinaldo Aguilar and vouchered
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<emphasis italics="true" pageId="200" pageNumber="201">R. Aguilar 10200</emphasis>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="200" pageNumber="201" type="distribution">
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<paragraph pageId="200" pageNumber="201">Distribution</paragraph>
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<paragraph pageId="200" pageNumber="201">
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(Fig.
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<figureCitation captionStart="Figure 179" captionStartId="F179" captionText="Figure 179. Distribution of Conostegia osaensis." figureDoi="10.3897/phytokeys.67.6703.figure179" httpUri="https://binary.pensoft.net/fig/96537" pageId="200" pageNumber="201">179</figureCitation>
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). Endemic to the Osa Peninsula in southern Costa Rica, 40-200 m in elevation.
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</paragraph>
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<caption ID-Zenodo-Dep="974013" doi="10.3897/phytokeys.67.6703.figure179" httpUri="https://binary.pensoft.net/fig/96537" pageId="200" pageNumber="201" start="Figure 179" startId="F179">
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<paragraph pageId="200" pageNumber="201">
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<emphasis bold="true" pageId="200" pageNumber="201">Figure 179.</emphasis>
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Distribution of
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph pageId="200" pageNumber="201">
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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is endemic to the Osa Peninsula of Costa Rica and quite distinctive as it tends to be a large tree up to about 25 m tall. In addition, the leaf abaxial surface is covered with a stellate lepidote indument and the flowers are calyptrate, not pleiostemonous, and have a short style. In the molecular phylogeny
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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falls sister to the clade comprised of
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="plumosa">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia plumosa</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Naudin" authorityYear="1850" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia speciosa" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="speciosa">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia speciosa</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia subcrustulata" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="subcrustulata">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia subcrustulata</emphasis>
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</taxonomicName>
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, and
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="xalapensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia xalapensis</emphasis>
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</taxonomicName>
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. With these taxa
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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shares the calyptrate calyx and the short style.
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Aguilar, Kriebel & Almeda" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia osaensis" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="species" species="osaensis">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia osaensis</emphasis>
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</taxonomicName>
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has the largest seeds of any species in
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia" order="Myrtales" pageId="200" pageNumber="201" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="200" pageNumber="201">Conostegia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="200" pageNumber="201" type="materials_examined">
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<paragraph pageId="200" pageNumber="201">Specimens examined.</paragraph>
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<paragraph pageId="200" pageNumber="201">
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<emphasis bold="true" pageId="200" pageNumber="201">
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<collectingCountry name="Costa Rica">COSTA RICA</collectingCountry>
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.
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<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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</emphasis>
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:
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<normalizedToken originalValue="Rincón">Rincon</normalizedToken>
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, cerca de Banegas,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
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al este del centro del pueblo de Banegas,
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<normalizedToken originalValue="Estación">Estacion</normalizedToken>
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<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
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Los Charcos, sendero Dendrobates, Aguilar 10200, 10228 (INB); Mogos,
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<normalizedToken originalValue="Bahía">Bahia</normalizedToken>
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Chal, entrada a Chocuaco a
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<quantity metricMagnitude="4" metricUnit="m" metricValue="3.5" unit="km" value="35.0">35 km</quantity>
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de Chacarita, finca de Carlos Rojas, Aguilar 6208 (INB).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |