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<mods:title id="DFA032054E6C2A68578C2AA120E18510">Systematic revision of Aluntiini Emeljanov, 1979 (Hemiptera: Fulgoromorpha: Dictyopharidae: Dictyopharinae): reclassification, phylogenetic analysis, and biogeography</mods:title>
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<mods:namePart id="48CBA055C98C67C74026F852E5C1A565">Song, Zhi-Shun</mods:namePart>
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<mods:namePart id="2663727B14954E06ECF32CE16353ED02">Liang, Ai-Ping</mods:namePart>
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<paragraph id="B97236EAFFFEFFDCFCC74F89FA8B9BA2" blockId="19.[878,1389,429,451]" box="[878,1389,429,451]" pageId="19" pageNumber="368">
<heading id="E23A8186FFFEFFDCFCC74F89FA8B9BA2" box="[878,1389,429,451]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="368" reason="4">
<taxonomicName id="7ECD4D69FFFEFFDCFCC74F89FB149BA2" ID-CoL="5JVG" authority="SONG &amp; LIANG" authorityName="Song &amp; Szwedo &amp; Wang &amp; Liang" authorityYear="2016" box="[878,1266,429,451]" class="Insecta" family="Dictyopharidae" genus="Madagascaritia" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="368" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8BB9EAF8FFFEFFDCFCC74F89FBD89BA2" bold="true" box="[878,1086,429,450]" italics="true" pageId="19" pageNumber="368">MADAGASCARITIA</emphasis>
SONG &amp; LIANG
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<taxonomicNameLabel id="908A5783FFFEFFDCFB534F94FA8B9BA2" box="[1274,1389,429,450]" pageId="19" pageNumber="368" rank="genus">GEN. NOV.</taxonomicNameLabel>
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<paragraph id="B97236EAFFFEFFDCFC934FF0FC259B89" blockId="19.[826,1441,468,551]" box="[826,963,468,489]" pageId="19" pageNumber="368">
<emphasis id="8BB9EAF8FFFEFFDCFC934FF0FC259B89" box="[826,963,468,489]" italics="true" pageId="19" pageNumber="368">
<typeStatus id="66768848FFFEFFDCFC934FF0FC889B89" box="[826,878,468,489]" pageId="19" pageNumber="368">Type</typeStatus>
species
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<paragraph id="B97236EAFFFEFFDCFC934FD6FBC39847" blockId="19.[826,1441,468,551]" pageId="19" pageNumber="368">
<taxonomicName id="7ECD4D69FFFEFFDCFC934FD6FAE09868" authority="Song &amp; Liang" authorityName="Song &amp; Szwedo &amp; Wang &amp; Liang" authorityYear="2016" box="[826,1286,498,520]" class="Insecta" family="Dictyopharidae" genus="Madagascaritia" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="368" phylum="Arthropoda" rank="species" species="angusta" status="sp. nov.">
<emphasis id="8BB9EAF8FFFEFFDCFC934FD6FBB39868" box="[826,1109,498,520]" italics="true" pageId="19" pageNumber="368">Madagascaritia angusta</emphasis>
Song &amp; Liang
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<taxonomicNameLabel id="908A5783FFFEFFDCFAB84FD6FA959867" box="[1297,1395,498,519]" pageId="19" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation.
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<emphasis id="8BB9EAF8FFFEFFDCFC934C75FC4C9806" box="[826,938,593,614]" italics="true" pageId="19" pageNumber="368">Diagnosis</emphasis>
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This new genus can be separated from other
<taxonomicName id="7ECD4D69FFFEFFDCFA924C4BFA4798E5" authorityName="Emeljanov" authorityYear="1979" box="[1339,1441,623,645]" class="Insecta" family="Dictyopharidae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="tribe" tribe="Aluntiini">Aluntiini</taxonomicName>
genera by the following combination of characters: cephalic process with dorsal surface very narrow and compressed laterally before the eyes, so it becomes nearly triangular in cross section; frons with intermediate carinae strongly elevated, median carina incomplete or complete but obscure in middle; tegmina with costal cell distinctly narrow, coriaceous, opaque, and bright yellow, with suboblique veins (although observed with difficulty because of sclerotic surface), without dendroid secondary veins; basal three-quarters with dendroid nodose secondary veins densely interspersed among longitudinal veins; Sc+R, M, and CuA first branching nearly abreast.
</paragraph>
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<emphasis id="8BB9EAF8FFFEFFDCFC934A19FC5D9E32" box="[826,955,1085,1106]" italics="true" pageId="19" pageNumber="368">Description</emphasis>
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<paragraph id="B97236EAFFFEFFDCFC934A7FFAB49EEF" blockId="19.[826,1442,1085,1902]" pageId="19" pageNumber="368">General colour ochraceous in dead specimens (probably viridescent or stramineous green in life).</paragraph>
<paragraph id="B97236EAFFFEFFDCFCFA4ABCFA479C18" blockId="19.[826,1442,1085,1902]" pageId="19" pageNumber="368">
Head (
<figureCitation id="21F62A6FFFFEFFDCFC364ABCFBFA9ECE" box="[927,1052,1176,1198]" captionStart="Figure 11" captionStartId="20.[144,223,1828,1847]" captionTargetBox="[209,1360,196,1793]" captionTargetId="figure-0@20.[194,1373,193,1798]" captionTargetPageId="20" captionText="Figure 11. Madagascaritia angusta sp. nov. A, head, pronotum, and mesonotum, dorsal view; B, head, lateral view; C, head, ventral view; D, pygofer and gonostyles, ventral view; E, male genitalia, lateral view; F, pygofer and segment X, dorsal view; G, aedeagus, ventral view; H, aedeagus, lateral view; I, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5357664" httpUri="https://zenodo.org/record/5357664/files/figure.png" pageId="19" pageNumber="368">Fig. 11AC</figureCitation>
) distinctly elongate, 1.51.8 times as long as pronotum and mesonotum combined. Cephalic process with dorsal surface very narrow and compressed laterally before the eyes in dorsal view (
<figureCitation id="21F62A6FFFFEFFDCFCEB4B36FC409F48" box="[834,934,1298,1320]" captionStart="Figure 11" captionStartId="20.[144,223,1828,1847]" captionTargetBox="[209,1360,196,1793]" captionTargetId="figure-0@20.[194,1373,193,1798]" captionTargetPageId="20" captionText="Figure 11. Madagascaritia angusta sp. nov. A, head, pronotum, and mesonotum, dorsal view; B, head, lateral view; C, head, ventral view; D, pygofer and gonostyles, ventral view; E, male genitalia, lateral view; F, pygofer and segment X, dorsal view; G, aedeagus, ventral view; H, aedeagus, lateral view; I, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5357664" httpUri="https://zenodo.org/record/5357664/files/figure.png" pageId="19" pageNumber="368">Fig. 11A</figureCitation>
), so it becomes nearly triangular in cross section. Vertex (
<figureCitation id="21F62A6FFFFEFFDCFC5D4B15FBB29F27" box="[1012,1108,1329,1351]" captionStart="Figure 11" captionStartId="20.[144,223,1828,1847]" captionTargetBox="[209,1360,196,1793]" captionTargetId="figure-0@20.[194,1373,193,1798]" captionTargetPageId="20" captionText="Figure 11. Madagascaritia angusta sp. nov. A, head, pronotum, and mesonotum, dorsal view; B, head, lateral view; C, head, ventral view; D, pygofer and gonostyles, ventral view; E, male genitalia, lateral view; F, pygofer and segment X, dorsal view; G, aedeagus, ventral view; H, aedeagus, lateral view; I, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5357664" httpUri="https://zenodo.org/record/5357664/files/figure.png" pageId="19" pageNumber="368">Fig. 11A</figureCitation>
) with median carina absent, only median furrowed suture visible; lateral carinae strongly ridged, convergent anteriorly between eyes, slightly sinuate in front of eyes, and then narrowed and nearly subparallel towards apex, but apex acuate; posterior margin weakly ridged, arcuately concave. Frons (
<figureCitation id="21F62A6FFFFEFFDCFCEB4BCCFC479F9F" box="[834,929,1512,1535]" captionStart="Figure 11" captionStartId="20.[144,223,1828,1847]" captionTargetBox="[209,1360,196,1793]" captionTargetId="figure-0@20.[194,1373,193,1798]" captionTargetPageId="20" captionText="Figure 11. Madagascaritia angusta sp. nov. A, head, pronotum, and mesonotum, dorsal view; B, head, lateral view; C, head, ventral view; D, pygofer and gonostyles, ventral view; E, male genitalia, lateral view; F, pygofer and segment X, dorsal view; G, aedeagus, ventral view; H, aedeagus, lateral view; I, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5357664" httpUri="https://zenodo.org/record/5357664/files/figure.png" pageId="19" pageNumber="368">Fig. 11C</figureCitation>
) with lateral carinae narrow and elongate; intermediate carinae strongly elevated, converging posteriorly and extending before eyes; median carina incomplete, or complete but obscure in middle. Clypeus, rostrum, eyes, ocelli, and antennae similar to
<taxonomicName id="7ECD4D69FFFEFFDCFAEC4847FA7C9C18" authorityName="Stal" authorityYear="1866" box="[1349,1434,1635,1656]" class="Insecta" family="Dictyopharidae" genus="Aluntia" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="8BB9EAF8FFFEFFDCFAEC4847FA7C9C18" box="[1349,1434,1635,1656]" italics="true" pageId="19" pageNumber="368">Aluntia</emphasis>
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.
</paragraph>
<paragraph id="B97236EAFFFEFFDCFCFA48A6FA479D0E" blockId="19.[826,1442,1085,1902]" pageId="19" pageNumber="368">
Pronotum and mesonotum (
<figureCitation id="21F62A6FFFFEFFDCFB0748A5FAF59CF8" box="[1198,1299,1665,1688]" captionStart="Figure 11" captionStartId="20.[144,223,1828,1847]" captionTargetBox="[209,1360,196,1793]" captionTargetId="figure-0@20.[194,1373,193,1798]" captionTargetPageId="20" captionText="Figure 11. Madagascaritia angusta sp. nov. A, head, pronotum, and mesonotum, dorsal view; B, head, lateral view; C, head, ventral view; D, pygofer and gonostyles, ventral view; E, male genitalia, lateral view; F, pygofer and segment X, dorsal view; G, aedeagus, ventral view; H, aedeagus, lateral view; I, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5357664" httpUri="https://zenodo.org/record/5357664/files/figure.png" pageId="19" pageNumber="368">Fig. 11A</figureCitation>
) similar to
<taxonomicName id="7ECD4D69FFFEFFDCFC934884FC769CD5" authorityName="Stal" authorityYear="1866" box="[826,912,1696,1717]" class="Insecta" family="Dictyopharidae" genus="Aluntia" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="8BB9EAF8FFFEFFDCFC934884FC769CD5" box="[826,912,1696,1717]" italics="true" pageId="19" pageNumber="368">Aluntia</emphasis>
</taxonomicName>
. Tegmina (
<figureCitation id="21F62A6FFFFEFFDCFBB84884FB849CD6" box="[1041,1122,1696,1718]" captionStart="Figure 2" captionStartId="7.[164,243,1509,1528]" captionTargetBox="[177,1434,208,1469]" captionTargetId="figure-110@7.[164,1444,193,1479]" captionTargetPageId="7" captionText="Figure 2. Tegmina (AF, G, I) and hindwings (H, J) of Aluntiini and Arjunini trib. nov. A, Aluntia longicephalica sp. nov.; B, Aluntia schimperii; C, Dendrophora borneensis; D, Dictyomorpha moluccana; E, Indodictyophara lobosa; F, Madagascaritia hova; G, H, Arjuna scriba; I, J, Pippax opilionoides. Abbreviations: ac, apical cell; dsv, dendroid secondary veins." figureDoi="http://doi.org/10.5281/zenodo.5357648" httpUri="https://zenodo.org/record/5357648/files/figure.png" pageId="19" pageNumber="368">Fig. 2F</figureCitation>
) elongate and broad, translucent, and dull whitish, nearly 2.5 times as long as broad; costal cell very narrow, mostly coriaceous, opaque, and bright yellow in colour, with suboblique veins (although observed with difficulty because of sclerotic surface), without dendroid secondary veins; apex rounded; basal two-thirds with dendroid nodose
</paragraph>
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